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<document id="DFE91C861FFB1B0ECF88CB2AFC93BF92" ID-CLB-Dataset="25925" ID-DOI="10.11646/zootaxa.4658.3.8" ID-GBIF-Dataset="65575e2a-b926-423a-a565-82d041ddf1f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3376150" ID-ZooBank="ECF3D8F3-3F41-48DB-B811-6B4265404EE6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566576285773" checkinUser="plazi" docAuthor="Jałoszyński, Paweł, Cheng, Zhi-Fei, Brajczewska, Nastazja &amp; Yin, Zi-Wei" docDate="2019" docId="03B687F9E270FF8857A56098FDCFFA49" docLanguage="en" docName="zootaxa.4658.3.8.pdf" docOrigin="Zootaxa 4658 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clidicus forceps Jałoszyński &amp; Cheng &amp; Brajczewska &amp; Yin 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="574" masterDocId="FF8FFF81E271FF8B57326539FFB3FFAE" masterDocTitle="Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="584" masterPageNumber="571" pageNumber="572" updateTime="1698752449993" updateUser="ExternalLinkService">
<mods:mods id="4D47CDE36634BD7EB94EC2BAE0C1EF22" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A4336E18EDC52C6234D4151DF9D848B7">
<mods:title id="89FEC93629C0FC6E366B507A466C0C07">Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="F7A983A13954063E59CBA6B2CE5C9D20" type="personal">
<mods:role id="3BCE14BDE8FC731E104C7D631EAD9892">
<mods:roleTerm id="4D015D0A044B72CC1EC37A312E324999">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="185D445F5E81D600F64D6EF909CD5A66">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:name id="91090CFDD358A3E2E0798DEFF650D4C4" type="personal">
<mods:role id="46D3109684ADA2D5CC8799C5AD444F8F">
<mods:roleTerm id="33396C99C4ECB735C8C35A5FF92D73D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D7ABB5A7176ECEED8287ABE3D05DA76">Cheng, Zhi-Fei</mods:namePart>
</mods:name>
<mods:name id="AF9057DDFB5DFAFFE1CD19156207A1E2" type="personal">
<mods:role id="7723D25F00281266ECDABE3A3E0700ED">
<mods:roleTerm id="96D12BE71F87D3F9B8EC25DFC3E88B03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D6E2784AE1D5B2B08433D5B36FDCF64">Brajczewska, Nastazja</mods:namePart>
</mods:name>
<mods:name id="520A7FE700D565C39E0802D029567BD8" type="personal">
<mods:role id="04DC18974F61C2FD08967B0844493141">
<mods:roleTerm id="1E7251E397CC5EACCC95C85CB9C1C6E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97B67F5180576F436B3A8966C41ACF5B">Yin, Zi-Wei</mods:namePart>
</mods:name>
<mods:typeOfResource id="FEA12035EFFA87E10B240A72C999D078">text</mods:typeOfResource>
<mods:relatedItem id="8CCBE29DF94391CF408280D38FC3C59C" type="host">
<mods:titleInfo id="9E11A8D5BE69FAFD8E8B7B9D6A233244">
<mods:title id="8705C21F28D5B97CC17BF16A774740C6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="63FB4764A9781DFC2D94D726CDFC278F">
<mods:date id="FAC51501BA9FD1BC34D0F80AF4165556">2019</mods:date>
<mods:detail id="32862AF68732C6973252DB3F174EB30E" type="pubDate">
<mods:number id="F6265506EE8251FB0FBDDFDB47FFA095">2019-08-23</mods:number>
</mods:detail>
<mods:detail id="8F4DDCAA1714260055F95C2764C42F65" type="volume">
<mods:number id="A900AEDAE6125DF48019795F7D5884C7">4658</mods:number>
</mods:detail>
<mods:detail id="322ECFFEFC4A5E03B459B96232223B48" type="issue">
<mods:number id="49D130AD4CE9D7E1CC799003FD56117C">3</mods:number>
</mods:detail>
<mods:extent id="377CF5E9FD5AE08BC3E03DE8619D3240" unit="page">
<mods:start id="346C101000F2A59437582734B19EE90D">571</mods:start>
<mods:end id="727B8BB9CE4FFA8B9F39345E82E6A317">584</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5F8760179FA3A970526767FAE606B435">journal article</mods:classification>
<mods:identifier id="9D349EBAEA462B4A2D7AB8512CE174E4" type="CLB-Dataset">25925</mods:identifier>
<mods:identifier id="8612D0B81D35304BEEE168125F88F4D1" type="DOI">10.11646/zootaxa.4658.3.8</mods:identifier>
<mods:identifier id="6298FD9641822919F5EA8D747F18ABC0" type="GBIF-Dataset">65575e2a-b926-423a-a565-82d041ddf1f9</mods:identifier>
<mods:identifier id="A16D507BCAC1D52E17F833945D7448C5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="80009087C0EAB460AC5685E81FE1FE18" type="Zenodo-Dep">3376150</mods:identifier>
<mods:identifier id="9168A95F705F1BFDC45DC21579C76E03" type="ZooBank">ECF3D8F3-3F41-48DB-B811-6B4265404EE6</mods:identifier>
</mods:mods>
<treatment id="03B687F9E270FF8857A56098FDCFFA49" ID-GBIF-Taxon="159243639" LSID="urn:lsid:plazi:treatment:03B687F9E270FF8857A56098FDCFFA49" httpUri="http://treatment.plazi.org/id/03B687F9E270FF8857A56098FDCFFA49" lastPageId="3" lastPageNumber="574" pageId="1" pageNumber="572">
<subSubSection id="C3056564E270FF8A57A56098FD5EFA15" box="[151,749,1440,1468]" pageId="1" pageNumber="572" type="nomenclature">
<paragraph id="8BA036EFE270FF8A57A56098FD5EFA15" blockId="1.[151,749,1440,1503]" box="[151,749,1440,1468]" pageId="1" pageNumber="572">
<heading id="D0E88183E270FF8A57A56098FD5EFA15" bold="true" box="[151,749,1440,1468]" fontSize="11" level="1" pageId="1" pageNumber="572" reason="1">
<emphasis id="B96BEAFDE270FF8A57A56098FD5EFA15" bold="true" box="[151,749,1440,1468]" pageId="1" pageNumber="572">
<taxonomicName id="4C1F4D6CE270FF8A57A56098FD10FA15" ID-CoL="8XXT4" authority="Cheng, Yin &amp; Jaloszynski" authorityName="Jałoszyński &amp; Cheng &amp; Brajczewska &amp; Yin" authorityYear="2019" box="[151,675,1440,1468]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="forceps" status="sp. nov.">
