503 lines
57 KiB
XML
503 lines
57 KiB
XML
<document id="C6D028462511775F6E51955AB149F1D5" ID-CLB-Dataset="25925" ID-DOI="10.11646/zootaxa.4658.3.8" ID-GBIF-Dataset="65575e2a-b926-423a-a565-82d041ddf1f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3376150" ID-ZooBank="ECF3D8F3-3F41-48DB-B811-6B4265404EE6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566576285773" checkinUser="plazi" docAuthor="Jałoszyński, Paweł, Cheng, Zhi-Fei, Brajczewska, Nastazja & Yin, Zi-Wei" docDate="2019" docId="03B687F9E274FF8C57A561B0FD64FB0D" docLanguage="en" docName="zootaxa.4658.3.8.pdf" docOrigin="Zootaxa 4658 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clidicus kalis Jałoszyński & Cheng & Brajczewska & Yin 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="578" masterDocId="FF8FFF81E271FF8B57326539FFB3FFAE" masterDocTitle="Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="584" masterPageNumber="571" pageNumber="576" updateTime="1698752449993" updateUser="ExternalLinkService">
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<mods:title id="FB818665DD6A6CC8EA7AC0FC1DD5C8F2">Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="59F22DDF62E03560AB2F5E11811E2E7F">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="2DB476E9DE9A7D3A99075EC8CACE7228">Cheng, Zhi-Fei</mods:namePart>
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<mods:namePart id="4C0EF11EDE8630E0DEA3A1EC88C39F80">Brajczewska, Nastazja</mods:namePart>
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<mods:namePart id="6771483FEAD5E908810FFD0585BEDF97">Yin, Zi-Wei</mods:namePart>
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<mods:title id="AF98CB4C34A904FBA77AE28E78B0977D">Zootaxa</mods:title>
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<mods:date id="6F74B3A7FF48F1D611E45FBB3C50D73C">2019</mods:date>
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<mods:number id="869FE48FE1BE9DF747F309202102EFC3">2019-08-23</mods:number>
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<mods:number id="5884A66E164753BD4DC4A36DCDAD58C6">4658</mods:number>
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<taxonomicName id="4C1F4D6CE274FF8E57A561B0FD36FB0D" ID-CoL="8XXT7" authority="Jaloszynski, Cheng & Yin" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[151,645,1160,1188]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="kalis" status="sp. nov.">
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<emphasis id="B96BEAFDE274FF8E57A561B0FE88FB0A" bold="true" box="[151,315,1161,1188]" italics="true" pageId="5" pageNumber="576">Clidicus kalis</emphasis>
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Jałoszyński, Cheng & Yin
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(
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<figureCitation id="13242A6AE274FF8E57AD6194FEACFB69" box="[159,287,1197,1223]" captionStart="FIGURES 14–20" captionStartId="6.[151,264,1277,1302]" captionTargetBox="[162,1430,181,1246]" captionTargetId="figure@6.[151,1436,181,1252]" captionTargetPageId="6" captionText="FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20)." httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="5" pageNumber="576">Figs 14–20</figureCitation>
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)
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</paragraph>
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<emphasis id="B96BEAFDE274FF8E57A561CDFEEDFAA1" bold="true" box="[151,350,1268,1295]" pageId="5" pageNumber="576">Material studied</emphasis>
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.
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<typeStatus id="54A4884DE274FF8E565F61CDFDA3FAA1" box="[365,528,1268,1295]" pageId="5" pageNumber="576" type="holotype">HOLOTYPE</typeStatus>
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:
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:
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<collectingCountry id="F308767FE274FF8E557061CDFD46FAA1" box="[578,757,1268,1295]" name="Philippines" pageId="5" pageNumber="576">PHILIPPINES</collectingCountry>
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</emphasis>
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, with two labels: “
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/
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/ Min- danao, prov.
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<collectingRegion id="49DBF80DE274FF8E561A6020FEC2FA9D" box="[296,369,1305,1331]" country="Philippines" name="Davao" pageId="5" pageNumber="576">Davao</collectingRegion>
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der Sul /
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<location id="8EC06034E274FF8E56ED6020FDE4FA9D" LSID="urn:lsid:plazi:treatment:03B687F9E274FF8C57A561B0FD64FB0D:8EC06034E274FF8E56ED6020FDE4FA9D" box="[479,599,1305,1331]" country="Philippines" latitude="6.9166665" longLatPrecision="1303" longitude="125.26667" name="Kapatagan" pageId="5" pageNumber="576" stateProvince="Davao">Kapatagan</location>
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/
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<emphasis id="B96BEAFDE274FF8E555F6023FD75FA9D" box="[621,710,1305,1331]" italics="true" pageId="5" pageNumber="576">
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<date id="FFA1102FE274FF8E555F6023FD75FA9D" box="[621,710,1305,1331]" pageId="5" pageNumber="576" value="2018-04">
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<collectingDate id="EFE5E9C7E274FF8E555F6023FD75FA9D" box="[621,710,1305,1331]" pageId="5" pageNumber="576" value="2018-04">IV 2018</collectingDate>
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</date>
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</emphasis>
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/ leg.
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<collectorName id="26EA5339E274FF8E543F6020FC79FA9D" box="[781,970,1305,1331]" pageId="5" pageNumber="576">Ismael Lumawig</collectorName>
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(local coll.)” [white, printed], “
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<taxonomicName id="4C1F4D6CE274FF8E52026020FA3EFA9D" authorityName="Laporte" authorityYear="1832" box="[1328,1421,1305,1331]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEAFDE274FF8E52026020FA3EFA9D" box="[1328,1421,1305,1331]" italics="true" pageId="5" pageNumber="576">Clidicus</emphasis>
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</taxonomicName>
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/
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<emphasis id="B96BEAFDE274FF8E57A56004FF7FFAF9" box="[151,204,1341,1367]" italics="true" pageId="5" pageNumber="576">kalis</emphasis>
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Jałoszyński
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<emphasis id="B96BEAFDE274FF8E56646006FE3EFAF9" box="[342,397,1343,1367]" italics="true" pageId="5" pageNumber="576">et al.</emphasis>
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</taxonomicName>
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/
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<typeStatus id="54A4884DE274FF8E56916004FDFEFAF9" box="[419,589,1341,1367]" pageId="5" pageNumber="576" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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)
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</materialsCitation>
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.
