267 lines
33 KiB
XML
267 lines
33 KiB
XML
<document id="5DE867FC879063B5233DDC7DDE750D8B" ID-CLB-Dataset="25925" ID-DOI="10.11646/zootaxa.4658.3.8" ID-GBIF-Dataset="65575e2a-b926-423a-a565-82d041ddf1f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3376150" ID-ZooBank="ECF3D8F3-3F41-48DB-B811-6B4265404EE6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566576285773" checkinUser="plazi" docAuthor="Jałoszyński, Paweł, Cheng, Zhi-Fei, Brajczewska, Nastazja & Yin, Zi-Wei" docDate="2019" docId="03B687F9E276FF8257A561CCFDBEFBBD" docLanguage="en" docName="zootaxa.4658.3.8.pdf" docOrigin="Zootaxa 4658 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clidicus occisor Jałoszyński & Cheng & Brajczewska & Yin 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="580" masterDocId="FF8FFF81E271FF8B57326539FFB3FFAE" masterDocTitle="Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="584" masterPageNumber="571" pageNumber="578" updateTime="1698752449993" updateUser="ExternalLinkService">
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<mods:titleInfo id="152B638519433D9F2A4BA75AD1F335A2">
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<mods:title id="19BBC1784F98F1C586A72B406B13D768">Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="E6B03579E34921E8BBA6ED0035666DD6">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="9B6E5DD413DC2C0F67C68624A32CD0B8">Cheng, Zhi-Fei</mods:namePart>
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<mods:namePart id="B02CB02368A31231BCE7AE4567C0BA71">Brajczewska, Nastazja</mods:namePart>
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<mods:roleTerm id="4B7E60FFC7F6B4F5C6501EDA9C049F18">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AE12C7085105F0BE526629A850FD6CBB">Yin, Zi-Wei</mods:namePart>
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<mods:title id="15D5EDC5BDA2F17E9B5C3BF6090BAFA7">Zootaxa</mods:title>
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<mods:part id="2A159E36FCFA2BF893027230A9991B4C">
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<mods:date id="4FAF577A6A8DB8005CF10A42A27068AD">2019</mods:date>
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<mods:number id="9B50DCC89685577701D45BE9852579A3">2019-08-23</mods:number>
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<mods:number id="A70B4867F9F7E0E2C387D5855D3C0A7E">4658</mods:number>
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<treatment id="03B687F9E276FF8257A561CCFDBEFBBD" ID-GBIF-Taxon="159243636" LSID="urn:lsid:plazi:treatment:03B687F9E276FF8257A561CCFDBEFBBD" httpUri="http://treatment.plazi.org/id/03B687F9E276FF8257A561CCFDBEFBBD" lastPageId="9" lastPageNumber="580" pageId="7" pageNumber="578">
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<subSubSection id="C3056564E276FF8C57A561CCFD8AFAA1" box="[151,569,1268,1296]" pageId="7" pageNumber="578" type="nomenclature">
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<paragraph id="8BA036EFE276FF8C57A561CCFD8AFAA1" blockId="7.[151,569,1268,1332]" box="[151,569,1268,1296]" pageId="7" pageNumber="578">
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<heading id="D0E88183E276FF8C57A561CCFD8AFAA1" bold="true" box="[151,569,1268,1296]" fontSize="11" level="1" pageId="7" pageNumber="578" reason="1">
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<emphasis id="B96BEAFDE276FF8C57A561CCFD8AFAA1" bold="true" box="[151,569,1268,1296]" pageId="7" pageNumber="578">
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<taxonomicName id="4C1F4D6CE276FF8C57A561CCFE5CFAA1" ID-CoL="8XXTB" authority="Jaloszynski" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[151,495,1268,1296]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="occisor" status="sp. nov.">
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<emphasis id="B96BEAFDE276FF8C57A561CCFEEBFAA1" bold="true" box="[151,344,1269,1296]" italics="true" pageId="7" pageNumber="578">Clidicus occisor</emphasis>
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Jałoszyński
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</taxonomicName>