<emphasis id="B96BEAFDE270FF8A57A56098FEE9FA15" bold="true" box="[151,346,1441,1468]" italics="true" pageId="1" pageNumber="572">Clidicus forceps</emphasis>
Cheng, Yin &amp; Jałoszyński
</taxonomicName>
<taxonomicNameLabel id="A2585786E270FF8A55996098FD5EFA15" box="[683,749,1441,1467]" pageId="1" pageNumber="572" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3056564E270FF8A57A560FCFEB8FA71" box="[151,267,1477,1503]" pageId="1" pageNumber="572" type="description">
<paragraph id="8BA036EFE270FF8A57A560FCFEB8FA71" blockId="1.[151,749,1440,1503]" box="[151,267,1477,1503]" pageId="1" pageNumber="572">
(
<figureCitation id="13242A6AE270FF8A57AD60FCFEB1FA71" box="[159,258,1477,1503]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1341,1366]" captionTargetBox="[154,1434,186,1300]" captionTargetId="figure@2.[151,1436,181,1316]" captionTargetPageId="2" captionText="FIGURES 1–7. Clidicus forceps Cheng et al. sp. n. Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7)." httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="1" pageNumber="572">Figs 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3056564E270FF8857A5633CFDCFFA49" lastPageId="3" lastPageNumber="574" pageId="1" pageNumber="572" type="materials_examined">
<paragraph id="8BA036EFE270FF8A57A5633CFB6EF8CA" blockId="1.[151,1437,1541,2037]" pageId="1" pageNumber="572">
<emphasis id="B96BEAFDE270FF8A57A5633CFED1F98E" bold="true" box="[151,354,1541,1568]" pageId="1" pageNumber="572">Material studied.</emphasis>
<materialsCitation id="3B773CB2E270FF8A565A633CFE64F9C6" ID-GBIF-Occurrence="2351005917" collectingDate="2017-02" collectionCode="HOLOTY-, SNUC" collectorName="Ismael Lumawig" country="Philippines" location="PHILIPPINES" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B96BEAFDE270FF8A565A633CFDA3F98E" bold="true" box="[360,528,1541,1568]" pageId="1" pageNumber="572">
<typeStatus id="54A4884DE270FF8A565A633CFDB8F98E" box="[360,523,1541,1568]" pageId="1" pageNumber="572" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
<specimenCount id="9D19FD66E270FF8A5527633EFD99F98F" box="[533,554,1543,1569]" pageId="1" pageNumber="572" type="male">♂</specimenCount>
:
<emphasis id="B96BEAFDE270FF8A550B633CFD58F98E" bold="true" box="[569,747,1541,1568]" pageId="1" pageNumber="572">
<collectingCountry id="F308767FE270FF8A550B633CFD58F98E" box="[569,747,1541,1568]" name="Philippines" pageId="1" pageNumber="572">PHILIPPINES</collectingCountry>
</emphasis>
, with two labels: “
<collectingCountry id="F308767FE270FF8A5485633FFBEDF98E" box="[951,1118,1542,1568]" name="Philippines" pageId="1" pageNumber="572">PHILIPPINES</collectingCountry>
/ Mindanao, prov. Lanao der Sur / Wao /
<emphasis id="B96BEAFDE270FF8A56256312FEDBF9EA" box="[279,360,1578,1604]" italics="true" pageId="1" pageNumber="572">
<date id="FFA1102FE270FF8A56256312FEDBF9EA" box="[279,360,1578,1604]" pageId="1" pageNumber="572" value="2017-02">
<collectingDate id="EFE5E9C7E270FF8A56256312FEDBF9EA" box="[279,360,1578,1604]" pageId="1" pageNumber="572" value="2017-02">II 2017</collectingDate>
</date>
</emphasis>
/ leg.
<collectorName id="26EA5339E270FF8A56986313FDD6F9EA" box="[426,613,1578,1604]" pageId="1" pageNumber="572">Ismael Lumawig</collectorName>
(local coll.)” [white, printed], “
<taxonomicName id="4C1F4D6CE270FF8A54F16313FB93F9EA" authorityName="Laporte" authorityYear="1832" box="[963,1056,1578,1604]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAFDE270FF8A54F16313FB93F9EA" box="[963,1056,1578,1604]" italics="true" pageId="1" pageNumber="572">Clidicus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C1F4D6CE270FF8A53016313FAA3F9EA" authority="Cheng et al." authorityName="Cheng" box="[1075,1296,1578,1604]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="forceps">
<emphasis id="B96BEAFDE270FF8A53016313FB37F9EA" box="[1075,1156,1578,1604]" italics="true" pageId="1" pageNumber="572">forceps</emphasis>
Cheng
<emphasis id="B96BEAFDE270FF8A53EB6312FAA3F9EA" box="[1241,1296,1578,1604]" italics="true" pageId="1" pageNumber="572">et al.</emphasis>
</taxonomicName>
/
<collectionCode id="ED0EAE2AE270FF8A52116313FA2FF9EB" box="[1315,1436,1578,1605]" pageId="1" pageNumber="572">HOLOTY-</collectionCode>
PUS” [red, printed] (
<collectionCode id="ED0EAE2AE270FF8A564C6377FE7CF9C6" box="[382,463,1614,1640]" pageId="1" pageNumber="572">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B773CB2E270FF8A56D36374FC5FF9C6" ID-GBIF-Occurrence="2351005937" box="[481,1004,1613,1641]" country="Philippines" location="PHILIPPINES" pageId="1" pageNumber="572" specimenCount="16" specimenCount-female="9" specimenCount-male="7" typeStatus="paratype">
<emphasis id="B96BEAFDE270FF8A56D36374FD37F9C6" bold="true" box="[481,644,1613,1640]" pageId="1" pageNumber="572">
<typeStatus id="54A4884DE270FF8A56D36374FD37F9C6" box="[481,644,1613,1640]" pageId="1" pageNumber="572" type="paratype">PARATYPES</typeStatus>
</emphasis>
(
<specimenCount id="9D19FD66E270FF8A55A16377FD67F9C7" box="[659,724,1614,1641]" pageId="1" pageNumber="572" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D19FD66E270FF8A55ED6377FCADF9C7" box="[735,798,1614,1641]" pageId="1" pageNumber="572" type="female">9 ♀♀</specimenCount>