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<materialsCitation id="3B773CB2E274FF8E54596005FE42FAD5" ID-GBIF-Occurrence="2416598861" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="USMB, SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="PHILIPPINES" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="49" specimenCount-female="30" specimenCount-male="19" stateProvince="Davao" typeStatus="holotype">
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<emphasis id="B96BEAFDE274FF8E54596005FBB9FAF9" bold="true" box="[875,1034,1340,1367]" pageId="5" pageNumber="576">PAPATYPES</emphasis>
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(
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,
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):
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<collectingCountry id="F308767FE274FF8E53D26005FA24FAF9" box="[1248,1431,1340,1367]" name="Philippines" pageId="5" pageNumber="576">PHILIPPINES</collectingCountry>
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:
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</emphasis>
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, same data as for holotype
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</materialsCitation>
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;
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<materialsCitation id="3B773CB2E274FF8E56CE6058FC3FFA31" ID-GBIF-Occurrence="2416598854" collectingDate="2018-03" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="USMB, SNUC" collectorName="Ismael Lumawig" country="Philippines" latitude="6.9166665" location="PHILIPPINES" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Davao" typeStatus="holotype">
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<specimenCount id="9D19FD66E274FF8E56CE6058FD9BFAD5" box="[508,552,1377,1403]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="9D19FD66E274FF8E55066058FDC7FAD5" box="[564,628,1377,1403]" pageId="5" pageNumber="576" type="female">2 ♀♀</specimenCount>
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, “
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<collectingCountry id="F308767FE274FF8E55B86058FC81FAD5" box="[650,818,1377,1403]" name="Philippines" pageId="5" pageNumber="576">PHILIPPINES</collectingCountry>
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/
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/ Mindanao, prov. Lanao der Sul / Wao /
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<emphasis id="B96BEAFDE274FF8E57D260BFFE88FA31" box="[224,315,1413,1439]" italics="true" pageId="5" pageNumber="576">
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<date id="FFA1102FE274FF8E57D260BFFE88FA31" box="[224,315,1413,1439]" pageId="5" pageNumber="576" value="2018-03">
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<collectingDate id="EFE5E9C7E274FF8E57D260BFFE88FA31" box="[224,315,1413,1439]" pageId="5" pageNumber="576" value="2018-03">III 2018</collectingDate>
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</date>
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</emphasis>
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/ leg.
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<collectorName id="26EA5339E274FF8E56B060BCFD8DFA31" box="[386,574,1413,1439]" pageId="5" pageNumber="576">Ismael Lumawig</collectorName>
|
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(local coll.)” [white, printed]
|
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</materialsCitation>
|
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;
|
||
<materialsCitation id="3B773CB2E274FF8E54AA60BCFAADFA31" ID-GBIF-Occurrence="2416598853" box="[920,1310,1413,1439]" collectingDate="2018-11" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="USMB, SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="1" specimenCount-female="58" specimenCount-male="1" stateProvince="Davao" typeStatus="holotype">
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<specimenCount id="9D19FD66E274FF8E54AA60BCFC77FA31" box="[920,964,1413,1439]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
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, same data except for
|
||
<date id="FFA1102FE274FF8E538D60BCFAADFA31" box="[1215,1310,1413,1439]" pageId="5" pageNumber="576" value="2018-11">
|
||
<collectingDate id="EFE5E9C7E274FF8E538D60BCFAADFA31" box="[1215,1310,1413,1439]" pageId="5" pageNumber="576" value="2018-11">XI 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E521860BCFE1AFA6D" ID-GBIF-Occurrence="2416598855" collectingDate="2018-09" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="USMB, SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="1" specimenCount-female="1" specimenCount-male="39" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E521860BCFAE7FA31" box="[1322,1364,1413,1439]" pageId="5" pageNumber="576" type="female">1 ♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E567A6090FE1AFA6D" box="[328,425,1449,1475]" pageId="5" pageNumber="576" value="2018-09">
|
||
<collectingDate id="EFE5E9C7E274FF8E567A6090FE1AFA6D" box="[328,425,1449,1475]" pageId="5" pageNumber="576" value="2018-09">IX 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E56846090FB2FFA6D" ID-GBIF-Occurrence="2416598867" box="[438,1180,1449,1475]" collectingDate="2019-02" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="USMB" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="1" specimenCount-female="58" specimenCount-male="1" stateProvince="Davao" typeStatus="paratype">
|
||
<specimenCount id="9D19FD66E274FF8E56846090FE50FA6D" box="[438,483,1449,1475]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
|
||
, same data, except for
|
||
<date id="FFA1102FE274FF8E55D86090FC8EFA6D" box="[746,829,1449,1475]" pageId="5" pageNumber="576" value="2019-02">
|
||
<collectingDate id="EFE5E9C7E274FF8E55D86090FC8EFA6D" box="[746,829,1449,1475]" pageId="5" pageNumber="576" value="2019-02">II 2019</collectingDate>
|
||
</date>
|
||
(
|
||
<typeStatus id="54A4884DE274FF8E547F6090FC09FA6D" box="[845,954,1449,1475]" pageId="5" pageNumber="576" type="paratype">paratypes</typeStatus>
|
||
in
|
||
<collectionCode id="ED0EAE2AE274FF8E54D26090FB9CFA6D" box="[992,1071,1449,1475]" pageId="5" pageNumber="576">USMB</collectionCode>
|
||
and cPJ)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E539B6090FDF7FA49" ID-GBIF-Occurrence="2416598863" collectingDate="2017-11" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="1" specimenCount-female="58" specimenCount-male="1" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E539B6090FB65FA6D" box="[1193,1238,1449,1475]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
|
||
, same data as for holotype, except for
|
||
<date id="FFA1102FE274FF8E564D60F4FE6FFA49" box="[383,476,1485,1511]" pageId="5" pageNumber="576" value="2017-11">
|
||
<collectingDate id="EFE5E9C7E274FF8E564D60F4FE6FFA49" box="[383,476,1485,1511]" pageId="5" pageNumber="576" value="2017-11">XI 2017</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E56D960F4FD8FFA49" box="[491,572,1485,1511]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E556260F4FCF8F9A5" ID-GBIF-Occurrence="2416598858" collectingDate="2016-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig" country="Philippines" latitude="6.9166665" location="Lanao del Sur" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Lanao del Sur" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E556260F4FDCFFA49" box="[592,636,1485,1511]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D19FD66E274FF8E55BA60F4FD7BFA49" box="[648,712,1485,1511]" pageId="5" pageNumber="576" type="female">4 ♀♀</specimenCount>
|
||
, “Philippines / Mindanao / prov.