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<taxonomicNameLabel id="A2585786E276FF8C56C461CCFD8AFAA1" box="[502,569,1269,1295]" pageId="7" pageNumber="578" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3056564E276FF8C57A56020FE94FA9A" box="[151,295,1305,1332]" pageId="7" pageNumber="578" type="description">
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<paragraph id="8BA036EFE276FF8C57A56020FE94FA9A" blockId="7.[151,569,1268,1332]" box="[151,295,1305,1332]" pageId="7" pageNumber="578">
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(
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<figureCitation id="13242A6AE276FF8C57AD6020FEACFA9A" box="[159,287,1305,1332]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1553,1578]" captionTargetBox="[165,1432,345,1516]" captionTargetId="figure@8.[151,1436,345,1528]" captionTargetPageId="8" captionText="FIGURES 21–26. Clidicus occisor Jałoszyński sp. n. Habitus of male (21); head of male in dorsal view (22), right antenna of male in dorsal view (23); aedeagus in ventral (24) and lateral (25) views; ‘sperm pump’ (26)." httpUri="https://zenodo.org/record/3376158/files/figure.png" pageId="7" pageNumber="578">Figs 21–26</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3056564E276FF8257A56059FDBEFBBD" lastPageId="9" lastPageNumber="580" pageId="7" pageNumber="578" type="materials_examined">
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<paragraph id="8BA036EFE276FF8C57A56059FA90FA6D" blockId="7.[151,1437,1376,2015]" pageId="7" pageNumber="578">
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<emphasis id="B96BEAFDE276FF8C57A56059FEECFAD5" bold="true" box="[151,351,1376,1403]" pageId="7" pageNumber="578">Material studied</emphasis>
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.
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<materialsCitation id="3B773CB2E276FF8C565C6059FA90FA6D" ID-GBIF-Occurrence="2351006010" collectingDate="1985-08-13" collectionCode="EUMJ" collectorName="K. Ishikawa" country="Philippines" elevation="50" location="Luan" pageId="7" pageNumber="578" specimenCount="1" specimenCount-male="1" stateProvince="South Cotabato" typeStatus="holotype">
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<emphasis id="B96BEAFDE276FF8C565C6059FDA5FAD5" bold="true" box="[366,534,1376,1403]" pageId="7" pageNumber="578">
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<typeStatus id="54A4884DE276FF8C565C6059FDA2FAD5" box="[366,529,1376,1403]" pageId="7" pageNumber="578" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D19FD66E276FF8C552C6058FD80FAD5" box="[542,563,1377,1403]" pageId="7" pageNumber="578" type="male">♂</specimenCount>
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:
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<emphasis id="B96BEAFDE276FF8C55776059FD4BFAD5" bold="true" box="[581,760,1376,1403]" pageId="7" pageNumber="578">
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<collectingCountry id="F308767FE276FF8C55776059FD4BFAD5" box="[581,760,1376,1403]" name="Philippines" pageId="7" pageNumber="578">PHILIPPINES</collectingCountry>
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</emphasis>
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||
, with three labels: “(PHILIPPINES) /
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<location id="8EC06034E276FF8C538D6058FB48FAD5" LSID="urn:lsid:plazi:treatment:03B687F9E276FF8257A561CCFDBEFBBD:8EC06034E276FF8C538D6058FB48FAD5" box="[1215,1275,1377,1403]" country="Philippines" name="Luan" pageId="7" pageNumber="578" stateProvince="South Cotabato">Luan</location>
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, alt.
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<quantity id="4CE79B0AE276FF8C52016059FADDFAD5" box="[1331,1390,1376,1403]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="578" unit="m" value="50.0">
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<elevation id="0032D1DCE276FF8C52016059FADDFAD5" box="[1331,1390,1376,1403]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="578" unit="m" value="50.0">50 m</elevation>
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</quantity>
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/ S.