):
<collectingCountry id="F308767FE270FF8A54076374FC5FF9C6" box="[821,1004,1613,1640]" name="Philippines" pageId="1" pageNumber="572">
<emphasis id="B96BEAFDE270FF8A54076374FC5FF9C6" bold="true" box="[821,1004,1613,1640]" pageId="1" pageNumber="572">PHILIPPINES</emphasis>
</collectingCountry>
</materialsCitation>
<emphasis id="B96BEAFDE270FF8A54DE6374FC42F9C6" bold="true" box="[1004,1009,1613,1640]" pageId="1" pageNumber="572">:</emphasis>
<materialsCitation id="3B773CB2E270FF8A54C56377FA2BF9C6" ID-GBIF-Occurrence="2351005846" box="[1015,1432,1614,1641]" collectingDate="2017-02" collectionCode="HOLOTY-, SNUC" collectorName="Ismael Lumawig" country="Philippines" location="PHILIPPINES" pageId="1" pageNumber="572" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A54C56377FB90F9C7" box="[1015,1059,1614,1641]" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D19FD66E270FF8A531C6377FBDFF9C7" box="[1070,1132,1614,1641]" pageId="1" pageNumber="572" type="female">2 ♀♀</specimenCount>
, same data as for holotype
</materialsCitation>
;
<materialsCitation id="3B773CB2E270FF8A57A5634BFCA1F922" ID-GBIF-Occurrence="2351005952" box="[151,786,1650,1677]" collectingDate="2016-12" collectionCode="SNUC" collectorName="Ismael Lumawig" country="Philippines" location="PHILIPPINES" pageId="1" pageNumber="572" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A57A5634BFF72F923" box="[151,193,1650,1677]" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
, same data as for holotype, except
<date id="FFA1102FE270FF8A5574634AFD18F922" box="[582,683,1650,1677]" pageId="1" pageNumber="572" value="2016-12">
<collectingDate id="EFE5E9C7E270FF8A5574634AFD18F922" box="[582,683,1650,1677]" pageId="1" pageNumber="572" value="2016-12">XII 2016</collectingDate>
</date>
(
<collectionCode id="ED0EAE2AE270FF8A558B634BFCBAF922" box="[697,777,1650,1676]" pageId="1" pageNumber="572">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B773CB2E270FF8A542F634BFA92F922" ID-GBIF-Occurrence="2351005827" box="[797,1313,1650,1677]" collectingDate="2017-10" collectionCode="HOLOTY-, SNUC" collectorName="Ismael Lumawig" country="Philippines" location="PHILIPPINES" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A542F634BFCFBF923" box="[797,840,1650,1677]" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
, same data as for holotype, except
<date id="FFA1102FE270FF8A53F9634AFA92F922" box="[1227,1313,1650,1677]" pageId="1" pageNumber="572" value="2017-10">
<collectingDate id="EFE5E9C7E270FF8A53F9634AFA92F922" box="[1227,1313,1650,1677]" pageId="1" pageNumber="572" value="2017-10">X 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B773CB2E270FF8A521E634BFDA5F91F" ID-GBIF-Occurrence="2351005798" collectingDate="2017-02" collectionCode="VI" collectorName="Ismael Lumawig" country="Philippines" location="PHILIPPINES" pageId="1" pageNumber="572" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A521E634BFAE5F923" box="[1324,1366,1650,1677]" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
, same data as for holotype, except for
<date id="FFA1102FE270FF8A56CB63AEFDA5F91F" box="[505,534,1687,1713]" pageId="1" pageNumber="572" value="2018-06">
<collectionCode id="ED0EAE2AE270FF8A56CB63AEFDA5F91F" box="[505,534,1687,1713]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="572">VI</collectionCode>
</date>
</materialsCitation>
2018 (
<collectionCode id="ED0EAE2AE270FF8A555663AFFD07F91E" box="[612,692,1686,1712]" pageId="1" pageNumber="572">SNUC</collectionCode>
);
<materialsCitation id="3B773CB2E270FF8A55F563AFFB6EF91F" ID-GBIF-Occurrence="2351005884" box="[711,1245,1686,1713]" collectingDate="2017-02" collectionCode="VII" collectorName="Ismael Lumawig" country="Philippines" location="PHILIPPINES" pageId="1" pageNumber="572" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A55F563AFFCB5F91F" box="[711,774,1686,1713]" pageId="1" pageNumber="572" type="female">2 ♀♀</specimenCount>
, same data as for holotype, except for
<date id="FFA1102FE270FF8A538463AEFB6EF91F" box="[1206,1245,1687,1713]" pageId="1" pageNumber="572" value="2018-07">
<collectionCode id="ED0EAE2AE270FF8A538463AEFB6EF91F" box="[1206,1245,1687,1713]" pageId="1" pageNumber="572">VII</collectionCode>
</date>
</materialsCitation>
2018 (
<collectionCode id="ED0EAE2AE270FF8A521863AFFAC9F91E" box="[1322,1402,1686,1712]" pageId="1" pageNumber="572">SNUC</collectionCode>
);
<materialsCitation id="3B773CB2E270FF8A52BC63AFFF4BF956" ID-GBIF-Occurrence="2351005817" collectingDate="2017-01" collectionCode="SNUC" collectorName="Ismael Lumawig" country="Philippines" location="Kalatungan" municipality="Mindanao" pageId="1" pageNumber="572" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bukidnon" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A52BC63AFFF1DF97B" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D19FD66E270FF8A57886383FF57F97B" box="[186,228,1722,1749]" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F308767FE270FF8A57C96383FE11F97A" box="[251,418,1722,1748]" name="Philippines" pageId="1" pageNumber="572">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6BC4AC95E270FF8A568B6383FD9EF97A" box="[441,557,1722,1748]" pageId="1" pageNumber="572">Mindanao</collectingMunicipality>
, prov.