|
||
<collectingRegion id="49DBF80DE274FF8E530F60F4FB68FA49" box="[1085,1243,1485,1511]" country="Philippines" name="Lanao del Sur" pageId="5" pageNumber="576">Lanao del Sur</collectingRegion>
|
||
/ Wao /
|
||
<emphasis id="B96BEAFDE274FF8E520B60F7FA2FFA49" box="[1337,1436,1485,1511]" italics="true" pageId="5" pageNumber="576">
|
||
<date id="FFA1102FE274FF8E520B60F7FA2FFA49" box="[1337,1436,1485,1511]" pageId="5" pageNumber="576" value="2016-12">
|
||
<collectingDate id="EFE5E9C7E274FF8E520B60F7FA2FFA49" box="[1337,1436,1485,1511]" pageId="5" pageNumber="576" value="2016-12">XII 2016</collectingDate>
|
||
</date>
|
||
</emphasis>
|
||
/ leg.
|
||
<collectorName id="26EA5339E274FF8E57EA60C8FE26F9A5" box="[216,405,1521,1547]" pageId="5" pageNumber="576">Ismael Lumawig</collectorName>
|
||
(local coll.)” [white, printed] (
|
||
<collectionCode id="ED0EAE2AE274FF8E55C160C8FCF7F9A5" box="[755,836,1521,1547]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E546560C8FAFAF9A5" ID-GBIF-Occurrence="2416598865" box="[855,1353,1521,1547]" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="2" specimenCount-female="58" specimenCount-male="2" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E546560C8FC2AF9A5" box="[855,921,1521,1547]" pageId="5" pageNumber="576" type="male">2 ♂♂</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E53A560C8FB52F9A5" box="[1175,1249,1521,1547]" pageId="5" pageNumber="576" value="2017-01">I 2017</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E53C260C8FAF2F9A5" box="[1264,1345,1521,1547]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E526760C8FDF8F981" ID-GBIF-Occurrence="2416598866" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="2" specimenCount-female="2" specimenCount-male="39" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E526760C8FA25F9A5" box="[1365,1430,1521,1547]" pageId="5" pageNumber="576" type="female">2 ♀♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E56BC632CFE52F981" box="[398,481,1557,1583]" pageId="5" pageNumber="576" value="2017-02">II 2017</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E56C1632CFDF7F981" box="[499,580,1557,1583]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E556B632CFB16F981" ID-GBIF-Occurrence="2416598870" box="[601,1189,1557,1583]" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E556B632CFD34F981" box="[601,647,1557,1583]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D19FD66E274FF8E55A7632CFD64F981" box="[661,727,1557,1583]" pageId="5" pageNumber="576" type="female">3 ♀♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E54EF632CFB88F981" box="[989,1083,1557,1583]" pageId="5" pageNumber="576" value="2017-04">IV 2017</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E537E632CFB2EF981" box="[1100,1181,1557,1583]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E5380632CFE6CF9FD" ID-GBIF-Occurrence="2416598852" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E5380632CFB53F981" box="[1202,1248,1557,1583]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D19FD66E274FF8E53DD632CFAA8F981" box="[1263,1307,1557,1583]" pageId="5" pageNumber="576" type="female">1 ♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E56226300FEC4F9FD" box="[272,375,1593,1619]" pageId="5" pageNumber="576" value="2017-07">VII 2017</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E56B46300FE64F9FD" box="[390,471,1593,1619]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E56D96300FC5DF9FD" ID-GBIF-Occurrence="2416598859" box="[491,1006,1593,1619]" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="1" specimenCount-female="1" specimenCount-male="39" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E56D96300FDA5F9FD" box="[491,534,1593,1619]" pageId="5" pageNumber="576" type="female">1 ♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E54276300FC36F9FD" box="[789,901,1593,1619]" pageId="5" pageNumber="576" value="2017-08">VIII 2017</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E54A76300FC55F9FD" box="[917,998,1593,1619]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E54C86300FE84F9D9" ID-GBIF-Occurrence="2416598864" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E54C86300FB8FF9FD" box="[1018,1084,1593,1619]" pageId="5" pageNumber="576" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D19FD66E274FF8E537A6300FB3AF9FD" box="[1096,1161,1593,1619]" pageId="5" pageNumber="576" type="female">3 ♀♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E52BA6300FF7CF9D9" pageId="5" pageNumber="576" value="2017-10">X 2017</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E57ED6364FE83F9D9" box="[223,304,1629,1655]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E56716364FC3FF9D9" ID-GBIF-Occurrence="2416598862" box="[323,908,1629,1655]" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="11" specimenCount-female="10" specimenCount-male="1" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E56716364FEC3F9D9" box="[323,368,1629,1655]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D19FD66E274FF8E564E6364FE78F9D9" box="[380,459,1629,1655]" pageId="5" pageNumber="576" type="female">10 ♀♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E55F56364FC97F9D9" box="[711,804,1629,1655]" pageId="5" pageNumber="576" value="2018-06">VI 2018</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E54066364FC36F9D9" box="[820,901,1629,1655]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E54AA6364FA2BF9D9" ID-GBIF-Occurrence="2416598857" box="[920,1432,1629,1655]" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="1" specimenCount-female="58" specimenCount-male="1" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E54AA6364FC76F9D9" box="[920,965,1629,1655]" pageId="5" pageNumber="576" type="male">1 ♂</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E53F26364FA83F9D9" box="[1216,1328,1629,1655]" pageId="5" pageNumber="576" value="2018-08">VIII 2018</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E520D6364FA23F9D9" box="[1343,1424,1629,1655]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E57A563B8FD34F935" ID-GBIF-Occurrence="2416598856" box="[151,647,1665,1691]" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig & N. Luzon" country="Philippines" latitude="6.9166665" location="Kapatagan" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="1" specimenCount-female="1" specimenCount-male="39" stateProvince="Davao" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E57A563B8FF71F935" box="[151,194,1665,1691]" pageId="5" pageNumber="576" type="female">1 ♀</specimenCount>
|
||
, same data except for
|
||
<date id="FFA1102FE274FF8E56F363B8FDADF935" box="[449,542,1665,1691]" pageId="5" pageNumber="576" value="2018-11">XI 2018</date>
|
||
(
|
||
<collectionCode id="ED0EAE2AE274FF8E551C63B8FDCCF935" box="[558,639,1665,1691]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E55A163B8FCCCF911" ID-GBIF-Occurrence="2416598860" collectingDate="2014-12" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="Ismael Lumawig" country="Philippines" latitude="6.9166665" location="Bukidnon" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="2" specimenCount-female="58" specimenCount-male="2" stateProvince="Bukidnon" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E55A163B8FD67F935" box="[659,724,1665,1691]" pageId="5" pageNumber="576" type="male">2 ♂♂</specimenCount>
|
||
, “
|
||
<collectingCountry id="F308767FE274FF8E55D863B8FCD8F935" box="[746,875,1665,1691]" name="Philippines" pageId="5" pageNumber="576">Philippines</collectingCountry>
|
||
/ Mindanao / prov.
|
||
<collectingRegion id="49DBF80DE274FF8E537963B8FB0AF935" box="[1099,1209,1665,1691]" country="Philippines" name="Bukidnon" pageId="5" pageNumber="576">Bukidnon</collectingRegion>
|
||
/ Kabanglasan /
|
||
<emphasis id="B96BEAFDE274FF8E524A63BBFF7CF911" italics="true" pageId="5" pageNumber="576">
|
||
<date id="FFA1102FE274FF8E524A63BBFF7CF911" pageId="5" pageNumber="576" value="2014-12">
|
||
<collectingDate id="EFE5E9C7E274FF8E524A63BBFF7CF911" pageId="5" pageNumber="576" value="2014-12">XII 2014</collectingDate>
|
||
</date>
|
||
</emphasis>
|
||
/ leg.
|
||
<collectorName id="26EA5339E274FF8E5626639CFE63F911" box="[276,464,1701,1727]" pageId="5" pageNumber="576">Ismael Lumawig</collectorName>
|
||
(local coll.)” [white, printed] (
|
||
<collectionCode id="ED0EAE2AE274FF8E5414639CFCC4F911" box="[806,887,1701,1727]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773CB2E274FF8E54B8639CFBB8F94D" ID-GBIF-Occurrence="2416598868" collectingDate="2017-01" collectingDateMax="2019-02" collectingDateMin="2014-12" collectionCode="SNUC" collectorName="N. Luzon & Ismael Lumawig" country="Philippines" latitude="6.9166665" location="Nueva Vizcaya" longLatPrecision="1303" longitude="125.26667" pageId="5" pageNumber="576" specimenCount="4" specimenCount-female="58" specimenCount-male="4" stateProvince="Nueva Vizcaya" typeStatus="holotype">
|
||
<specimenCount id="9D19FD66E274FF8E54B8639CFC79F911" box="[906,970,1701,1727]" pageId="5" pageNumber="576" type="male">4 ♂♂</specimenCount>
|
||
, “Philippines /
|
||
<collectorName id="26EA5339E274FF8E5347639CFB6CF911" box="[1141,1247,1701,1727]" pageId="5" pageNumber="576">N. Luzon</collectorName>
|
||
/
|
||
<collectingRegion id="49DBF80DE274FF8E53C1639CFA2FF911" box="[1267,1436,1701,1727]" country="Philippines" name="Nueva Vizcaya" pageId="5" pageNumber="576">Nueva Vizcaya</collectingRegion>
|
||
/ Kayapa /
|
||
<emphasis id="B96BEAFDE274FF8E562363F3FEE9F94D" box="[273,346,1737,1763]" italics="true" pageId="5" pageNumber="576">
|
||
<date id="FFA1102FE274FF8E562363F3FEE9F94D" box="[273,346,1737,1763]" pageId="5" pageNumber="576" value="2017-01">
|
||
<collectingDate id="EFE5E9C7E274FF8E562363F3FEE9F94D" box="[273,346,1737,1763]" pageId="5" pageNumber="576" value="2017-01">I 2017</collectingDate>
|
||
</date>
|
||
</emphasis>
|
||
/ leg.