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<collectingRegion id="49DBF80DE276FF8C57A560BCFE8DFA31" box="[151,318,1413,1439]" country="Philippines" name="South Cotabato" pageId="7" pageNumber="578">Cotabato Prov.</collectingRegion>
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/ Mindanao /
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||
<date id="FFA1102FE276FF8C56E860BCFDCAFA31" box="[474,633,1412,1439]" pageId="7" pageNumber="578" value="1985-08-13">
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||
<collectingDate id="EFE5E9C7E276FF8C56E860BCFDCAFA31" box="[474,633,1412,1439]" pageId="7" pageNumber="578" value="1985-08-13">13. VIII. 1985</collectingDate>
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||
</date>
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||
/
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||
<collectorName id="26EA5339E276FF8C55BD60BCFCA5FA31" box="[655,790,1413,1439]" pageId="7" pageNumber="578">K. Ishikawa</collectorName>
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||
leg.” [white, printed], “Tullgren sample / of leaf litter / in natural forest” [white, printed], “
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<taxonomicName id="4C1F4D6CE276FF8C553A6090FDD6FA6D" authorityName="Laporte" authorityYear="1832" box="[520,613,1449,1475]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEAFDE276FF8C553A6090FDD6FA6D" box="[520,613,1449,1475]" italics="true" pageId="7" pageNumber="578">Clidicus</emphasis>
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</taxonomicName>
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/
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||
<taxonomicName id="4C1F4D6CE276FF8C55496093FCE6FA6D" authority="Jaloszynski" authorityName="Jaloszynski" box="[635,853,1449,1475]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="occisor">
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<emphasis id="B96BEAFDE276FF8C55496093FD7EFA6D" box="[635,717,1450,1475]" italics="true" pageId="7" pageNumber="578">occisor</emphasis>
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Jałoszyński
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</taxonomicName>
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/
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<typeStatus id="54A4884DE276FF8C54596090FBA6FA6D" box="[875,1045,1449,1475]" pageId="7" pageNumber="578" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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<collectionCode id="ED0EAE2AE276FF8C53FA6090FAA4FA6D" box="[1224,1303,1449,1475]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="7" pageNumber="578">EUMJ</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA036EFE276FF8C57F560F5FBF9F9D9" blockId="7.[151,1437,1376,2015]" pageId="7" pageNumber="578">
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<emphasis id="B96BEAFDE276FF8C57F560F5FEF1FA49" bold="true" box="[199,322,1484,1511]" pageId="7" pageNumber="578">Diagnosis.</emphasis>
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Pronotum with evenly distributed, conspicuously deep and very dense punctures (distances between most punctures shorter than their diameters), lacking median impunctate line or area, cuticle between punctures finely shagreened and moderately glossy, pronotal disc with a sharply marked, narrow median longitudinal groove in posterior 1/4; aedeagus with symmetrical parameres, external margin of each broadly and deeply emarginate in subapical region, in lateral view parameres in subapical region not expanded dorsad.
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</paragraph>
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<paragraph id="8BA036EFE276FF8C57F563B9FC84F94D" blockId="7.[151,1437,1376,2015]" pageId="7" pageNumber="578">
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<emphasis id="B96BEAFDE276FF8C57F563B9FEEAF935" bold="true" box="[199,345,1664,1691]" pageId="7" pageNumber="578">Description.</emphasis>
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Body of male (
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<figureCitation id="13242A6AE276FF8C553163B8FDE0F935" box="[515,595,1665,1691]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1553,1578]" captionTargetBox="[165,1432,345,1516]" captionTargetId="figure@8.[151,1436,345,1528]" captionTargetPageId="8" captionText="FIGURES 21–26. Clidicus occisor Jałoszyński sp. n. Habitus of male (21); head of male in dorsal view (22), right antenna of male in dorsal view (23); aedeagus in ventral (24) and lateral (25) views; ‘sperm pump’ (26)." httpUri="https://zenodo.org/record/3376158/files/figure.png" pageId="7" pageNumber="578">Fig. 21</figureCitation>
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) elongate and slender, head flattened, prothorax and elytra strongly convex; legs and antennae long and slender, BL
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<quantity id="4CE79B0AE276FF8C5569639DFD70F911" box="[603,707,1700,1727]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" pageId="7" pageNumber="578" unit="mm" value="5.25">5.25 mm</quantity>
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; cuticle moderately glossy (more so on head), pigmentation uni- formly reddish brown, setae indistinctly lighter than cuticle.