<collectingRegion id="49DBF80DE270FF8A55456383FD56F97A" box="[631,741,1722,1748]" country="Philippines" name="Bukidnon" pageId="1" pageNumber="572">Bukidnon</collectingRegion>
/
<location id="8EC06034E270FF8A55C96383FCCFF97A" LSID="urn:lsid:plazi:treatment:03B687F9E270FF8857A56098FDCFFA49:8EC06034E270FF8A55C96383FCCFF97A" box="[763,892,1722,1748]" country="Philippines" municipality="Mindanao" name="Kalatungan" pageId="1" pageNumber="572" stateProvince="Bukidnon">Kalatungan</location>
/
<emphasis id="B96BEAFDE270FF8A54A36382FC6AF97A" box="[913,985,1722,1748]" italics="true" pageId="1" pageNumber="572">
<date id="FFA1102FE270FF8A54A36382FC6AF97A" box="[913,985,1722,1748]" pageId="1" pageNumber="572" value="2017-01">
<collectingDate id="EFE5E9C7E270FF8A54A36382FC6AF97A" box="[913,985,1722,1748]" pageId="1" pageNumber="572" value="2017-01">I 2017</collectingDate>
</date>
</emphasis>
/ leg.
<collectorName id="26EA5339E270FF8A532D6383FB6DF97A" box="[1055,1246,1722,1748]" pageId="1" pageNumber="572">Ismael Lumawig</collectorName>
” [white, printed] (
<collectionCode id="ED0EAE2AE270FF8A57AD63E7FF5CF956" box="[159,239,1758,1784]" pageId="1" pageNumber="572">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B773CB2E270FF8A563163E7FDADF8B2" ID-GBIF-Occurrence="2351005897" collectingDate="2017-03" collectionCode="SNUC" collectorName="Ismael Lumawig" country="Philippines" location="Kalatungan" municipality="Mindanao" pageId="1" pageNumber="572" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bukidnon" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A563163E7FE9CF957" box="[259,303,1758,1785]" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D19FD66E270FF8A560963E7FED6F957" box="[315,357,1758,1785]" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F308767FE270FF8A564E63E7FD90F956" box="[380,547,1758,1784]" name="Philippines" pageId="1" pageNumber="572">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6BC4AC95E270FF8A550B63E7FD1EF956" box="[569,685,1758,1784]" pageId="1" pageNumber="572">Mindanao</collectingMunicipality>
, prov.
<collectingRegion id="49DBF80DE270FF8A55C563E7FCD6F956" box="[759,869,1758,1784]" country="Philippines" name="Bukidnon" pageId="1" pageNumber="572">Bukidnon</collectingRegion>
/
<location id="8EC06034E270FF8A544963E7FC4FF956" LSID="urn:lsid:plazi:treatment:03B687F9E270FF8857A56098FDCFFA49:8EC06034E270FF8A544963E7FC4FF956" box="[891,1020,1758,1784]" country="Philippines" municipality="Mindanao" name="Kalatungan" pageId="1" pageNumber="572" stateProvince="Bukidnon">Kalatungan</location>
/
<emphasis id="B96BEAFDE270FF8A532363E6FBD8F956" box="[1041,1131,1758,1784]" italics="true" pageId="1" pageNumber="572">
<date id="FFA1102FE270FF8A532363E6FBD8F956" box="[1041,1131,1758,1784]" pageId="1" pageNumber="572" value="2017-03">
<collectingDate id="EFE5E9C7E270FF8A532363E6FBD8F956" box="[1041,1131,1758,1784]" pageId="1" pageNumber="572" value="2017-03">III 2017</collectingDate>
</date>
</emphasis>
/ leg.
<collectorName id="26EA5339E270FF8A538363E7FADEF956" box="[1201,1389,1758,1784]" pageId="1" pageNumber="572">Ismael Lumawig</collectorName>
(local coll.)” [white, printed] (
<collectionCode id="ED0EAE2AE270FF8A56F4623BFDA5F8B2" box="[454,534,1794,1820]" pageId="1" pageNumber="572">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B773CB2E270FF8A551B623BFB6BF8CA" ID-GBIF-Occurrence="2351005864" collectingDate="2019-05-04" collectorName="Shavrin A. V." country="Philippines" county="Mindanao" elevation="900" latitude="7.2764726" location="Pantukan" longLatPrecision="1" longitude="126.17022" municipality="Araibo" pageId="1" pageNumber="572" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Compostela" typeStatus="paratype">
<specimenCount id="9D19FD66E270FF8A551B623BFDDAF8B3" box="[553,617,1794,1821]" pageId="1" pageNumber="572" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D19FD66E270FF8A5547623BFD2DF8B3" box="[629,670,1794,1821]" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F308767FE270FF8A5587623BFCD2F8B2" box="[693,865,1794,1820]" name="Philippines" pageId="1" pageNumber="572">PHILIPPINES</collectingCountry>
:
<collectingCounty id="62C14E63E270FF8A5458623BFC6DF8B2" box="[874,990,1794,1820]" pageId="1" pageNumber="572">Mindanao</collectingCounty>
,
<collectingMunicipality id="6BC4AC95E270FF8A54D5623BFB84F8B2" box="[999,1079,1794,1820]" pageId="1" pageNumber="572">Araibo</collectingMunicipality>
, /
<location id="8EC06034E270FF8A5362623BFB08F8B2" LSID="urn:lsid:plazi:treatment:03B687F9E270FF8857A56098FDCFFA49:8EC06034E270FF8A5362623BFB08F8B2" box="[1104,1211,1794,1820]" country="Philippines" county="Mindanao" latitude="7.2764726" longLatPrecision="1" longitude="126.17022" municipality="Araibo" name="Pantukan" pageId="1" pageNumber="572" stateProvince="Compostela">Pantukan</location>
,
<collectingRegion id="49DBF80DE270FF8A53F7623BFA2BF8B2" box="[1221,1432,1794,1820]" country="Philippines" name="Compostela" pageId="1" pageNumber="572">Compostela Valley</collectingRegion>
,
<quantity id="4CE79B0AE270FF8A57A5621FFF69F8EE" box="[151,218,1830,1856]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="572" unit="m" value="900.0">
<elevation id="0032D1DCE270FF8A57A5621FFF69F8EE" box="[151,218,1830,1856]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="572" unit="m" value="900.0">900m</elevation>
</quantity>
, /
<location id="8EC06034E270FF8A57C0621FFE10F8EF" LSID="urn:lsid:plazi:treatment:03B687F9E270FF8857A56098FDCFFA49:8EC06034E270FF8A57C0621FFE10F8EF" box="[242,419,1830,1857]" country="Philippines" county="Mindanao" latitude="7.2764726" longLatPrecision="1" longitude="126.17022" municipality="Araibo" name="Candalaga Mts." pageId="1" pageNumber="572" stateProvince="Compostela">Candalaga Mts.</location>
,
<geoCoordinate id="EE2B5028E270FF8A569C621CFD8CF8EE" box="[430,575,1829,1856]" degrees="7" direction="north" minutes="16" orientation="latitude" pageId="1" pageNumber="572" precision="1" seconds="35.3" value="7.2764726">7°16’35.3”N</geoCoordinate>
, /
<geoCoordinate id="EE2B5028E270FF8A556B621FFCB0F8EE" box="[601,771,1830,1856]" degrees="126" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="572" precision="1" seconds="12.8" value="126.17022">126°10’12.8”E</geoCoordinate>
,
<date id="FFA1102FE270FF8A543C621CFCC1F8EE" box="[782,882,1829,1856]" pageId="1" pageNumber="572" value="2019-05-04">
<collectingDate id="EFE5E9C7E270FF8A543C621CFCC1F8EE" box="[782,882,1829,1856]" pageId="1" pageNumber="572" value="2019-05-04">4.5.2019</collectingDate>
</date>
, /
<collectorName id="26EA5339E270FF8A54B9621FFBA9F8EE" box="[907,1050,1830,1856]" pageId="1" pageNumber="572">Shavrin A.V.</collectorName>
leg., secondary broad-leaved / for- est, sifted from wet litter near river and / under wet leaves near rocks” [white, printed] (cPJ, cAS)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA036EFE270FF8A57F56254FE41F85A" blockId="1.[151,1437,1541,2037]" pageId="1" pageNumber="572">
<emphasis id="B96BEAFDE270FF8A57F56254FEF1F826" bold="true" box="[199,322,1901,1928]" pageId="1" pageNumber="572">Diagnosis.</emphasis>
Pronotum with evenly distributed, conspicuously deep and dense punctures (distances between most punctures half as long as their diameters), cuticle between punctures glossy, pronotal disc lacking posterior median longitudinal groove; aedeagus with strongly divergent distad, symmetrical subtriangular parameres, each with its very tip slightly curved mesad.
</paragraph>
<caption id="DF606667E273FF8957A56004FADEFAD4" httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="2" pageNumber="573" startId="2.[151,264,1341,1366]" targetBox="[154,1434,186,1300]" targetPageId="2">
<paragraph id="8BA036EFE273FF8957A56004FADEFAD4" blockId="2.[151,1436,1341,1402]" pageId="2" pageNumber="573">
<emphasis id="B96BEAFDE273FF8957A56004FE88FAF8" bold="true" box="[151,315,1341,1366]" pageId="2" pageNumber="573">FIGURES 1–7.</emphasis>
<taxonomicName id="4C1F4D6CE273FF8956736007FDDAFAF8" authority="Cheng et al." authorityName="Jałoszyński &amp; Cheng &amp; Brajczewska &amp; Yin" authorityYear="2019" box="[321,617,1341,1366]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="573" phylum="Arthropoda" rank="species" species="forceps" status="sp. nov.">
<emphasis id="B96BEAFDE273FF8956736007FE55FAF8" box="[321,486,1342,1366]" italics="true" pageId="2" pageNumber="573">Clidicus forceps</emphasis>
Cheng
<emphasis id="B96BEAFDE273FF8955076006FDDAFAF8" box="[565,617,1342,1366]" italics="true" pageId="2" pageNumber="573">et al.</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2585786E273FF8955476007FD1FFAFB" box="[629,684,1342,1365]" pageId="2" pageNumber="573" rank="species">sp. n.</taxonomicNameLabel>
Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7).
</paragraph>
</caption>
<paragraph id="8BA036EFE273FF8957F5609EFD39F9A4" blockId="2.[151,1436,1447,2015]" pageId="2" pageNumber="573">
<emphasis id="B96BEAFDE273FF8957F5609EFEEAFA6C" bold="true" box="[199,345,1447,1474]" pageId="2" pageNumber="573">Description.</emphasis>
Body of male (
<figureCitation id="13242A6AE273FF8955356091FDF4FA6C" box="[519,583,1448,1474]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1341,1366]" captionTargetBox="[154,1434,186,1300]" captionTargetId="figure@2.[151,1436,181,1316]" captionTargetPageId="2" captionText="FIGURES 1–7. Clidicus forceps Cheng et al. sp. n. Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7)." httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="2" pageNumber="573">Fig. 1</figureCitation>
) elongate and slender, head flattened, prothorax and elytra strongly convex; legs and antennae long and slender, BL
<quantity id="4CE79B0AE273FF89557E60F2FD5DFA49" box="[588,750,1483,1511]" metricMagnitude="-3" metricUnit="m" metricValue="5.43" metricValueMax="5.71" metricValueMin="5.15" pageId="2" pageNumber="573" unit="mm" value="5.43" valueMax="5.71" valueMin="5.15">5.15–5.71 mm</quantity>
(mean
<quantity id="4CE79B0AE273FF89540D60F2FC15FA49" box="[831,934,1483,1511]" metricMagnitude="-3" metricUnit="m" metricValue="5.43" pageId="2" pageNumber="573" unit="mm" value="5.43">5.43 mm</quantity>
); cuticle glossy, pigmentation uniformly dark brown, setae indistinctly lighter than cuticle.