|
||
<collectorName id="26EA5339E274FF8E56AB63F0FDE6F94D" box="[409,597,1737,1763]" pageId="5" pageNumber="576">Ismael Lumawig</collectorName>
|
||
(local coll.)” [white, printed] (
|
||
<collectionCode id="ED0EAE2AE274FF8E549C63F0FC4CF94D" box="[942,1023,1737,1763]" pageId="5" pageNumber="576">SNUC</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE274FF8E57F563D5FA8DF839" blockId="5.[151,1437,1268,1943]" pageId="5" pageNumber="576">
|
||
<emphasis id="B96BEAFDE274FF8E57F563D5FEF1F8A9" bold="true" box="[199,322,1772,1799]" pageId="5" pageNumber="576">Diagnosis.</emphasis>
|
||
Pronotum with unevenly distributed, inconspicuously small, shallow and sparse punctures (distances between most punctures larger than their diameters), at middle of disc with impunctate or nearly impunctate small irregular oval area or line, cuticle on pronotum between punctures glossy, pronotal disc with a narrow, shallow and diffuse, barely noticeable median longitudinal impression in posterior 1/6–1/4; aedeagus with asymmetrical parameres, both in ventral view distinctly undulated, the right one with its apex indistinctly curved mesad.
|
||
</paragraph>
|
||
<caption id="DF606667E277FF8D57A561C4FAAFFA94" httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,264,1277,1302]" targetBox="[162,1430,181,1246]" targetPageId="6">
|
||
<paragraph id="8BA036EFE277FF8D57A561C4FAAFFA94" blockId="6.[151,1436,1277,1338]" pageId="6" pageNumber="577">
|
||
<emphasis id="B96BEAFDE277FF8D57A561C4FEE7FAB8" bold="true" box="[151,340,1277,1302]" pageId="6" pageNumber="577">FIGURES 14–20.</emphasis>
|
||
<taxonomicName id="4C1F4D6CE277FF8D566861C7FD12FAB8" authority="Jaloszynski et al." authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[346,673,1278,1302]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="kalis" status="sp. nov.">
|
||
<emphasis id="B96BEAFDE277FF8D566861C7FE57FAB8" box="[346,484,1278,1302]" italics="true" pageId="6" pageNumber="577">Clidicus kalis</emphasis>
|
||
Jałoszyński
|
||
<emphasis id="B96BEAFDE277FF8D555E61C6FD12FAB8" box="[620,673,1278,1302]" italics="true" pageId="6" pageNumber="577">et al.</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2585786E277FF8D559561C6FD6DFAB8" box="[679,734,1279,1302]" pageId="6" pageNumber="577" rank="species">sp. n.</taxonomicNameLabel>
|
||
Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA036EFE277FF8D57F56051FB40FA64" blockId="6.[151,1437,1384,2023]" pageId="6" pageNumber="577">
|
||
Description. Body of male (
|
||
<figureCitation id="13242A6AE277FF8D55346051FDEAFA2D" box="[518,601,1384,1411]" captionStart="FIGURES 14–20" captionStartId="6.[151,264,1277,1302]" captionTargetBox="[162,1430,181,1246]" captionTargetId="figure@6.[151,1436,181,1252]" captionTargetPageId="6" captionText="FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20)." httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="6" pageNumber="577">Fig. 14</figureCitation>
|
||
) elongated and slender, head flattened, prothorax and elytra convex; legs and antennae long and slender, BL
|
||
<quantity id="4CE79B0AE277FF8D551760B5FD7AFA09" box="[549,713,1420,1447]" metricMagnitude="-3" metricUnit="m" metricValue="6.645" metricValueMax="6.88" metricValueMin="6.41" pageId="6" pageNumber="577" unit="mm" value="6.645" valueMax="6.88" valueMin="6.41">6.41–6.88 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE277FF8D542C60B5FC35FA09" box="[798,902,1420,1447]" metricMagnitude="-3" metricUnit="m" metricValue="6.63" pageId="6" pageNumber="577" unit="mm" value="6.63">6.63 mm</quantity>
|
||
); cuticle glossy on head and pronotum, slightly less so on elytral disc, pigmentation uniformly reddish brown, setae indistinctly lighter than cuticle.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE277FF8D57F560EDFAD0F8D5" blockId="6.[151,1437,1384,2023]" pageId="6" pageNumber="577">
|
||
Head (
|
||
<figureCitation id="13242A6AE277FF8D562360EDFED2FA41" box="[273,353,1492,1519]" captionStart="FIGURES 14–20" captionStartId="6.[151,264,1277,1302]" captionTargetBox="[162,1430,181,1246]" captionTargetId="figure@6.[151,1436,181,1252]" captionTargetPageId="6" captionText="FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20)." httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="6" pageNumber="577">Fig. 15</figureCitation>
|
||
) broadest near middle of tempora, HL
|
||
<quantity id="4CE79B0AE277FF8D543C60EDFC03FA41" box="[782,944,1492,1519]" metricMagnitude="-3" metricUnit="m" metricValue="1.115" metricValueMax="1.18" metricValueMin="1.05" pageId="6" pageNumber="577" unit="mm" value="1.115" valueMax="1.18" valueMin="1.05">1.05–1.18 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE277FF8D533060ECFBDAFA41" box="[1026,1129,1493,1519]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="6" pageNumber="577" unit="mm" value="1.12">1.12 mm</quantity>
|
||
), HW
|
||
<quantity id="4CE79B0AE277FF8D539D60EDFAE2FA41" box="[1199,1361,1492,1519]" metricMagnitude="-3" metricUnit="m" metricValue="1.4649999999999999" metricValueMax="1.5" metricValueMin="1.43" pageId="6" pageNumber="577" unit="mm" value="1.4649999999999999" valueMax="1.5" valueMin="1.43">1.43–1.50 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE277FF8D57A560C0FF4CF9BD" box="[151,255,1529,1555]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" pageId="6" pageNumber="577" unit="mm" value="1.47">1.47 mm</quantity>
|
||