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</paragraph>
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<paragraph id="8BA036EFE276FF8357F563D4FE3AFEED" blockId="7.[151,1437,1376,2015]" lastBlockId="8.[151,1436,153,323]" lastPageId="8" lastPageNumber="579" pageId="7" pageNumber="578">
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||
Head (
|
||
<figureCitation id="13242A6AE276FF8C562163D4FED6F8A9" box="[275,357,1773,1799]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1553,1578]" captionTargetBox="[165,1432,345,1516]" captionTargetId="figure@8.[151,1436,345,1528]" captionTargetPageId="8" captionText="FIGURES 21–26. Clidicus occisor Jałoszyński sp. n. Habitus of male (21); head of male in dorsal view (22), right antenna of male in dorsal view (23); aedeagus in ventral (24) and lateral (25) views; ‘sperm pump’ (26)." httpUri="https://zenodo.org/record/3376158/files/figure.png" pageId="7" pageNumber="578">Fig. 22</figureCitation>
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) broadest slightly behind half of tempora, HL
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||
<quantity id="4CE79B0AE276FF8C544A63D4FC6CF8A9" box="[888,991,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="578" unit="mm" value="0.9">0.90 mm</quantity>
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, HW
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||
<quantity id="4CE79B0AE276FF8C531063D5FB38F8A9" box="[1058,1163,1772,1799]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="578" unit="mm" value="1.25">1.25 mm</quantity>
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; tempora in dorsal view 3 times as long as eyes, in anterior third straight and slightly divergent posterad, posteriorly strongly and evenly rounded; vertex with a deep subtrapezoidal posteromedian impression and a pair of posterior pits marking sites where dorsal tentorial arms are fused with dorsal wall of head. Sides of vertex convex, frons between antennal insertions nearly flat, entire frons and vertex with narrow median longitudinal groove, supraantennal tubercles small but distinctly elevated; compound eyes small, nearly round, strongly convex but in dorsal view not projecting from head silhouette, finely faceted. Punctures on sides of vertex and frons dense, distinct but shallow and with diffuse mar- gins, mesal margin of each eye and posterolateral margin of each supraantennal tubercle with an irregular group of dense punctures; supraantennal tubercles and posteromedian vertexal impression virtually impunctate. Setae long, sparse, suberect to erect. Antennae (
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<figureCitation id="13242A6AE279FF83551A65D8FDCAFF55" box="[552,633,225,251]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1553,1578]" captionTargetBox="[165,1432,345,1516]" captionTargetId="figure@8.[151,1436,345,1528]" captionTargetPageId="8" captionText="FIGURES 21–26. Clidicus occisor Jałoszyński sp. n. Habitus of male (21); head of male in dorsal view (22), right antenna of male in dorsal view (23); aedeagus in ventral (24) and lateral (25) views; ‘sperm pump’ (26)." httpUri="https://zenodo.org/record/3376158/files/figure.png" pageId="8" pageNumber="579">Fig. 23</figureCitation>
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) slender, scape slightly longer than head, antennomeres II–VII elongate (
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<collectionCode id="ED0EAE2AE279FF8357AD643CFF74FEB1" box="[159,199,261,287]" pageId="8" pageNumber="579">VII</collectionCode>
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indistinctly so), VIII about as long as broad, IX slightly transverse, X distinctly transverse, XI about as long as broad; AnL
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<quantity id="4CE79B0AE279FF83562F6410FE37FEED" box="[285,388,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="3.73" pageId="8" pageNumber="579" unit="mm" value="3.73">3.73 mm</quantity>
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.
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</paragraph>
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<caption id="DF606667E279FF8357A56328FBF6F9E0" httpUri="https://zenodo.org/record/3376158/files/figure.png" pageId="8" pageNumber="579" startId="8.[151,264,1553,1578]" targetBox="[165,1432,345,1516]" targetPageId="8">
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<paragraph id="8BA036EFE279FF8357A56328FBF6F9E0" blockId="8.[151,1436,1553,1614]" pageId="8" pageNumber="579">
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||
<emphasis id="B96BEAFDE279FF8357A56328FEE6F984" bold="true" box="[151,341,1553,1578]" pageId="8" pageNumber="579">FIGURES 21–26.</emphasis>
|
||
<taxonomicName id="4C1F4D6CE279FF8356696328FDCEF984" authority="Jaloszynski" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[347,637,1553,1578]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="occisor" status="sp. nov.">
|
||
<emphasis id="B96BEAFDE279FF8356696328FDB2F987" box="[347,513,1553,1577]" italics="true" pageId="8" pageNumber="579">Clidicus occisor</emphasis>
|
||
Jałoszyński
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2585786E279FF8355B1632BFD09F987" box="[643,698,1554,1577]" pageId="8" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
|
||
Habitus of male (21); head of male in dorsal view (22), right antenna of male in dorsal view (23); aedeagus in ventral (24) and lateral (25) views; ‘sperm pump’ (26).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA036EFE279FF8357F56345FCDDF938" blockId="8.[151,1437,1660,2010]" box="[199,878,1660,1686]" pageId="8" pageNumber="579">Labrum with straight anterior margin, lacking median notch.</paragraph>
|
||
<paragraph id="8BA036EFE279FF8357F56399FA9FF83C" blockId="8.[151,1437,1660,2010]" pageId="8" pageNumber="579">
|
||
Pronotum with subpentagonal disc and short posterior ‘collar’, broadest near anterior fourth;
|
||
<collectionCode id="ED0EAE2AE279FF8353E96399FB48F914" box="[1243,1275,1696,1722]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="579" type="Museum">PL</collectionCode>
|
||
<quantity id="4CE79B0AE279FF8352336399FAD4F914" box="[1281,1383,1696,1722]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="8" pageNumber="579" unit="mm" value="1.28">1.28 mm</quantity>
|
||
,
|
||
<collectionCode id="ED0EAE2AE279FF8352416399FA2FF914" box="[1395,1436,1696,1722]" country="Thailand" name="Paleontological Collections" pageId="8" pageNumber="579">PW</collectionCode>
|
||
<quantity id="4CE79B0AE279FF8357A563FDFF4CF970" box="[151,255,1732,1758]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="8" pageNumber="579" unit="mm" value="1.23">1.23 mm</quantity>
|
||
; ‘collar’ demarcated by a narrow transverse groove with nine small pits; additionally two larger pits are present on each side of pronotum in its constricted portion; base with a well-marked median longitudinal groove at anterior margin of transverse antebasal groove, subequal to 1/4 of the length of pronotal disc. Punctures on disc distinct, much larger and deeper than those on head, almost evenly distributed, separated by spaces shorter than diameters of punctures, sides of pronotum (above each procoxal cavity) indistinctly punctate; setae moderately long, sparse, suberect. Cuticle between punctures on pronotal disc finely shagreened, less glossy than on head.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE279FF8257F562A5FAD1FF55" blockId="8.[151,1437,1660,2010]" lastBlockId="9.[151,1437,153,1043]" lastPageId="9" lastPageNumber="580" pageId="8" pageNumber="579">
|
||
Elytra oval, broadest slightly in front of middle, EL
|
||
<quantity id="4CE79B0AE279FF83542D62A5FC35F818" box="[799,902,1948,1974]" metricMagnitude="-3" metricUnit="m" metricValue="3.08" pageId="8" pageNumber="579" unit="mm" value="3.08">3.08 mm</quantity>
|
||
, EW
|
||
<quantity id="4CE79B0AE279FF8354FA62A2FB9CF818" box="[968,1071,1947,1974]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="8" pageNumber="579" unit="mm" value="1.85">1.85 mm</quantity>
|
||
, EI 1.66; each elytron with five dorsal and one lateral rows of large and deep, densely distributed pits, cuticle between pits finely shagreened, less glossy than that on head, between rows with very fine and sparse setiferous punctures; setae longer and slightly thicker than those on head and pronotum, moderately dense, erect. Apex of each elytron obliquely truncated, with lateral corner rounded and indistinct, adsutural corner strongly projecting posterad, sharp-angled and pointed.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE278FF8257F5643CFE2CFEB1" blockId="9.[151,1437,153,1043]" box="[199,415,261,287]" pageId="9" pageNumber="580">Hind wings absent.</paragraph>
|
||
<paragraph id="8BA036EFE278FF8257F56410FDE0FEED" blockId="9.[151,1437,153,1043]" box="[199,595,297,323]" pageId="9" pageNumber="580">Legs long and slender, unmodified.</paragraph>
|
||
<paragraph id="8BA036EFE278FF8257F56474FED9FDB5" blockId="9.[151,1437,153,1043]" pageId="9" pageNumber="580">
|
||
Aedeagus (
|
||
<figureCitation id="13242A6AE278FF8256756474FE78FEC9" box="[327,459,332,359]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1553,1578]" captionTargetBox="[165,1432,345,1516]" captionTargetId="figure@8.[151,1436,345,1528]" captionTargetPageId="8" captionText="FIGURES 21–26. Clidicus occisor Jałoszyński sp. n. Habitus of male (21); head of male in dorsal view (22), right antenna of male in dorsal view (23); aedeagus in ventral (24) and lateral (25) views; ‘sperm pump’ (26)." httpUri="https://zenodo.org/record/3376158/files/figure.png" pageId="9" pageNumber="580">Figs 24–25</figureCitation>
|
||
) strongly elongate but moderately slender, AeL
|
||
<quantity id="4CE79B0AE278FF8254C86474FBD0FEC9" box="[1018,1123,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="9" pageNumber="580" unit="mm" value="1.23">1.23 mm</quantity>
|
||
; parameres of equal length and shape, short, in ventral view lateral margin of each broadly and deeply emarginate in subapical region, apices distinctly divergent distad, rounded, in lateral view parameres in subapical region not expanded dorsad; flagellum differentiated from ductus ejaculatorius by a rapid broadening between proximal third and middle of median lobe. ‘Sperm pump’ (
|
||
<figureCitation id="13242A6AE278FF82567764E4FE25FE56" box="[325,406,477,504]" captionStart="FIGURES 21–26" captionStartId="8.[151,264,1553,1578]" captionTargetBox="[165,1432,345,1516]" captionTargetId="figure@8.[151,1436,345,1528]" captionTargetPageId="8" captionText="FIGURES 21–26. Clidicus occisor Jałoszyński sp. n. Habitus of male (21); head of male in dorsal view (22), right antenna of male in dorsal view (23); aedeagus in ventral (24) and lateral (25) views; ‘sperm pump’ (26)." httpUri="https://zenodo.org/record/3376158/files/figure.png" pageId="9" pageNumber="580">Fig. 26</figureCitation>
|
||
)
|
||
<quantity id="4CE79B0AE278FF82569764E4FDBBFE59" box="[421,520,477,503]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="580" unit="mm" value="0.68">0.68 mm</quantity>
|
||
in length, with two large ‘funnels’, distal funnel much shorter and slightly broader than proximal one.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE278FF8257F5671CFE11FD91" blockId="9.[151,1437,153,1043]" box="[199,418,549,575]" pageId="9" pageNumber="580">
|
||
<emphasis id="B96BEAFDE278FF8257F5671CFEADFD91" bold="true" box="[199,286,549,575]" pageId="9" pageNumber="580">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE278FF8257F56771FDFEFDCD" blockId="9.[151,1437,153,1043]" box="[199,589,584,611]" pageId="9" pageNumber="580">
|
||
<emphasis id="B96BEAFDE278FF8257F56771FEEAFDCD" bold="true" box="[199,345,584,611]" pageId="9" pageNumber="580">Distribution</emphasis>
|
||
. Southern Mindanao.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE278FF8257F56754FDB0FD05" blockId="9.[151,1437,153,1043]" pageId="9" pageNumber="580">
|
||
<emphasis id="B96BEAFDE278FF8257F56754FEFDFD29" bold="true" box="[199,334,621,647]" pageId="9" pageNumber="580">Etymology.</emphasis>
|
||
The Latin noun in apposition
|
||
<taxonomicName id="4C1F4D6CE278FF8255926757FD41FD29" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[672,754,622,647]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="occisor">
|
||
<emphasis id="B96BEAFDE278FF8255926757FD41FD29" box="[672,754,622,647]" italics="true" pageId="9" pageNumber="580">occisor</emphasis>
|
||
</taxonomicName>
|
||
means “a slayer, murderer”; refers to the typical of all
|
||
<taxonomicName id="4C1F4D6CE278FF8252696754FF0FFD05" authorityName="Laporte" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEAFDE278FF8252696754FF0FFD05" italics="true" pageId="9" pageNumber="580">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
spp. evil-looking mandibles.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE278FF8257F5678CFDBEFBBD" blockId="9.[151,1437,153,1043]" pageId="9" pageNumber="580">
|
||
<emphasis id="B96BEAFDE278FF8257F5678CFE88FD61" bold="true" box="[199,315,693,719]" pageId="9" pageNumber="580">Remarks.</emphasis>
|
||
This species is most similar to
|
||
<taxonomicName id="4C1F4D6CE278FF8255AD678CFCAFFD61" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[671,796,693,719]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B96BEAFDE278FF8255AD678CFCAFFD61" box="[671,796,693,719]" italics="true" pageId="9" pageNumber="580">C. shavrini</emphasis>
|
||
</taxonomicName>
|
||
described below; they both have symmetrical parameres of a similar shape. However, the pronotum of
|
||