</paragraph>
<paragraph id="8BA036EFE273FF8957F5632DFF67F871" blockId="2.[151,1436,1447,2015]" pageId="2" pageNumber="573">
Head (
<figureCitation id="13242A6AE273FF895623632DFEE6F980" box="[273,341,1556,1582]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1341,1366]" captionTargetBox="[154,1434,186,1300]" captionTargetId="figure@2.[151,1436,181,1316]" captionTargetPageId="2" captionText="FIGURES 1–7. Clidicus forceps Cheng et al. sp. n. Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7)." httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="2" pageNumber="573">Fig. 2</figureCitation>
) broadest near middle of tempora, HL
<quantity id="4CE79B0AE273FF89543A632AFC19F981" box="[776,938,1555,1583]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="2" pageNumber="573" unit="mm" value="0.925" valueMax="0.95" valueMin="0.9">0.90–0.95 mm</quantity>
(mean
<quantity id="4CE79B0AE273FF8954CC632DFBD6F981" box="[1022,1125,1556,1583]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="573" unit="mm" value="0.93">0.93 mm</quantity>
), HW
<quantity id="4CE79B0AE273FF89539F632AFAE3F981" box="[1197,1360,1555,1583]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" metricValueMax="1.33" metricValueMin="1.25" pageId="2" pageNumber="573" unit="mm" value="1.29" valueMax="1.33" valueMin="1.25">1.25–1.33 mm</quantity>
(mean
<quantity id="4CE79B0AE273FF8957A56301FEB2F9FD" box="[151,257,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="573" unit="mm" value="1.3">1.30 mm</quantity>
); tempora in dorsal view nearly 3 times as long as eyes, in anterior third nearly straight and indistinctly divergent posterad, posteriorly strongly and evenly rounded; vertex with a deep subtrapezoidal posteromedian im- pression and a pair of indistinct posterior pits marking sites where dorsal tentorial arms are fused with dorsal wall of head. Sides of vertex convex, frons near each antennal insertion slightly impressed, entire frons and vertex with a narrow median longitudinal groove, supraantennal tubercles small but distinctly elevated; clypeus transverse and convex, demarcated from frons by a deep and distinct frontoclypeal groove; compound eyes small, nearly round, strongly convex but in dorsal view not projecting from head silhouette, finely faceted. Punctures on sides of vertex and frons dense, indistinct and shallow and with diffuse margins, mesal margin of each eye and posterolateral margin of each supraanternnal tubercle with an irregular group of dense punctures; supraantennal tubercles virtually impunctate. Setae moderately long, sparse, suberect to erect. Antennae (
<figureCitation id="13242A6AE273FF8954F46245FBB9F838" box="[966,1034,1916,1942]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1341,1366]" captionTargetBox="[154,1434,186,1300]" captionTargetId="figure@2.[151,1436,181,1316]" captionTargetPageId="2" captionText="FIGURES 1–7. Clidicus forceps Cheng et al. sp. n. Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7)." httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="2" pageNumber="573">Fig. 3</figureCitation>
) slender, scape slightly longer than head, antennomeres II–IX elongate (IX indistinctly so), X–XI distinctly transverse; AnL
<quantity id="4CE79B0AE273FF8953446299FAABF815" box="[1142,1304,1952,1979]" metricMagnitude="-3" metricUnit="m" metricValue="3.27" metricValueMax="3.44" metricValueMin="3.1" pageId="2" pageNumber="573" unit="mm" value="3.27" valueMax="3.44" valueMin="3.1">3.10–3.44 mm</quantity>
(mean
<quantity id="4CE79B0AE273FF8952596299FF74F871" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="2" pageNumber="573" unit="mm" value="3.26">3.26 mm</quantity>
).
</paragraph>
<paragraph id="8BA036EFE272FF8857F565A0FB01FF1D" blockId="3.[151,1437,153,1511]" box="[199,1202,153,179]" pageId="3" pageNumber="574">Labrum with jagged anterior margin bearing a shallow subtriangular anteromedian notch.</paragraph>
<paragraph id="8BA036EFE272FF8857F56584FCF5FE25" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
Pronotum with subpentagonal disc and short posterior ‘collar’, broadest near anterior third;
<collectionCode id="ED0EAE2AE272FF8853E06584FB41FF79" box="[1234,1266,189,215]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="574" type="Museum">PL</collectionCode>
<quantity id="4CE79B0AE272FF8853CB6584FA2EFF79" box="[1273,1437,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" metricValueMax="1.4" metricValueMin="1.22" pageId="3" pageNumber="574" unit="mm" value="1.31" valueMax="1.4" valueMin="1.22">1.22–1.40 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8857D765D8FEFEFF55" box="[229,333,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="3" pageNumber="574" unit="mm" value="1.33">1.33 mm</quantity>
),
<collectionCode id="ED0EAE2AE272FF88565365D8FE39FF55" box="[353,394,225,251]" country="Thailand" name="Paleontological Collections" pageId="3" pageNumber="574">PW</collectionCode>
<quantity id="4CE79B0AE272FF8856A065D9FD85FF55" box="[402,566,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.36" metricValueMin="1.25" pageId="3" pageNumber="574" unit="mm" value="1.305" valueMax="1.36" valueMin="1.25">1.25–1.36 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8855B965D8FD40FF55" box="[651,755,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="574" unit="mm" value="1.3">1.30 mm</quantity>
); ‘collar’ demarcated by a narrow transverse groove with 14 irregular and diffuse pits; additionally, three larger pits are present on each side of pronotum in its constricted por- tion. Punctures on disc similar to those on the most densely punctate regions of head, almost evenly distributed, separated by spaces twice as wide as diameters of punctures; setae moderately long, dense, and suberect. Cuticle between punctures on pronotal disc as glossy as that on head.
</paragraph>
<paragraph id="8BA036EFE272FF8857F564ACFEC7FDCD" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
Elytra oval, broadest slightly in front of middle, EL 3.00–
<quantity id="4CE79B0AE272FF88547964ACFC1DFE01" box="[843,942,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="574" unit="mm" value="3.4">3.40 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF88533364ACFBDAFE01" box="[1025,1129,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="3.17" pageId="3" pageNumber="574" unit="mm" value="3.17">3.17 mm</quantity>
), EW
<quantity id="4CE79B0AE272FF88539F64ACFAE3FE01" box="[1197,1360,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="2.025" metricValueMax="2.12" metricValueMin="1.93" pageId="3" pageNumber="574" unit="mm" value="2.025" valueMax="2.12" valueMin="1.93">1.93–2.12 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8857A56480FEB3FE7D" box="[151,256,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.99" pageId="3" pageNumber="574" unit="mm" value="1.99">1.99 mm</quantity>
), EI 1.53–1.64; each elytron with five dorsal and one lateral rows of large and deep, densely distributed pits, cuticle between pits glossy, between rows with very fine and sparse setiferous punctures; setae longer and slightly thicker than those on head and pronotum, moderately dense, suberect. Apex of each elytron obliquely truncated, with lateral corner rounded and indistinct, and with adsutural corner strongly projecting posterad, sharpangled and pointed.
</paragraph>
<paragraph id="8BA036EFE272FF8857F56754FE2CFD29" blockId="3.[151,1437,153,1511]" box="[199,415,621,647]" pageId="3" pageNumber="574">Hind wings absent.</paragraph>
<paragraph id="8BA036EFE272FF8857F567A8FDE0FD05" blockId="3.[151,1437,153,1511]" box="[199,595,657,683]" pageId="3" pageNumber="574">Legs long and slender, unmodified.</paragraph>
<paragraph id="8BA036EFE272FF8857F5678CFED9FC2D" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
Aedeagus (
<figureCitation id="13242A6AE272FF885677678CFE1AFD61" box="[325,425,692,719]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1341,1366]" captionTargetBox="[154,1434,186,1300]" captionTargetId="figure@2.[151,1436,181,1316]" captionTargetPageId="2" captionText="FIGURES 1–7. Clidicus forceps Cheng et al. sp. n. Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7)." httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="3" pageNumber="574">Figs 4–5</figureCitation>
) strongly elongate but moderately slender, in lateral view strongly curved, AeL
<quantity id="4CE79B0AE272FF88521E678CFF70FD5D" metricMagnitude="-3" metricUnit="m" metricValue="1.115" metricValueMax="1.17" metricValueMin="1.06" pageId="3" pageNumber="574" unit="mm" value="1.115" valueMax="1.17" valueMin="1.06">1.06–1.17 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF88562967E0FE36FD5D" box="[283,389,729,755]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="3" pageNumber="574" unit="mm" value="1.12">1.12 mm</quantity>
); parameres long, symmetrical, in ventral view each paramere with subtriangular apex, api- ces strongly divergent distad, but with their very tips slightly curved mesad and rounded; flagellum differentiated from ductus ejaculatorius by a rapid broadening of walls near basal third of median lobe. ‘Sperm pump’ (
<figureCitation id="13242A6AE272FF88527D6618FA27FC92" box="[1359,1428,801,828]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1341,1366]" captionTargetBox="[154,1434,186,1300]" captionTargetId="figure@2.[151,1436,181,1316]" captionTargetPageId="2" captionText="FIGURES 1–7. Clidicus forceps Cheng et al. sp. n. Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7)." httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="3" pageNumber="574">Fig. 6</figureCitation>
)
<quantity id="4CE79B0AE272FF8857A5667CFE89FCF1" box="[151,314,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.7" metricValueMin="7.6" pageId="3" pageNumber="574" unit="mm" value="0.815" valueMax="0.87" valueMin="0.76">0.76–0.87 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8856BC667CFE40FCF1" box="[398,499,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="574" unit="mm" value="0.8">0.80 mm</quantity>
) in length, with two large ‘funnels’, distal funnel much shorter and slightly brooder than proximal one.
</paragraph>
<paragraph id="8BA036EFE272FF8857F566B4FCA5FBBD" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
<emphasis id="B96BEAFDE272FF8857F566B4FE96FC09" bold="true" box="[199,293,909,935]" pageId="3" pageNumber="574">Female.</emphasis>
Externally very similar to male, on average smaller and with broader elytra; BL
<quantity id="4CE79B0AE272FF88539F66B5FAE3FC09" box="[1197,1360,908,935]" metricMagnitude="-3" metricUnit="m" metricValue="5.404999999999999" metricValueMax="5.52" metricValueMin="5.29" pageId="3" pageNumber="574" unit="mm" value="5.404999999999999" valueMax="5.52" valueMin="5.29">5.29–5.52 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8857A56689FEB2FC65" box="[151,257,944,971]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="574" unit="mm" value="5.4">5.40 mm</quantity>
); HL 0.98–1.00 mm (mean
<quantity id="4CE79B0AE272FF8855726688FD1AFC65" box="[576,681,945,971]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="574" unit="mm" value="0.98">0.98 mm</quantity>
), HW
<quantity id="4CE79B0AE272FF8855C76688FC29FC65" box="[757,922,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" metricValueMax="1.44" metricValueMin="1.32" pageId="3" pageNumber="574" unit="mm" value="1.38" valueMax="1.44" valueMin="1.32">1.32–1.44 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8854C36688FBE8FC65" box="[1009,1115,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="3" pageNumber="574" unit="mm" value="1.39">1.39 mm</quantity>
), AnL
<quantity id="4CE79B0AE272FF88539B6688FAFDFC65" box="[1193,1358,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="3.27" metricValueMax="3.44" metricValueMin="3.1" pageId="3" pageNumber="574" unit="mm" value="3.27" valueMax="3.44" valueMin="3.1">3.10–3.44 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8857A566ECFF4CFC41" box="[151,255,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" pageId="3" pageNumber="574" unit="mm" value="3.28">3.28 mm</quantity>
);
<collectionCode id="ED0EAE2AE272FF88562166ECFE80FC41" box="[275,307,981,1007]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="574" type="Museum">PL</collectionCode>
<quantity id="4CE79B0AE272FF88560B66ECFE6FFC41" box="[313,476,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.4" metricValueMin="1.21" pageId="3" pageNumber="574" unit="mm" value="1.305" valueMax="1.4" valueMin="1.21">1.21–1.40 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF88550266ECFD2BFC41" box="[560,664,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="574" unit="mm" value="1.29">1.29 mm</quantity>
),
<collectionCode id="ED0EAE2AE272FF88559866ECFD60FC41" box="[682,723,981,1007]" country="Thailand" name="Paleontological Collections" pageId="3" pageNumber="574">PW</collectionCode>
<quantity id="4CE79B0AE272FF8855E966EDFCCDFC41" box="[731,894,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.36" metricValueMin="1.25" pageId="3" pageNumber="574" unit="mm" value="1.305" valueMax="1.36" valueMin="1.25">1.25–1.36 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8854E366ECFB8AFC41" box="[977,1081,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="574" unit="mm" value="1.31">1.31 mm</quantity>
); EL
<quantity id="4CE79B0AE272FF88534766ECFAABFC41" box="[1141,1304,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="3.115" metricValueMax="3.21" metricValueMin="3.02" pageId="3" pageNumber="574" unit="mm" value="3.115" valueMax="3.21" valueMin="3.02">3.02–3.21 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF88525966ECFF74FBBD" metricMagnitude="-3" metricUnit="m" metricValue="3.12" pageId="3" pageNumber="574" unit="mm" value="3.12">3.12 mm</quantity>
), EW
<quantity id="4CE79B0AE272FF88563E66C0FE1CFBBD" box="[268,431,1017,1043]" metricMagnitude="-3" metricUnit="m" metricValue="2.005" metricValueMax="2.04" metricValueMin="1.97" pageId="3" pageNumber="574" unit="mm" value="2.005" valueMax="2.04" valueMin="1.97">1.97–2.04 mm</quantity>
(mean 2.00 mm), EI 1.51–1.60.
</paragraph>
<paragraph id="8BA036EFE272FF8857F56124FD54FB0D" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
Spermatheca (
<figureCitation id="13242A6AE272FF88565A6124FE1FFB99" box="[360,428,1053,1079]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1341,1366]" captionTargetBox="[154,1434,186,1300]" captionTargetId="figure@2.[151,1436,181,1316]" captionTargetPageId="2" captionText="FIGURES 1–7. Clidicus forceps Cheng et al. sp. n. Habitus of male (1); head of male in dorsal view (2), left antenna of male in dorsal view (3); aedeagus in ventral (4) and lateral (5) views; ‘sperm pump’ (6); female terminalia and spermatheca (7)." httpUri="https://zenodo.org/record/3376152/files/figure.png" pageId="3" pageNumber="574">Fig. 7</figureCitation>
) with oval capsular part slightly flattened at insertion site of ductus spermathecae, which is connected to a projection slightly irregular in shape, width (diameter) of capsular part
<quantity id="4CE79B0AE272FF8853946179FAFFFBF5" box="[1190,1356,1088,1115]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="3" pageNumber="574" unit="mm" value="0.145" valueMax="0.15" valueMin="0.14">0.14–0.15 mm</quantity>
(mean
<quantity id="4CE79B0AE272FF8857A5615DFEBCFBD1" box="[151,271,1124,1151]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="574" unit="mm" value="0.145">0.145 mm</quantity>
); accessory gland with connecting duct conspicuously long; ductus spermathecae moderately strongly entangled; bursa copulatrix elongate, subrectangular.
</paragraph>
<paragraph id="8BA036EFE272FF8857F56195FC85FB69" blockId="3.[151,1437,153,1511]" box="[199,822,1196,1223]" pageId="3" pageNumber="574">
<emphasis id="B96BEAFDE272FF8857F56195FED2FB69" bold="true" box="[199,353,1196,1223]" pageId="3" pageNumber="574">Distribution.</emphasis>
North-central and east-central Mindanao.
</paragraph>
<paragraph id="8BA036EFE272FF8857F561E8FE89FABE" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
<emphasis id="B96BEAFDE272FF8857F561E8FEFDFB45" bold="true" box="[199,334,1233,1259]" pageId="3" pageNumber="574">Etymology.</emphasis>
The name refers to the shape of parameres which resemble forceps (the Latin “
<taxonomicName id="4C1F4D6CE272FF8853E361E8FA91FB45" authorityName="Jałoszyński &amp; Cheng &amp; Brajczewska &amp; Yin" authorityYear="2019" box="[1233,1314,1233,1259]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="forceps">
<emphasis id="B96BEAFDE272FF8853E361E8FA91FB45" box="[1233,1314,1233,1259]" italics="true" pageId="3" pageNumber="574">forceps</emphasis>
</taxonomicName>
” is a noun in apposition).
</paragraph>
<paragraph id="8BA036EFE272FF8857F56020FE63FA31" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
<emphasis id="B96BEAFDE272FF8857F56020FE88FA9D" bold="true" box="[199,315,1305,1331]" pageId="3" pageNumber="574">Remarks.</emphasis>
<taxonomicName id="4C1F4D6CE272FF8856736020FE46FA9D" authorityName="Jałoszyński &amp; Cheng &amp; Brajczewska &amp; Yin" authorityYear="2019" box="[321,501,1305,1331]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="forceps">
<emphasis id="B96BEAFDE272FF8856736020FE46FA9D" box="[321,501,1305,1331]" italics="true" pageId="3" pageNumber="574">Clidicus forceps</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C1F4D6CE272FF88558C6020FCE4FA9D" box="[702,855,1305,1331]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="crocodylus">
<emphasis id="B96BEAFDE272FF88558C6020FD62FA9D" box="[702,721,1305,1331]" italics="true" pageId="3" pageNumber="574">C</emphasis>
.
<emphasis id="B96BEAFDE272FF8855EF6020FCE4FA9D" box="[733,855,1305,1331]" italics="true" pageId="3" pageNumber="574">crocodylus</emphasis>
</taxonomicName>
. However, the aedeagus of
<taxonomicName id="4C1F4D6CE272FF8853B56020FB44FA9D" authorityName="Jałoszyński &amp; Cheng &amp; Brajczewska &amp; Yin" authorityYear="2019" box="[1159,1271,1305,1331]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="forceps">
<emphasis id="B96BEAFDE272FF8853B56020FB44FA9D" box="[1159,1271,1305,1331]" italics="true" pageId="3" pageNumber="574">C. forceps</emphasis>
</taxonomicName>
is less slender, and the parameres have a different shape. The apex of each paramere is distinctly bent mesad, so that there is a distinct angle on the lateral parameral margin in its subapical region. In
<taxonomicName id="4C1F4D6CE272FF8854936058FB8FFAD5" box="[929,1084,1377,1403]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="crocodylus">
<emphasis id="B96BEAFDE272FF8854936058FB8FFAD5" box="[929,1084,1377,1403]" italics="true" pageId="3" pageNumber="574">C. crocodylus</emphasis>
</taxonomicName>
, the parameral apices are more slender and not bent mesad.
</paragraph>
<paragraph id="8BA036EFE272FF8857F56090FDCFFA49" blockId="3.[151,1437,153,1511]" pageId="3" pageNumber="574">
Some adults of this species were collected in a secondary broad-leaved forest, by sifting leaf litter near river and from under wet leaves near rocks (
<figureCitation id="13242A6AE272FF88552960F4FDDCFA49" box="[539,623,1485,1511]" captionStart="FIGURES 34–37" captionStartId="11.[151,264,1944,1969]" captionTargetBox="[151,1436,957,1919]" captionTargetId="figure@11.[151,1436,957,1919]" captionTargetPageId="11" captionText="FIGURES 34–37. Collecting sites of Clidicus forceps Cheng et al. sp. n. (34) and C. shavrini Jałoszyński sp. n. (35–37). Can- dalaga Mts., Compostela Valley (34); Mt. Talomo (35–37)." httpUri="https://zenodo.org/record/3376162/files/figure.png" pageId="3" pageNumber="574">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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