); tempora in dorsal view approximately 3 times as long as eyes, in anterior third straight and distinctly di- vergent posterad, posteriorly evenly, strongly rounded; vertex with a deep subtriangular posteromedian impression and a pair of shallow pits marking the sites where dorsal tentorial arms are fused with the head wall, sides of vertex convex; frons between antennal insertions nearly flat, vertex and posterior portion of frons with narrow median longitudinal groove, which is more pronounced posteriorly; supraantennal tubercles small but distinctly elevated; compound eyes small, nearly round, in dorsal view indistinctly projecting laterally from the head silhouette, finely faceted. Punctures on sides of vertex and frons distinct but shallow and with diffuse margins, except for groups of several densely distributed punctures near mesal margin of each eye and posterolateral margin of each supraantennal tubercle; posteromedian impression on vertex, median line on frons and vertex, and supraantennal tubercles indis- tinctly punctate or impunctate. Setae long, sparse, suberect to erect. Antennae (
|
||
<figureCitation id="13242A6AE277FF8D53286205FBDFF8F9" box="[1050,1132,1852,1879]" captionStart="FIGURES 14–20" captionStartId="6.[151,264,1277,1302]" captionTargetBox="[162,1430,181,1246]" captionTargetId="figure@6.[151,1436,181,1252]" captionTargetPageId="6" captionText="FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20)." httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="6" pageNumber="577">Fig. 16</figureCitation>
|
||
) slender, all antennomeres elongate (antennomere XI indistinctly), scape slightly longer than head, AnL
|
||
<quantity id="4CE79B0AE277FF8D54CA6258FB28F8D5" box="[1016,1179,1889,1915]" metricMagnitude="-3" metricUnit="m" metricValue="4.085" metricValueMax="4.29" metricValueMin="3.88" pageId="6" pageNumber="577" unit="mm" value="4.085" valueMax="4.29" valueMin="3.88">3.88–4.29 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE277FF8D53DD6258FAE4F8D5" box="[1263,1367,1889,1915]" metricMagnitude="-3" metricUnit="m" metricValue="4.08" pageId="6" pageNumber="577" unit="mm" value="4.08">4.08 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE277FF8D57F562BDFB74F830" blockId="6.[151,1437,1384,2023]" box="[199,1223,1924,1951]" pageId="6" pageNumber="577">Labrum with straight anterior margin bearing an indistinct, barely noticeable median notch.</paragraph>
|
||
<paragraph id="8BA036EFE277FF8C57F56290FA34FEB1" blockId="6.[151,1437,1384,2023]" lastBlockId="7.[151,1437,153,1187]" lastPageId="7" lastPageNumber="578" pageId="6" pageNumber="577">
|
||
Pronotum with subpentagonal disc and short posterior ‘collar’, broadest near anterior third;
|
||
<collectionCode id="ED0EAE2AE277FF8D53E06290FB41F86D" box="[1234,1266,1961,1987]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="6" pageNumber="577" type="Museum">PL</collectionCode>
|
||
<quantity id="4CE79B0AE277FF8D53CB6291FA2EF86D" box="[1273,1437,1960,1987]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" metricValueMax="1.74" metricValueMin="1.5" pageId="6" pageNumber="577" unit="mm" value="1.62" valueMax="1.74" valueMin="1.5">1.50–1.74 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE277FF8D57D662F5FEFFF849" box="[228,332,1996,2023]" metricMagnitude="-3" metricUnit="m" metricValue="1.61" pageId="6" pageNumber="577" unit="mm" value="1.61">1.61 mm</quantity>
|
||
),
|
||
<collectionCode id="ED0EAE2AE277FF8D566D62F4FE3BF849" box="[351,392,1997,2023]" country="Thailand" name="Paleontological Collections" pageId="6" pageNumber="577">PW</collectionCode>
|
||
<quantity id="4CE79B0AE277FF8D56BD62F5FD81F849" box="[399,562,1996,2023]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="6" pageNumber="577" unit="mm" value="1.525" valueMax="1.6" valueMin="1.45">1.45–1.60 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE277FF8D55B762F5FD5EF849" box="[645,749,1996,2023]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="6" pageNumber="577" unit="mm" value="1.52">1.52 mm</quantity>
|
||
); ‘collar’ demarcated by a narrow transverse groove with 6–8 diffuse pits; additionally, two or three larger pits are present on each side of pronotum in its constricted portion. Punctures on disc distinct but small, shallow and diffuse, unevenly distributed, with variable spaces between punctures, but most punctures are separated by spaces wider than their diameters; elongate median area or line nearly impunctate or at least with punctures sparser than on sides; setae similar to those on head but indistinctly thicker.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F56410FB5DFE59" blockId="7.[151,1437,153,1187]" pageId="7" pageNumber="578">
|
||
Elytra approximately shield-shaped, broadest at or indistinctly behind middle, EL
|
||
<quantity id="4CE79B0AE276FF8C53426411FAA7FEED" box="[1136,1300,296,323]" metricMagnitude="-3" metricUnit="m" metricValue="3.86" metricValueMax="4.05" metricValueMin="3.67" pageId="7" pageNumber="578" unit="mm" value="3.86" valueMax="4.05" valueMin="3.67">3.67–4.05 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C52596410FF74FEC9" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="578" unit="mm" value="3.9">3.90 mm</quantity>
|
||
), EW
|
||
<quantity id="4CE79B0AE276FF8C56396475FE1DFEC9" box="[267,430,332,359]" metricMagnitude="-3" metricUnit="m" metricValue="2.275" metricValueMax="2.4" metricValueMin="2.15" pageId="7" pageNumber="578" unit="mm" value="2.275" valueMax="2.4" valueMin="2.15">2.15–2.40 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C55336474FDDBFEC9" box="[513,616,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" pageId="7" pageNumber="578" unit="mm" value="2.28">2.28 mm</quantity>