<taxonomicName id="4C1F4D6CE278FF8255A867E0FCBEFD5D" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[666,781,729,755]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="occisor">
|
||
<emphasis id="B96BEAFDE278FF8255A867E0FCBEFD5D" box="[666,781,729,755]" italics="true" pageId="9" pageNumber="580">C. occisor</emphasis>
|
||
</taxonomicName>
|
||
is more strongly punctate, the punctures are larger, deeper and denser than those in
|
||
<taxonomicName id="4C1F4D6CE278FF82569F67C4FD99FCB9" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[429,554,765,791]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B96BEAFDE278FF82569F67C4FD99FCB9" box="[429,554,765,791]" italics="true" pageId="9" pageNumber="580">C. shavrini</emphasis>
|
||
</taxonomicName>
|
||
, and the longitudinal median groove is present (absent in
|
||
<taxonomicName id="4C1F4D6CE278FF82538667C4FA82FCB9" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[1204,1329,765,791]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B96BEAFDE278FF82538667C4FA82FCB9" box="[1204,1329,765,791]" italics="true" pageId="9" pageNumber="580">C. shavrini</emphasis>
|
||
</taxonomicName>
|
||
). The aedeagi, despite superficial similarity, clearly differ: in
|
||
<taxonomicName id="4C1F4D6CE278FF8255D76618FCE4FC95" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[741,855,801,827]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="occisor">
|
||
<emphasis id="B96BEAFDE278FF8255D76618FCE4FC95" box="[741,855,801,827]" italics="true" pageId="9" pageNumber="580">C. occisor</emphasis>
|
||
</taxonomicName>
|
||
the subapical lateral emargination of each paramere is much deeper and consequently the parameral apex is distinctly more slender than that in
|
||
<taxonomicName id="4C1F4D6CE278FF8253AE667CFAA9FCF1" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[1180,1306,837,863]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B96BEAFDE278FF8253AE667CFAA9FCF1" box="[1180,1306,837,863]" italics="true" pageId="9" pageNumber="580">C. shavrini</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, in lateral view, the parameres in
|
||
<taxonomicName id="4C1F4D6CE278FF8255366650FDC4FC2D" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[516,631,873,899]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="occisor">
|
||
<emphasis id="B96BEAFDE278FF8255366650FDC4FC2D" box="[516,631,873,899]" italics="true" pageId="9" pageNumber="580">C. occisor</emphasis>
|
||
</taxonomicName>
|
||
are not expanded dorsad, whereas in
|
||
<taxonomicName id="4C1F4D6CE278FF82532C6650FB28FC2D" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[1054,1179,873,899]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B96BEAFDE278FF82532C6650FB28FC2D" box="[1054,1179,873,899]" italics="true" pageId="9" pageNumber="580">C. shavrini</emphasis>
|
||
</taxonomicName>
|
||
parameral apices form a rapid, rounded expansion. Also the aedeagus in lateral view is almost evenly curved in
|
||
<taxonomicName id="4C1F4D6CE278FF82534266B4FB51FC09" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[1136,1250,909,935]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="occisor">
|
||
<emphasis id="B96BEAFDE278FF82534266B4FB51FC09" box="[1136,1250,909,935]" italics="true" pageId="9" pageNumber="580">C. occisor</emphasis>
|
||
</taxonomicName>
|
||
, vs. rapidly bent in the proximal third in
|
||
<taxonomicName id="4C1F4D6CE278FF8256AF6688FDABFC65" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[413,536,945,971]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B96BEAFDE278FF8256AF6688FDABFC65" box="[413,536,945,971]" italics="true" pageId="9" pageNumber="580">C. shavrini</emphasis>
|
||
</taxonomicName>
|
||
. The ‘sperm pump’ differs markedly between these two species; in
|
||
<taxonomicName id="4C1F4D6CE278FF8252306688FAC7FC65" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[1282,1396,945,971]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="occisor">
|
||
<emphasis id="B96BEAFDE278FF8252306688FAC7FC65" box="[1282,1396,945,971]" italics="true" pageId="9" pageNumber="580">C. occisor</emphasis>
|
||
</taxonomicName>
|
||
the proximal funnel is elongate and narrower than the transverse distal funnel, vs. both funnels strongly transverse and of equal diameters in
|
||
<taxonomicName id="4C1F4D6CE278FF8256B866C0FDB5FBBD" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[394,518,1017,1043]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B96BEAFDE278FF8256B866C0FDB5FBBD" box="[394,518,1017,1043]" italics="true" pageId="9" pageNumber="580">C. shavrini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |