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<document id="91962AA2A5DCF5EDBA111CF439D0298E" ID-DOI="10.4202/app.00061.2014" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626402" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707249081010" checkinUser="felipe" docAuthor="Denayer, Julien" docDate="2016" docId="03B687FCFFE91949FFBDFBAC329C867B" docLanguage="en" docName="ActaPalaeontolPol.61.1.51-70.pdf" docOrigin="Acta Palaeontologica Polonica 61 (1)" docSource="http://dx.doi.org/10.4202/app.00061.2014" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Pseudoendophyllum Onoprienko 1979" docType="treatment" docVersion="3" lastPageNumber="60" masterDocId="FF8FFF84FFE01940FFDFFFE0367F8762" masterDocTitle="Rugose corals across the Devonian-Carboniferous boundary in NW Turkey" masterLastPageNumber="70" masterPageNumber="51" pageNumber="60" updateTime="1713405178588" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DD0A40F18100060EB00BDDDE0CB063EC">Rugose corals across the Devonian-Carboniferous boundary in NW Turkey</mods:title>
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<mods:identifier id="B81162B3AF531C097B3B2F9BE92A9302" type="DOI">10.4202/app.00061.2014</mods:identifier>
<mods:identifier id="CE7A19B7E631093293FA62ACA782A031" type="ISSN">1732-2421</mods:identifier>
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<treatment id="03B687FCFFE91949FFBDFBAC329C867B" LSID="urn:lsid:plazi:treatment:03B687FCFFE91949FFBDFBAC329C867B" httpUri="http://treatment.plazi.org/id/03B687FCFFE91949FFBDFBAC329C867B" lastPageNumber="60" pageId="9" pageNumber="60">
<subSubSection id="C3056561FFE91949FFBDFBAC34BF830A" box="[98,704,1100,1128]" pageId="9" pageNumber="60" type="nomenclature">
<paragraph id="8BA036EAFFE91949FFBDFBAC34BF830A" blockId="9.[98,704,1100,1128]" box="[98,704,1100,1128]" pageId="9" pageNumber="60">
<heading id="D0E88186FFE91949FFBDFBAC34BF830A" box="[98,704,1100,1128]" fontSize="12" level="2" pageId="9" pageNumber="60" reason="3">
Genus
<taxonomicName id="4C1F4D69FFE91949FF64FBAC34BF830A" authority="Onoprienko, 1979 b" authorityName="Onoprienko" authorityYear="1979" box="[187,704,1100,1128]" class="Anthozoa" family="Kyphophyllidae" genus="Pseudoendophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FF64FBAC37C0830A" box="[187,447,1100,1128]" italics="true" pageId="9" pageNumber="60">Pseudoendophyllum</emphasis>
<bibRefCitation id="EF8E4B1BFFE91949FE18FBAC34BF830A" author="Onoprienko, Y." box="[455,704,1100,1128]" pageId="9" pageNumber="60" pagination="29 - 32" refId="ref15512" refString="Onoprienko, Y. 1979 b. Question about relationship between genera Endophyllum and Tabulophyllum (Rugosa) [in Russian]. In: M. N. Gramm (ed.), Iskopaemye Bespozvonocnye Dal'nego Vostoka (dannye po novym nahodkam), 29 - 32. Akademia Nauk SSSR, Dal'nevostocnyj Naucnyj Centr, Vladivostok." type="book chapter" year="1979">Onoprienko, 1979b</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3056561FFE91949FFBDFB98329C867B" pageId="9" pageNumber="60" type="materials_examined">
<paragraph id="8BA036EAFFE91949FFBDFB9837E783C8" blockId="9.[98,770,1144,1194]" pageId="9" pageNumber="60">
<materialsCitation id="3B773CB7FFE91949FFBDFB9837EB83C8" country="Russia" location="Famennian of Novaya Zemlya" pageId="9" pageNumber="60" specimenCount="1" typeStatus="holotype">
<emphasis id="B96BEAF8FFE91949FFBDFB9836A783EF" box="[98,216,1144,1165]" italics="true" pageId="9" pageNumber="60">
<typeStatus id="54A48848FFE91949FFBDFB9836F083EF" box="[98,143,1144,1165]" pageId="9" pageNumber="60">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C1F4D69FFE91949FF3AFB98343983EF" authority="Gorsky, 1935" authorityName="Gorsky" authorityYear="1935" box="[229,582,1144,1165]" class="Pucciniomycetes" family="Pucciniaceae" genus="Endophyllum" kingdom="Fungi" order="Pucciniales" pageId="9" pageNumber="60" phylum="Basidiomycota" rank="species" species="nalivkini">
<emphasis id="B96BEAF8FFE91949FF3AFB9837C183EF" box="[229,446,1144,1165]" italics="true" pageId="9" pageNumber="60">Endophyllum nalivkini</emphasis>
<bibRefCitation id="EF8E4B1BFFE91949FE1CFB98343983EF" author="Gorsky, I. &amp; Gorskij, I." box="[451,582,1144,1165]" pageId="9" pageNumber="60" pagination="6 - 122" refId="ref14024" refString="Gorsky, I. [Gorskij, I.] 1935. Some Coelenterata from the Lower Carboniferous beds of the Nowaya Zemlya [in Russian]. Trudy Arkticeskogo Instituta 28: 6 - 122." type="journal article" year="1935">Gorsky, 1935</bibRefCitation>
</taxonomicName>
; uppermost
<location id="8EC06031FFE91949FD66FB98373883C8" LSID="urn:lsid:plazi:treatment:03B687FCFFE91949FFBDFBAC329C867B:8EC06031FFE91949FD66FB98373883C8" country="Russia" name="Famennian of Novaya Zemlya" pageId="9" pageNumber="60">Famennian of Novaya Zemlya</location>
,
<collectingCountry id="F308767AFFE91949FE8EFB7537EB83C8" box="[337,404,1173,1194]" name="Russia" pageId="9" pageNumber="60">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA036EAFFE91949FFBDFB5D378883B4" blockId="9.[98,503,1213,1238]" box="[98,503,1213,1238]" pageId="9" pageNumber="60">
<emphasis id="B96BEAF8FFE91949FFBDFB5D36AB83B4" box="[98,212,1213,1238]" italics="true" pageId="9" pageNumber="60">Diagnosis</emphasis>
.—See
<bibRefCitation id="EF8E4B1BFFE91949FEF8FB5D378C83B4" author="Berkowski, B." box="[295,499,1213,1238]" pageId="9" pageNumber="60" pagination="1 - 87" refId="ref12904" refString="Berkowski, B. 2002. Famennian Rugosa and Heterocorallia from Southern Poland. Palaeontologia Polonica 61: 1 - 87." type="journal article" year="2002">Berkowski (2002)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA036EAFFE91949FFBDFB08329C867B" blockId="9.[98,771,1256,1960]" lastBlockId="9.[808,1481,160,281]" pageId="9" pageNumber="60">
<emphasis id="B96BEAF8FFE91949FFBDFB0836BA8263" box="[98,197,1256,1281]" italics="true" pageId="9" pageNumber="60">Remarks</emphasis>
.—
<bibRefCitation id="EF8E4B1BFFE91949FF3AFB0837AD8263" author="Onoprienko, Y." box="[229,466,1256,1281]" pageId="9" pageNumber="60" pagination="29 - 32" refId="ref15512" refString="Onoprienko, Y. 1979 b. Question about relationship between genera Endophyllum and Tabulophyllum (Rugosa) [in Russian]. In: M. N. Gramm (ed.), Iskopaemye Bespozvonocnye Dal'nego Vostoka (dannye po novym nahodkam), 29 - 32. Akademia Nauk SSSR, Dal'nevostocnyj Naucnyj Centr, Vladivostok." type="book chapter" year="1979">Onoprienko (1979b)</bibRefCitation>
created
<taxonomicName id="4C1F4D69FFE91949FDE2FB0836F38243" authorityName="Onoprienko" authorityYear="1979" class="Anthozoa" family="Kyphophyllidae" genus="Pseudoendophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FDE2FB0836F38243" italics="true" pageId="9" pageNumber="60">Pseudoendophyllum</emphasis>
</taxonomicName>
for the corals attributed to
<taxonomicName id="4C1F4D69FFE91949FE0AFAE8357D8243" authority="Torley, 1933" authorityName="Torley" authorityYear="1933" box="[469,770,1288,1313]" class="Pucciniomycetes" family="Pucciniaceae" genus="Endophyllum" kingdom="Fungi" order="Pucciniales" pageId="9" pageNumber="60" phylum="Basidiomycota" rank="genus">
<emphasis id="B96BEAF8FFE91949FE0AFAE834148243" box="[469,619,1288,1313]" italics="true" pageId="9" pageNumber="60">Endophyllum</emphasis>
<bibRefCitation id="EF8E4B1BFFE91949FDAAFAE8357D8243" author="Torley, K." box="[629,770,1288,1313]" pageId="9" pageNumber="60" pagination="630 - 633" refId="ref17084" refString="Torley, K. 1933. Uber Endophyllum bowerbanki M. - E. and H. Deutsche Geologische Gesellschaft Zeitschrift 85: 630 - 633." type="journal article" year="1933">Torley, 1933</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EF8E4B1BFFE91949FF55FAC937548220" author="Gorsky, I. &amp; Gorskij, I." box="[138,299,1321,1346]" pageId="9" pageNumber="60" pagination="6 - 122" refId="ref14024" refString="Gorsky, I. [Gorskij, I.] 1935. Some Coelenterata from the Lower Carboniferous beds of the Nowaya Zemlya [in Russian]. Trudy Arkticeskogo Instituta 28: 6 - 122." type="journal article" year="1935">Gorsky (1935</bibRefCitation>
,
<bibRefCitation id="EF8E4B1BFFE91949FEE3FAC9370C8220" author="Gorsky, I. &amp; Gorskij, I." box="[316,371,1321,1346]" pageId="9" pageNumber="60" pagination="1 - 221" refId="ref14061" refString="Gorsky, I. [Gorskij, I.] 1938. Carboniferous corals from Novaya Zemlya [in Russian]. In: D. V. Nalivkin (ed.), Paleontologia Sovetskoj Arktiki. Vsesouznij Arkticeskij Institut, Trudy, Leningrad 93: 1 - 221." type="journal article" year="1938">1938</bibRefCitation>
) in the uppermost Famennian of Novaya Zemlya. As noticed by
<bibRefCitation id="EF8E4B1BFFE91949FE0CFAA934DE8200" author="Berkowski, B." box="[467,673,1353,1378]" pageId="9" pageNumber="60" pagination="1 - 87" refId="ref12904" refString="Berkowski, B. 2002. Famennian Rugosa and Heterocorallia from Southern Poland. Palaeontologia Polonica 61: 1 - 87." type="journal article" year="2002">Berkowski (2002)</bibRefCitation>
,
<bibRefCitation id="EF8E4B1BFFE91949FD70FAA936DE82E0" author="Gorsky, I. &amp; Gorskij, I." pageId="9" pageNumber="60" pagination="6 - 122" refId="ref14024" refString="Gorsky, I. [Gorskij, I.] 1935. Some Coelenterata from the Lower Carboniferous beds of the Nowaya Zemlya [in Russian]. Trudy Arkticeskogo Instituta 28: 6 - 122." type="journal article" year="1935">Gorsky 1935</bibRefCitation>
) described his corals as
<taxonomicName id="4C1F4D69FFE91949FE05FA89340D82E0" authorityName="Torley" authorityYear="1933" box="[474,626,1385,1410]" class="Pucciniomycetes" family="Pucciniaceae" genus="Endophyllum" kingdom="Fungi" order="Pucciniales" pageId="9" pageNumber="60" phylum="Basidiomycota" rank="genus">
<emphasis id="B96BEAF8FFE91949FE05FA89340D82E0" box="[474,626,1385,1410]" italics="true" pageId="9" pageNumber="60">Endophyllum</emphasis>
</taxonomicName>
because he could not highlight any difference with the Devonian genus. Nevertheless, no
<taxonomicName id="4C1F4D69FFE91949FEBDFA4A378782A1" authorityName="Torley" authorityYear="1933" box="[354,504,1450,1475]" class="Pucciniomycetes" family="Pucciniaceae" genus="Endophyllum" kingdom="Fungi" order="Pucciniales" pageId="9" pageNumber="60" phylum="Basidiomycota" rank="genus">
<emphasis id="B96BEAF8FFE91949FEBDFA4A378782A1" box="[354,504,1450,1475]" italics="true" pageId="9" pageNumber="60">Endophyllum</emphasis>
</taxonomicName>
is known between the Frasnian and the uppermost Famennian, so the uppermost Famennian
<taxonomicName id="4C1F4D69FFE91949FEF4FA0B37BE8166" authorityName="Torley" authorityYear="1933" box="[299,449,1515,1540]" class="Pucciniomycetes" family="Pucciniaceae" genus="Endophyllum" kingdom="Fungi" order="Pucciniales" pageId="9" pageNumber="60" phylum="Basidiomycota" rank="genus">
<emphasis id="B96BEAF8FFE91949FEF4FA0B37BE8166" box="[299,449,1515,1540]" italics="true" pageId="9" pageNumber="60">Endophyllum</emphasis>
</taxonomicName>
are possibly homeomorphs of the Middle Devonian ones and should thus be considered as Elvis taxa (
<bibRefCitation id="EF8E4B1BFFE91949FEE3F9CB37888126" author="Berkowski, B." box="[316,503,1579,1604]" pageId="9" pageNumber="60" pagination="285 - 290" refId="ref12875" refString="Berkowski, B. 2001. Famennian colonial Rugosa from southern Poland: recovery and extinction. Bulletin of the Tohoku University Museum 1: 285 - 290." type="journal article" year="2001">Berkowski 2001</bibRefCitation>
). In
<collectingCountry id="F308767AFFE91949FDF3F9CB34048126" box="[556,635,1579,1604]" name="Poland" pageId="9" pageNumber="60">Poland</collectingCountry>
,
<bibRefCitation id="EF8E4B1BFFE91949FD56F9CB36DC8107" author="Berkowski, B." pageId="9" pageNumber="60" pagination="1 - 87" refId="ref12904" refString="Berkowski, B. 2002. Famennian Rugosa and Heterocorallia from Southern Poland. Palaeontologia Polonica 61: 1 - 87." type="journal article" year="2002">Berkowski 2002</bibRefCitation>
) documented two species from the uppermost part of Middle Famennian (
<taxonomicName id="4C1F4D69FFE91949FEAFF98C34A581E7" authority="Zone" authorityName="Zone" box="[368,730,1644,1669]" family="Palmatolepidae" genus="Palmatolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="marginifera">
<emphasis id="B96BEAF8FFE91949FEAFF98C34E981E7" box="[368,662,1644,1669]" italics="true" pageId="9" pageNumber="60">Palmatolepis marginifera</emphasis>
Zone
</taxonomicName>
) to uppermost Famennian (
<taxonomicName id="4C1F4D69FFE91949FEA0F96C348881C7" authority="Zone" authorityName="Zone" box="[383,759,1676,1701]" genus="Siphonodella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="praesulcata">
<emphasis id="B96BEAF8FFE91949FEA0F96C34D481C7" box="[383,683,1676,1701]" italics="true" pageId="9" pageNumber="60">Siphonodella praesulcata</emphasis>
Zone
</taxonomicName>
). The origin of the genus should consequently be situated, at least in the Early Famennian.
<bibRefCitation id="EF8E4B1BFFE91949FDDBF92D357D8184" author="Onoprienko, Y." box="[516,770,1741,1766]" pageId="9" pageNumber="60" pagination="29 - 32" refId="ref15512" refString="Onoprienko, Y. 1979 b. Question about relationship between genera Endophyllum and Tabulophyllum (Rugosa) [in Russian]. In: M. N. Gramm (ed.), Iskopaemye Bespozvonocnye Dal'nego Vostoka (dannye po novym nahodkam), 29 - 32. Akademia Nauk SSSR, Dal'nevostocnyj Naucnyj Centr, Vladivostok." type="book chapter" year="1979">Onoprienko’s (1979b)</bibRefCitation>
choice to create
<taxonomicName id="4C1F4D69FFE91949FEEAF90D34628064" authorityName="Onoprienko" authorityYear="1979" box="[309,541,1773,1798]" class="Anthozoa" family="Kyphophyllidae" genus="Pseudoendophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FEEAF90D34628064" box="[309,541,1773,1798]" italics="true" pageId="9" pageNumber="60">Pseudoendophyllum</emphasis>
</taxonomicName>
is justified if the Middle and Upper Devonian forms have a different origin.
<bibRefCitation id="EF8E4B1BFFE91949FF45F8CE37148025" author="Berkowski, B." box="[154,363,1838,1863]" pageId="9" pageNumber="60" pagination="1 - 87" refId="ref12904" refString="Berkowski, B. 2002. Famennian Rugosa and Heterocorallia from Southern Poland. Palaeontologia Polonica 61: 1 - 87." type="journal article" year="2002">Berkowski (2002)</bibRefCitation>
proposes two hypotheses for this origin: (i)
<taxonomicName id="4C1F4D69FFE91949FF09F8AE37C48005" authorityName="Onoprienko" authorityYear="1979" box="[214,443,1870,1895]" class="Anthozoa" family="Kyphophyllidae" genus="Pseudoendophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FF09F8AE37C48005" box="[214,443,1870,1895]" italics="true" pageId="9" pageNumber="60">Pseudoendophyllum</emphasis>
</taxonomicName>
evolved from
<taxonomicName id="4C1F4D69FFE91949FDBFF8AE357D8005" authorityName="Birenheide" authorityYear="1962" box="[608,770,1870,1895]" class="Anthozoa" family="Endophyllidae" genus="Smithiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FDBFF8AE357D8005" box="[608,770,1870,1895]" italics="true" pageId="9" pageNumber="60">Smithiphyllum</emphasis>
</taxonomicName>
by acquiring a cerioid habit; (ii)
<taxonomicName id="4C1F4D69FFE91949FE37F88E34B280E5" authorityName="Onoprienko" authorityYear="1979" box="[488,717,1902,1927]" class="Anthozoa" family="Kyphophyllidae" genus="Pseudoendophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FE37F88E34B280E5" box="[488,717,1902,1927]" italics="true" pageId="9" pageNumber="60">Pseudoendophyllum</emphasis>
</taxonomicName>
is a
<taxonomicName id="4C1F4D69FFE91949FFBDF86F377380CA" authorityName="Fenton &amp; Fenton" authorityYear="1924" box="[98,268,1935,1960]" class="Anthozoa" family="Chonophyllidae" genus="Tabulophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FFBDF86F377380CA" box="[98,268,1935,1960]" italics="true" pageId="9" pageNumber="60">Tabulophyllum</emphasis>
</taxonomicName>
that became colonial (this second option is also favoured by
<bibRefCitation id="EF8E4B1BFFE91949FC27FF4032B687DB" author="Jell, J. S. &amp; Hill, D." box="[1016,1225,160,185]" pageId="9" pageNumber="60" pagination="1 - 16" refId="ref14431" refString="Jell, J. S. and Hill, D. 1970. The Devonian coral fauna of the Point Hibbs Limestone, Tasmania. Papers and Proceedings of the Royal Society of Tasmania 104: 1 - 16." type="journal article" year="1970">Jell and Hill 1970</bibRefCitation>
for the origin of the Middle Devonian
<taxonomicName id="4C1F4D69FFE91949FC2CFF2032F887BB" authorityName="Torley" authorityYear="1933" box="[1011,1159,192,217]" class="Pucciniomycetes" family="Pucciniaceae" genus="Endophyllum" kingdom="Fungi" order="Pucciniales" pageId="9" pageNumber="60" phylum="Basidiomycota" rank="genus">
<emphasis id="B96BEAF8FFE91949FC2CFF2032F887BB" box="[1011,1159,192,217]" italics="true" pageId="9" pageNumber="60">Endophyllum</emphasis>
</taxonomicName>
).
<taxonomicName id="4C1F4D69FFE91949FB42FF20333A87BB" authorityName="Fenton &amp; Fenton" authorityYear="1924" box="[1181,1349,192,217]" class="Anthozoa" family="Chonophyllidae" genus="Tabulophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FB42FF20333A87BB" box="[1181,1349,192,217]" italics="true" pageId="9" pageNumber="60">Tabulophyllum</emphasis>
</taxonomicName>
being present in the Famennian (Poty 1984;
<bibRefCitation id="EF8E4B1BFFE91949FB7BFF003320879B" author="Berkowski, B." box="[1188,1375,224,249]" pageId="9" pageNumber="60" pagination="1 - 87" refId="ref12904" refString="Berkowski, B. 2002. Famennian Rugosa and Heterocorallia from Southern Poland. Palaeontologia Polonica 61: 1 - 87." type="journal article" year="2002">Berkowski 2002</bibRefCitation>
), it might be at the origin of
<taxonomicName id="4C1F4D69FFE91949FC28FEE032A4867B" authorityName="Onoprienko" authorityYear="1979" box="[1015,1243,256,281]" class="Anthozoa" family="Kyphophyllidae" genus="Pseudoendophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="60" phylum="Cnidaria" rank="genus">
<emphasis id="B96BEAF8FFE91949FC28FEE032A4867B" box="[1015,1243,256,281]" italics="true" pageId="9" pageNumber="60">Pseudoendophyllum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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