|
||
), EI 1.61–1.81; each elytron with five dorsal and one lateral rows of large and deep, densely distributed pits, cuticle between pits slightly less glossy than on head and pronotum, between rows with very fine and sparse setiferous punctures. Apex of each elytron obliquely truncated, with moderately distinct, obtuse-angled lateral corner and distinct adsutural corner, which is strongly projecting posterad, sharp-angled and pointed. Setae on elytra distinctly longer and thicker than those on pronotum, suberect to erect.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F56738FE2CFDB5" blockId="7.[151,1437,153,1187]" box="[199,415,513,539]" pageId="7" pageNumber="578">Hind wings absent.</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F5671CFDE0FD91" blockId="7.[151,1437,153,1187]" box="[199,595,549,575]" pageId="7" pageNumber="578">Legs long and slender, unmodified.</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F56770FC41FD5D" blockId="7.[151,1437,153,1187]" pageId="7" pageNumber="578">
|
||
Aedeagus (
|
||
<figureCitation id="13242A6AE276FF8C56746770FE75FDCD" box="[326,454,585,611]" captionStart="FIGURES 14–20" captionStartId="6.[151,264,1277,1302]" captionTargetBox="[162,1430,181,1246]" captionTargetId="figure@6.[151,1436,181,1252]" captionTargetPageId="6" captionText="FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20)." httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="7" pageNumber="578">Figs 17–18</figureCitation>
|
||
) strongly elongate but moderately slender, AeL
|
||
<quantity id="4CE79B0AE276FF8C54D96771FB3CFDCD" box="[1003,1167,584,611]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="7" pageNumber="578" unit="mm" value="1.5" valueMax="1.55" valueMin="1.45">1.45–1.55 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C53D76771FAFDFDCD" box="[1253,1358,584,611]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="7" pageNumber="578" unit="mm" value="1.52">1.52 mm</quantity>
|
||
); para- meres of unequal length and shape, in ventral view both undulated and with similar, slightly curved mesad apices, parameres distinctly divergent distad; flagellum differentiated from ductus ejaculatorius by a rapid broadening in proximal third of median lobe. ‘Sperm pump’ (
|
||
<figureCitation id="13242A6AE276FF8C559D678CFCB1FD61" box="[687,770,693,719]" captionStart="FIGURES 14–20" captionStartId="6.[151,264,1277,1302]" captionTargetBox="[162,1430,181,1246]" captionTargetId="figure@6.[151,1436,181,1252]" captionTargetPageId="6" captionText="FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20)." httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="7" pageNumber="578">Fig. 19</figureCitation>
|
||
)
|
||
<quantity id="4CE79B0AE276FF8C5421678CFC04FD61" box="[787,951,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" metricValueMax="1.08" metricValueMin="0.94" pageId="7" pageNumber="578" unit="mm" value="1.01" valueMax="1.08" valueMin="0.94">0.94–1.08 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C533C678CFBC7FD61" box="[1038,1140,693,719]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" pageId="7" pageNumber="578" unit="mm" value="0.99">0.99 mm</quantity>
|
||
) in length, with two large ‘funnels’, distal funnel much shorter and slightly brooder than proximal one.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F567C4FCFCFC2D" blockId="7.[151,1437,153,1187]" pageId="7" pageNumber="578">
|
||
<emphasis id="B96BEAFDE276FF8C57F567C4FEADFCB9" bold="true" box="[199,286,765,791]" pageId="7" pageNumber="578">Female</emphasis>
|
||
. Externally very similar to male, on average smaller and with shorter antennae; BL
|
||
<quantity id="4CE79B0AE276FF8C53C467C4FA2EFCB9" box="[1270,1437,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="6.625" metricValueMax="6.92" metricValueMin="6.33" pageId="7" pageNumber="578" unit="mm" value="6.625" valueMax="6.92" valueMin="6.33">6.33–6.92 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C57D46619FEE3FC95" box="[230,336,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="6.59" pageId="7" pageNumber="578" unit="mm" value="6.59">6.59 mm</quantity>
|
||
); HL
|
||
<quantity id="4CE79B0AE276FF8C56A06618FD84FC95" box="[402,567,801,827]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" metricValueMax="1.18" metricValueMin="1.1" pageId="7" pageNumber="578" unit="mm" value="1.14" valueMax="1.18" valueMin="1.1">1.10–1.18 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C55BC6619FD4BFC95" box="[654,760,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="578" unit="mm" value="1.15">1.15 mm</quantity>
|
||
), HW
|
||
<quantity id="4CE79B0AE276FF8C54766619FC5AFC95" box="[836,1001,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="1.615" metricValueMax="1.7" metricValueMin="1.53" pageId="7" pageNumber="578" unit="mm" value="1.615" valueMax="1.7" valueMin="1.53">1.53–1.70 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C53726618FB19FC95" box="[1088,1194,801,827]" metricMagnitude="-3" metricUnit="m" metricValue="1.61" pageId="7" pageNumber="578" unit="mm" value="1.61">1.61 mm</quantity>
|
||
), AnL
|
||
<quantity id="4CE79B0AE276FF8C53CA6619FA2EFC95" box="[1272,1437,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="3.815" metricValueMax="4.05" metricValueMin="3.58" pageId="7" pageNumber="578" unit="mm" value="3.815" valueMax="4.05" valueMin="3.58">3.58–4.05 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C57D1667CFEF9FCF1" box="[227,330,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="3.81" pageId="7" pageNumber="578" unit="mm" value="3.81">3.81 mm</quantity>
|
||
);
|
||
<collectionCode id="ED0EAE2AE276FF8C566E667CFECFFCF1" box="[348,380,837,863]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="578" type="Museum">PL</collectionCode>
|
||
<quantity id="4CE79B0AE276FF8C56B0667DFD90FCF1" box="[386,547,836,863]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.74" metricValueMin="1.51" pageId="7" pageNumber="578" unit="mm" value="1.625" valueMax="1.74" valueMin="1.51">1.51–1.74 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C5546667CFD68FCF1" box="[628,731,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="578" unit="mm" value="1.63">1.63 mm</quantity>
|
||
),
|
||
<collectionCode id="ED0EAE2AE276FF8C55DF667CFCA5FCF1" box="[749,790,837,863]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="578">PW</collectionCode>
|
||
<quantity id="4CE79B0AE276FF8C542E667DFC0DFCF1" box="[796,958,836,863]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="7" pageNumber="578" unit="mm" value="1.525" valueMax="1.6" valueMin="1.45">1.45–1.60 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C533D667DFBC5FCF1" box="[1039,1142,836,863]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" pageId="7" pageNumber="578" unit="mm" value="1.51">1.51 mm</quantity>
|
||
); EL
|
||
<quantity id="4CE79B0AE276FF8C539D667CFAE2FCF1" box="[1199,1361,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="3.8649999999999998" metricValueMax="4.1" metricValueMin="3.63" pageId="7" pageNumber="578" unit="mm" value="3.8649999999999998" valueMax="4.1" valueMin="3.63">3.63–4.10 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C57A56650FF4CFC2D" box="[151,255,873,899]" metricMagnitude="-3" metricUnit="m" metricValue="3.81" pageId="7" pageNumber="578" unit="mm" value="3.81">3.81 mm</quantity>
|
||
), EW
|
||
<quantity id="4CE79B0AE276FF8C56766651FE54FC2D" box="[324,487,872,899]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" metricValueMax="2.45" metricValueMin="2.23" pageId="7" pageNumber="578" unit="mm" value="2.34" valueMax="2.45" valueMin="2.23">2.23–2.45 mm</quantity>
|
||
(mean
|
||
<quantity id="4CE79B0AE276FF8C55096650FD10FC2D" box="[571,675,873,899]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="7" pageNumber="578" unit="mm" value="2.36">2.36 mm</quantity>
|
||
), EI 1.52–1.70.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F566B4FC11FC41" blockId="7.[151,1437,153,1187]" pageId="7" pageNumber="578">
|
||
Spermatheca (
|
||
<figureCitation id="13242A6AE276FF8C565A66B4FE09FC09" box="[360,442,909,935]" captionStart="FIGURES 14–20" captionStartId="6.[151,264,1277,1302]" captionTargetBox="[162,1430,181,1246]" captionTargetId="figure@6.[151,1436,181,1252]" captionTargetPageId="6" captionText="FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20)." httpUri="https://zenodo.org/record/3376156/files/figure.png" pageId="7" pageNumber="578">Fig. 20</figureCitation>
|
||
) with ovoid capsular part, elongate, broadened and slightly irregular in shape projection connected to ductus spermathecae, width of capsular part
|
||
<quantity id="4CE79B0AE276FF8C54026689FC14FC65" box="[816,935,944,971]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="578" unit="mm" value="0.125">0.125 mm</quantity>
|
||
; accessory gland with connecting duct con- spicuously long; ductus spermathecae moderately strongly entangled.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F566C1FBB2FBBD" blockId="7.[151,1437,153,1187]" box="[199,1025,1016,1043]" pageId="7" pageNumber="578">
|
||
<emphasis id="B96BEAFDE276FF8C57F566C1FEEAFBBD" bold="true" box="[199,345,1016,1043]" pageId="7" pageNumber="578">Distribution</emphasis>
|
||
. South-central and north-central Mindanao, northern Luzon.
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F56124FDAFFBF2" blockId="7.[151,1437,153,1187]" pageId="7" pageNumber="578">
|
||
<emphasis id="B96BEAFDE276FF8C57F56124FEFDFB99" bold="true" box="[199,334,1053,1079]" pageId="7" pageNumber="578">Etymology.</emphasis>
|
||
The name refers to undulated parameres which resemble the kalis, a traditional Philippine sword (the name is a noun in apposition).
|
||
</paragraph>
|
||
<paragraph id="8BA036EFE276FF8C57F5615CFD64FB0D" blockId="7.[151,1437,153,1187]" pageId="7" pageNumber="578">
|
||
<emphasis id="B96BEAFDE276FF8C57F5615CFE88FBD1" bold="true" box="[199,315,1125,1151]" pageId="7" pageNumber="578">Remarks.</emphasis>
|
||
This species is most similar to
|
||
<taxonomicName id="4C1F4D6CE276FF8C55A5615CFC9EFBD1" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[663,813,1125,1151]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="interfector">
|
||
<emphasis id="B96BEAFDE276FF8C55A5615CFC9EFBD1" box="[663,813,1125,1151]" italics="true" pageId="7" pageNumber="578">C. interfector</emphasis>
|
||
</taxonomicName>
|
||
; see remarks for that species. Also in
|
||
<taxonomicName id="4C1F4D6CE276FF8C53E0615CFA94FBD1" authorityName="Jałoszyński & Cheng & Brajczewska & Yin" authorityYear="2019" box="[1234,1319,1125,1151]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="kalis">
|
||
<emphasis id="B96BEAFDE276FF8C53E0615CFA94FBD1" box="[1234,1319,1125,1151]" italics="true" pageId="7" pageNumber="578">C. kalis</emphasis>
|
||
</taxonomicName>
|
||
the length of parameres is variable between local populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |