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<document id="7B700A27A0E1D2661B2875C09F2DE7AB" ID-DOI="10.11646/zootaxa.3972.2.5" ID-GBIF-Dataset="2adbbfb9-f544-45b6-9269-748b09f702d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="239574" ID-ZooBank="6A51EB89-0FE5-4F49-B427-E75459D8A3AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461187349600" checkinUser="plazi" docAuthor="Johnson, Robert A. &amp; Cover, Stefan P." docDate="2015" docId="03B687EEFFFB7C62FF17B0D9650AD8AC" docLanguage="en" docName="zt03972p249.pdf" docOrigin="Zootaxa 3972 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pogonomyrmex aterrimus Wheeler 1936, NEW STATUS" docType="treatment" docVersion="8" lastPageNumber="234" masterDocId="FF8FFF96FFF97C61FF80B1016435DF39" masterDocTitle="A taxonomic revision of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae) on Hispaniola" masterLastPageNumber="249" masterPageNumber="231" pageNumber="233" updateTime="1698599853834" updateUser="plazi">
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<mods:title id="45D99EC586C0C6CDCF05E006C4048AC9">A taxonomic revision of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae) on Hispaniola</mods:title>
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<mods:namePart id="46F6E81D16AD0FDEBC98E3AC26358A8C">Johnson, Robert A.</mods:namePart>
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<mods:namePart id="30884FBF46C5E896F619373F66D96FAF">Cover, Stefan P.</mods:namePart>
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<mods:date id="A17CE21DB705C546DFE038B81ABDFBDB">2015</mods:date>
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<treatment id="03B687EEFFFB7C62FF17B0D9650AD8AC" ID-DOI="http://doi.org/10.5281/zenodo.6095460" ID-GBIF-Taxon="119628751" ID-Zenodo-Dep="6095460" LSID="urn:lsid:plazi:treatment:03B687EEFFFB7C62FF17B0D9650AD8AC" httpUri="http://treatment.plazi.org/id/03B687EEFFFB7C62FF17B0D9650AD8AC" lastPageId="3" lastPageNumber="234" pageId="2" pageNumber="233">
<subSubSection id="C3056573FFFB7C63FF17B0D9651BDD2D" pageId="2" pageNumber="233" type="nomenclature">
<paragraph id="8BA036F8FFFB7C63FF17B0D96772DECA" blockId="2.[151,839,472,533]" box="[151,839,472,499]" pageId="2" pageNumber="233">
<heading id="D0E88194FFFB7C63FF17B0D96772DECA" bold="true" box="[151,839,472,499]" fontSize="11" level="1" pageId="2" pageNumber="233" reason="1">
<emphasis id="B96BEAEAFFFB7C63FF17B0D96772DECA" bold="true" box="[151,839,472,499]" pageId="2" pageNumber="233">
<taxonomicName id="4C1F4D7BFFFB7C63FF17B0D966B0DECA" authority="Wheeler 1936" authorityName="Wheeler" authorityYear="1936" box="[151,645,472,499]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="aterrimus" status="stat. nov.">
<emphasis id="B96BEAEAFFFB7C63FF17B0D965E7DECB" bold="true" box="[151,466,472,498]" italics="true" pageId="2" pageNumber="233">Pogonomyrmex aterrimus</emphasis>
<bibRefCitation id="EF8E4B09FFFB7C63FE59B0D866B0DECA" author="Wheeler" box="[473,645,473,499]" pageId="2" pageNumber="233" refString="Wheeler, W. M. (1936) Ants from Hispaniola and Mona Island. Bulletin of the Museum of Comparative Zoology, 80, 195 - 211." type="journal article" year="1936">Wheeler 1936</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A2585791FFFB7C63FD0CB0D86772DECA" box="[652,839,473,499]" pageId="2" pageNumber="233">NEW STATUS</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF17B0FD651BDD2D" blockId="2.[151,839,472,533]" box="[151,302,508,533]" pageId="2" pageNumber="233">
(
<figureCitation id="13242A7DFFFB7C63FF1EB0FD6510DD2D" box="[158,293,508,533]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[151,250,1941,1963]" captionStartId-1="5.[151,250,1930,1952]" captionStartId-2="6.[151,250,1938,1960]" captionTargetBox-0="[382,1172,195,1896]" captionTargetBox-1="[410,1144,193,1893]" captionTargetBox-2="[389,1155,193,1893]" captionTargetId-0="figure@4.[382,1179,195,1896]" captionTargetId-1="figure@5.[410,1150,193,1894]" captionTargetId-2="figure@6.[389,1161,193,1894]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Photograph of Pogonomyrmex aterrimus Wheeler lectotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217240). Photographs by Erin Prado from www. antweb. org." captionText-1="FIGURE 2. Photograph of Pogonomyrmex “ darlingtoni ” Wheeler lectotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217241). Photographs by Erin Prado from www. antweb. org." captionText-2="FIGURE 3. Photograph of Pogonomyrmex aterrimus Wheeler dealate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217248). Photographs by Erin Prado from www. antweb. org." httpUri-0="https://zenodo.org/record/239575/files/figure.png" httpUri-1="https://zenodo.org/record/239576/files/figure.png" httpUri-2="https://zenodo.org/record/239577/files/figure.png" pageId="2" pageNumber="233">Figures 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3056573FFFB7C63FF17B34460E5DC57" pageId="2" pageNumber="233" type="reference_group">
<paragraph id="8BA036F8FFFB7C63FF17B3446682DDAE" blockId="2.[151,1436,580,878]" pageId="2" pageNumber="233">
<treatmentCitationGroup id="AB0F11D6FFFB7C63FF17B3446682DDAE" infered="true" pageId="2" pageNumber="233">
<taxonomicName id="4C1F4D7BFFFB7C63FF17B34467B8DD63" ID-CoL="5KK5K" authority="Wheeler, 1936: 197" authorityName="Wheeler" authorityPageNumber="197" authorityYear="1936" box="[151,909,580,602]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="subSpecies" species="schmitti" subGenus="Ephebomyrmex" subSpecies="aterrimus">
<emphasis id="B96BEAEAFFFB7C63FF17B3446503DD63" box="[151,310,581,602]" italics="true" pageId="2" pageNumber="233">Pogonomyrmex</emphasis>
(
<emphasis id="B96BEAEAFFFB7C63FEC8B34565D0DD63" box="[328,485,580,602]" italics="true" pageId="2" pageNumber="233">Ephebomyrmex</emphasis>
)
<emphasis id="B96BEAEAFFFB7C63FE78B3456680DD63" box="[504,693,580,602]" italics="true" pageId="2" pageNumber="233">schmitti aterrimus</emphasis>
<treatmentCitation id="0ABE10E9FFFB7C63FD3EB34567B8DD63" author="Wheeler" box="[702,909,580,602]" page="197" pageId="2" pageNumber="233" year="1936">
<bibRefCitation id="EF8E4B09FFFB7C63FD3EB345676CDD63" author="Wheeler" box="[702,857,580,602]" pageId="2" pageNumber="233" refString="Wheeler, W. M. (1936) Ants from Hispaniola and Mona Island. Bulletin of the Museum of Comparative Zoology, 80, 195 - 211." type="journal article" year="1936">Wheeler, 1936</bibRefCitation>
: 197
</treatmentCitation>
</taxonomicName>
(worker).
<typeStatus id="54A4885AFFFB7C63FB80B3456068DD63" box="[1024,1117,580,602]" pageId="2" pageNumber="233" type="syntype">Syntypes</typeStatus>
examined:
<specimenCount id="9D19FD71FFFB7C63FB5AB3456176DD63" box="[1242,1347,580,602]" pageId="2" pageNumber="233" type="worker">3 workers</specimenCount>
[MCZ];
<collectingCountry id="F3087668FFFB7C63FF47B362653BDD41" box="[199,270,611,632]" name="Haiti" pageId="2" pageNumber="233">HAITI</collectingCountry>
, between La Visite and Kenscoff,
<quantity id="4CE79B1DFFFB7C63FD0DB363670BDD41" box="[653,830,610,632]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" metricValueMax="2.1336" metricValueMin="1.524" pageId="2" pageNumber="233" unit="ft" value="6000.0" valueMax="7000.0" valueMin="5000.0">5,0007,000 feet</quantity>
(Dr. Darlington leg., autumn 1934) (MCZ worker here designated
<typeStatus id="54A4885AFFFB7C63FEB9B38065E0DDAE" box="[313,469,641,663]" pageId="2" pageNumber="233" type="lectotype">
<emphasis id="B96BEAEAFFFB7C63FEB9B38065E0DDAE" bold="true" box="[313,469,641,663]" pageId="2" pageNumber="233">LECTOTYPE</emphasis>
</typeStatus>
[CASENT0217240]).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF17B3A1609FDD8F" blockId="2.[151,1436,580,878]" box="[151,1194,672,694]" pageId="2" pageNumber="233">
<taxonomicName id="4C1F4D7BFFFB7C63FF17B3A16651DD8F" authority="Wheeler" authorityName="Wheeler" box="[151,612,672,694]" class="Insecta" family="Formicidae" genus="Ephebomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="subSpecies" species="schmitti" subSpecies="aterrimus">
<emphasis id="B96BEAEAFFFB7C63FF17B3A165C1DD8F" box="[151,500,672,694]" italics="true" pageId="2" pageNumber="233">Ephebomyrmex schmitti aterrimus</emphasis>
(Wheeler)
</taxonomicName>
; Kempf, 1972: 106. First combination in
<taxonomicName id="4C1F4D7BFFFB7C63FB87B3A16091DD8F" box="[1031,1188,672,694]" class="Insecta" family="Formicidae" genus="Ephebomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAEAFFFB7C63FB87B3A16091DD8F" box="[1031,1188,672,694]" italics="true" pageId="2" pageNumber="233">Ephebomyrmex</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF17B3C160F4DDED" blockId="2.[151,1436,580,878]" box="[151,1217,702,725]" pageId="2" pageNumber="233">
<taxonomicName id="4C1F4D7BFFFB7C63FF17B3C16661DDED" ID-CoL="5KK5K" authority="Wheeler" authorityName="Wheeler" box="[151,596,702,725]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="subSpecies" species="schmitti" subSpecies="aterrimus">
<emphasis id="B96BEAEAFFFB7C63FF17B3C165C0DDEC" box="[151,501,702,725]" italics="true" pageId="2" pageNumber="233">Pogonomyrmex schmitti aterrimus</emphasis>
Wheeler
</taxonomicName>
; Bolton, 1995: 339. Revived combination in
<taxonomicName id="4C1F4D7BFFFB7C63FB9CB3C1608FDDEC" ID-CoL="6TNK" box="[1052,1210,704,725]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAEAFFFB7C63FB9CB3C1608FDDEC" box="[1052,1210,704,725]" italics="true" pageId="2" pageNumber="233">Pogonomyrmex</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF17B3DF6020DC09" blockId="2.[151,1436,580,878]" pageId="2" pageNumber="233">
<treatmentCitationGroup id="AB0F11D6FFFB7C63FF17B3DF6020DC09" infered="true" pageId="2" pageNumber="233">
<taxonomicName id="4C1F4D7BFFFB7C63FF17B3DF67A0DDCA" ID-CoL="5KK5L" authority="Wheeler, 1936: 197" authorityName="Wheeler" authorityPageNumber="197" authorityYear="1936" box="[151,917,733,755]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="subSpecies" species="schmitti" subGenus="Ephebomyrmex" subSpecies="darlingtoni">
<emphasis id="B96BEAEAFFFB7C63FF17B3DF6503DDCA" box="[151,310,734,755]" italics="true" pageId="2" pageNumber="233">Pogonomyrmex</emphasis>
(
<emphasis id="B96BEAEAFFFB7C63FEC7B3DC65D1DDCA" box="[327,484,733,755]" italics="true" pageId="2" pageNumber="233">Ephebomyrmex</emphasis>
)
<emphasis id="B96BEAEAFFFB7C63FE75B3DC66F5DDCA" box="[501,704,733,755]" italics="true" pageId="2" pageNumber="233">schmitti darlingtoni</emphasis>
<treatmentCitation id="0ABE10E9FFFB7C63FD49B3DC67A0DDCA" author="Wheeler" box="[713,917,733,755]" page="197" pageId="2" pageNumber="233" year="1936">
<bibRefCitation id="EF8E4B09FFFB7C63FD49B3DC6756DDCA" author="Wheeler" box="[713,867,733,755]" pageId="2" pageNumber="233" refString="Wheeler, W. M. (1936) Ants from Hispaniola and Mona Island. Bulletin of the Museum of Comparative Zoology, 80, 195 - 211." type="journal article" year="1936">Wheeler, 1936</bibRefCitation>
: 197
</treatmentCitation>
</taxonomicName>
(worker).
<typeStatus id="54A4885AFFFB7C63FB86B3DC6056DDCA" box="[1030,1123,733,755]" pageId="2" pageNumber="233" type="syntype">Syntypes</typeStatus>
examined:
<specimenCount id="9D19FD71FFFB7C63FB5DB3DC6170DDCA" box="[1245,1349,733,755]" pageId="2" pageNumber="233" type="worker">6 workers</specimenCount>
[MCZ];
<collectingCountry id="F3087668FFFB7C63FF47B3FD653BDC28" box="[199,270,764,785]" name="Haiti" pageId="2" pageNumber="233">HAITI</collectingCountry>
, northeastern foothills of Massif de la Hotte,
<quantity id="4CE79B1DFFFB7C63FD79B3FD679DDC2B" box="[761,936,764,786]" metricMagnitude="2" metricUnit="m" metricValue="9.144" metricValueMax="12.192" metricValueMin="6.096" pageId="2" pageNumber="233" unit="ft" value="3000.0" valueMax="4000.0" valueMin="2000.0">2,0004,000 feet</quantity>
(Dr. Darlington leg.,
<date id="FFA11038FFFB7C63FB12B3FD613CDC2B" box="[1170,1289,764,786]" pageId="2" pageNumber="233" value="2010-10">10 October</date>
, 1934) (MCZ worker here designated
<typeStatus id="54A4885AFFFB7C63FE39B21B6661DC09" box="[441,596,794,816]" pageId="2" pageNumber="233" type="lectotype">
<emphasis id="B96BEAEAFFFB7C63FE39B21B6661DC09" bold="true" box="[441,596,794,816]" pageId="2" pageNumber="233">LECTOTYPE</emphasis>
</typeStatus>
[CASENT0217241]).
<emphasis id="B96BEAEAFFFB7C63FCBCB21B6020DC09" bold="true" box="[828,1045,794,816]" pageId="2" pageNumber="233">NEW SYNONOMY</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF17B238608FDC76" blockId="2.[151,1436,580,878]" box="[151,1210,825,847]" pageId="2" pageNumber="233">
<taxonomicName id="4C1F4D7BFFFB7C63FF17B2386646DC76" authority="Wheeler" authorityName="Wheeler" box="[151,627,825,847]" class="Insecta" family="Formicidae" genus="Ephebomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="subSpecies" species="schmitti" subSpecies="darlingtoni">
<emphasis id="B96BEAEAFFFB7C63FF17B2386636DC76" box="[151,515,825,847]" italics="true" pageId="2" pageNumber="233">Ephebomyrmex schmitti darlingtoni</emphasis>
(Wheeler)
</taxonomicName>
; Kempf, 1972: 106. First combination in
<taxonomicName id="4C1F4D7BFFFB7C63FB96B2386086DC76" box="[1046,1203,825,847]" class="Insecta" family="Formicidae" genus="Ephebomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAEAFFFB7C63FB96B2386086DC76" box="[1046,1203,825,847]" italics="true" pageId="2" pageNumber="233">Ephebomyrmex</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF17B25860E5DC57" blockId="2.[151,1436,580,878]" box="[151,1232,856,878]" pageId="2" pageNumber="233">
<taxonomicName id="4C1F4D7BFFFB7C63FF17B2586656DC57" ID-CoL="5KK5L" authority="Wheeler" authorityName="Wheeler" box="[151,611,856,878]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="subSpecies" species="schmitti" subSpecies="darlingtoni">
<emphasis id="B96BEAEAFFFB7C63FF17B2586630DC57" box="[151,517,856,878]" italics="true" pageId="2" pageNumber="233">Pogonomyrmex schmitti darlingtoni</emphasis>
Wheeler
</taxonomicName>
; Bolton, 1995: 340. Revived combination in
<taxonomicName id="4C1F4D7BFFFB7C63FBACB25860FEDC57" ID-CoL="6TNK" box="[1068,1227,857,878]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAEAFFFB7C63FBACB25860FEDC57" box="[1068,1227,857,878]" italics="true" pageId="2" pageNumber="233">Pogonomyrmex</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3056573FFFB7C62FF17B29B65BDDECC" lastPageId="3" lastPageNumber="234" pageId="2" pageNumber="233" type="description">
<paragraph id="8BA036F8FFFB7C63FF17B29B614BDB7A" blockId="2.[151,1437,922,2027]" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FF17B29B6549DC8A" bold="true" box="[151,380,922,947]" pageId="2" pageNumber="233">Worker. Diagnosis.</emphasis>
Small (HW =
<quantity id="4CE79B1DFFFB7C63FDA8B29B66E5DC8A" box="[552,720,922,947]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" metricValueMax="1.35" metricValueMin="1.11" pageId="2" pageNumber="233" unit="mm" value="1.23" valueMax="1.35" valueMin="1.11">1.111.35 mm</quantity>
), dorsum of mesosoma rugoreticulate to vermiculate; posterior surface of petiolar node viewed from behind and above elongate, clearly longer than wide, petiolar node notably narrower than postpetiole (PW/PPW = 0.780.90); anterior surface of petiolar node partly to completely sculptured, dull to weakly shining; in profile, anterior surface of petiolar node forming a right to weakly obtuse angle with peduncle of petiole; dorsum of postpetiole and first gastral tergum strongly punctate, dull (
<figureCitation id="13242A7DFFFB7C63FA8AB52B615BDB7A" box="[1290,1390,1066,1091]" captionStart="FIGURE 1" captionStartId="4.[151,250,1941,1963]" captionTargetBox="[382,1172,195,1896]" captionTargetId="figure@4.[382,1179,195,1896]" captionTargetPageId="4" captionText="FIGURE 1. Photograph of Pogonomyrmex aterrimus Wheeler lectotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217240). Photographs by Erin Prado from www. antweb. org." httpUri="https://zenodo.org/record/239575/files/figure.png" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FA8AB52B615BDB7A" bold="true" box="[1290,1390,1066,1091]" pageId="2" pageNumber="233">Figure 1</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF47B54F670ADBEA" blockId="2.[151,1437,922,2027]" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FF47B54F6548DB5E" bold="true" box="[199,381,1102,1127]" pageId="2" pageNumber="233">Measurements.</emphasis>
<typeStatus id="54A4885AFFFB7C63FE04B54F65DFDB5E" box="[388,490,1102,1127]" pageId="2" pageNumber="233" type="lectotype">lectotype</typeStatus>
(
<emphasis id="B96BEAEAFFFB7C63FE7BB54E663CDB5F" box="[507,521,1103,1126]" italics="true" pageId="2" pageNumber="233">n</emphasis>
= 10 + 2
<typeStatus id="54A4885AFFFB7C63FDF6B54F6728DB5E" box="[630,797,1102,1127]" pageId="2" pageNumber="233" type="paralectotype">paralectotypes</typeStatus>
). HL 1.29 (1.291.47); HW 1.11 (1.151.33); MOD 0.21 (0.230.25); OMD 0.27 (0.250.34); SL 0.90 (0.911.01); PNW 0.80 (0.810.93); HFL 1.23 (1.171.36); ML 1.50 (1.471.72); PW 0.39 (0.350.45); PPW 0.51 (0.450.53). Indices: SI 81.08 (69.9284.87); CI 86.05 (88.32 95.00); OI 18.92 (18.0520.83); HFI 110.81 (87.97111.57).
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF47B5DE610CDA5A" blockId="2.[151,1437,922,2027]" pageId="2" pageNumber="233">
Measurements for
<emphasis id="B96BEAEAFFFB7C63FE1EB5DE6599DBCF" box="[414,428,1247,1270]" italics="true" pageId="2" pageNumber="233">P</emphasis>
. “
<taxonomicName id="4C1F4D7BFFFB7C63FE49B5DF6672DBCF" box="[457,583,1246,1270]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B96BEAEAFFFB7C63FE49B5DF6672DBCF" box="[457,583,1246,1270]" italics="true" pageId="2" pageNumber="233">darlingtoni</emphasis>
</taxonomicName>
”.
<typeStatus id="54A4885AFFFB7C63FDE4B5DF66FEDBCE" box="[612,715,1246,1271]" pageId="2" pageNumber="233" type="lectotype">lectotype</typeStatus>
(
<emphasis id="B96BEAEAFFFB7C63FD5DB5DE66DEDBCF" box="[733,747,1247,1270]" italics="true" pageId="2" pageNumber="233">n</emphasis>
= 5
<typeStatus id="54A4885AFFFB7C63FCA6B5DF67FFDBCE" box="[806,970,1246,1271]" pageId="2" pageNumber="233" type="paralectotype">paralectotypes</typeStatus>
) (see below). HL 1.29 (1.291.44); HW 1.21 (1.131.35); MOD 0.27 (0.250.27); OMD 0.28 (0.270.31); SL 0.96 (0.881.06); PNW 0.80 (0.790.86); HFL 1.32 (1.251.46); ML 1.59 (1.581.72); PW 0.37 (0.360.43); PPW 0.45 (0.440.49). Indices: SI 79.34 (77.8879.03); CI 93.80 (87.6094.03); OI 22.31 (19.8422.12); HFI 109.09 (102.22114.96) (
<figureCitation id="13242A7DFFFB7C63FB44B44B611DDA5A" box="[1220,1320,1354,1379]" captionStart="FIGURE 2" captionStartId="5.[151,250,1930,1952]" captionTargetBox="[410,1144,193,1893]" captionTargetId="figure@5.[410,1150,193,1894]" captionTargetPageId="5" captionText="FIGURE 2. Photograph of Pogonomyrmex “ darlingtoni ” Wheeler lectotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217241). Photographs by Erin Prado from www. antweb. org." httpUri="https://zenodo.org/record/239576/files/figure.png" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FB44B44B611DDA5A" bold="true" box="[1220,1320,1354,1379]" pageId="2" pageNumber="233">Figure 2</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF47B46F6610D92E" blockId="2.[151,1437,922,2027]" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FF47B46F6627DABE" bold="true" box="[199,530,1390,1415]" pageId="2" pageNumber="233">Queen (dealate). Diagnosis.</emphasis>
With caste-specific morphology of the mesosoma related to wing-bearing and presence of ocelli on head. Small (HW =
<quantity id="4CE79B1DFFFB7C63FDF6B49266D5DA92" box="[630,736,1427,1451]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="2" pageNumber="233" unit="mm" value="1.32">1.32 mm</quantity>
), but notably larger than conspecific workers; in dorsal view, petiolar node elongate (width/length &lt;0.85), relatively narrow compared to postpetiole (PW/PPW &lt;0.75); anterior surface of petiolar node moderately sculptured; dorsum of postpetiole and most of first gastral tergum strongly granulate-punctate, dull (
<figureCitation id="13242A7DFFFB7C63FE31B4FF6620D92E" box="[433,533,1534,1559]" captionStart="FIGURE 3" captionStartId="6.[151,250,1938,1960]" captionTargetBox="[389,1155,193,1893]" captionTargetId="figure@6.[389,1161,193,1894]" captionTargetPageId="6" captionText="FIGURE 3. Photograph of Pogonomyrmex aterrimus Wheeler dealate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217248). Photographs by Erin Prado from www. antweb. org." httpUri="https://zenodo.org/record/239577/files/figure.png" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FE31B4FF6620D92E" bold="true" box="[433,533,1534,1559]" pageId="2" pageNumber="233">Figure 3</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF47B72367F5D966" blockId="2.[151,1437,922,2027]" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FF47B7236543D902" bold="true" box="[199,374,1570,1595]" pageId="2" pageNumber="233">Measurements</emphasis>
(
<emphasis id="B96BEAEAFFFB7C63FE06B72265A1D903" box="[390,404,1571,1594]" italics="true" pageId="2" pageNumber="233">n</emphasis>
= 1). HL 1.45; HW 1.32; MOD 0.28; OMD 0.29; SL 1.06; PNW 0.96; HFL 1.36; ML 1.90; PW 0.42; PPW 0.59. Indices: SI 80.30; CI 91.03; OI 21.21; HFI 103.03.
</paragraph>
<paragraph id="8BA036F8FFFB7C63FF47B76B6607D862" blockId="2.[151,1437,922,2027]" pageId="2" pageNumber="233">
<emphasis id="B96BEAEAFFFB7C63FF47B76B656CD9BA" bold="true" box="[199,345,1642,1667]" pageId="2" pageNumber="233">Description.</emphasis>
With caste-specific morphology of the mesosoma related to wing-bearing and presence of ocelli on head. In full-face view, head longer than broad (CI = 91.03), posterior margin flat. Longitudinal rugae on cephalic dorsum prominent, wavy to irregular, area posterolateral to frontal carinae and posterior margin weakly to moderately rugoreticulate; interrugae strongly granulate, dull. Mandible with six teeth, dorsal surface coarsely rugose. Eye relatively small, situated anterior to middle of head. Base of antennal scape weakly to moderately granulate, distal portion strongly granulate, dull. Psammophore poorly-developed, consisting of short hairs scattered across ventral side of head.
</paragraph>
<paragraph id="8BA036F8FFFB7C62FF47B66765C2DEE9" blockId="2.[151,1437,922,2027]" lastBlockId="3.[151,1437,151,1941]" lastPageId="3" lastPageNumber="234" pageId="2" pageNumber="233">
All mesosomal surfaces with subparallel, weakly irregular rugae to rugoreticulate. Pronotal collar rugoreticulate, mesoscutellum and scutellum with irregular longitudinal rugae. Dorsum of propodeum with wavy to irregular transverse rugae, sides rugoreticulate. Propodeum with long, well-developed superior and inferior spines, inferior spines about 0.60.7x the length of superior spines. In profile, petiolar node asymmetrical with anterior surface notably shorter than posterior surface, apex of node subangulate. In dorsal view, petiolar node longer than wide (width/length &lt;0.85), widest near middle, tapering to a rounded to subangulate anterior margin. Anterior surface of petiolar node moderately sculptured; sides and posterior surface rugoreticulate and “lumpy” in appearance, interrugae moderately granulate. In dorsal view, postpetiole widest near posterior margin, tapering to anterior margin, maximum width about equal to length, relatively narrow compared to petiolar node (PW/PPW &lt;0.75), all surfaces strongly granulate, dull. Anterior one-half of first gastral tergum strongly granulate, dull, posterior portion moderately coriarious, weakly shining. Moderately abundant yellowish-brown to brownish hairs on entire body, longest hairs on head and mesosoma approach to slightly exceed MOD. Entire body concolorous dark reddish-brown (
<figureCitation id="13242A7DFFFA7C62FE03B0B665D2DEE9" box="[387,487,439,464]" captionStart="FIGURE 2" captionStartId="5.[151,250,1930,1952]" captionTargetBox="[410,1144,193,1893]" captionTargetId="figure@5.[410,1150,193,1894]" captionTargetPageId="5" captionText="FIGURE 2. Photograph of Pogonomyrmex “ darlingtoni ” Wheeler lectotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217241). Photographs by Erin Prado from www. antweb. org." httpUri="https://zenodo.org/record/239576/files/figure.png" pageId="3" pageNumber="234">
<emphasis id="B96BEAEAFFFA7C62FE03B0B665D2DEE9" bold="true" box="[387,487,439,464]" pageId="3" pageNumber="234">Figure 2</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8BA036F8FFFA7C62FF47B0DD65BDDECC" blockId="3.[151,1437,151,1941]" box="[199,392,476,501]" pageId="3" pageNumber="234">
<emphasis id="B96BEAEAFFFA7C62FF47B0DD653EDECC" bold="true" box="[199,267,476,501]" pageId="3" pageNumber="234">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3056573FFFA7C62FF47B0FE65F7DD91" pageId="3" pageNumber="234" type="materials_examined">
<paragraph id="8BA036F8FFFA7C62FF47B0FE65F7DD91" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="234">
<emphasis id="B96BEAEAFFFA7C62FF47B0FE67C6DD2E" bold="true" box="[199,1011,511,536]" pageId="3" pageNumber="234">
Additional material examined.
<collectingCountry id="F3087668FFFA7C62FDBEB0FE67B2DD21" box="[574,903,511,536]" name="Dominican Republic" pageId="3" pageNumber="234">DOMINICAN REPUBLIC</collectingCountry>
:
<emphasis id="B96BEAEAFFFA7C62FC14B30167C6DD2E" bold="true" box="[916,1011,512,535]" italics="true" pageId="3" pageNumber="234">La Vega</emphasis>
</emphasis>
: Constanza, 3,0004,000,
<date id="FFA11038FFFA7C62FAACB30161A9DD21" box="[1324,1436,512,536]" pageId="3" pageNumber="234" value="1938-08">Aug 1938</date>
(PJ Darlington;
<collectionCode id="ED0EAE3DFFFA7C62FED0B32565A7DD05" LSID="urn:lsid:biocol.org:col:33791" box="[336,402,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="234">MCZ</collectionCode>
).
<emphasis id="B96BEAEAFFFA7C62FE27B3226612DD05" bold="true" box="[423,551,547,572]" italics="true" pageId="3" pageNumber="234">Pedernales</emphasis>
: Parque Nacional Sierra Bahoruco,
<quantity id="4CE79B1DFFFA7C62FC4CB325602DDD05" box="[972,1048,548,573]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="234" unit="m" value="760.0">760 m</quantity>
,
<date id="FFA11038FFFA7C62FBA7B325608FDD05" box="[1063,1210,548,572]" pageId="3" pageNumber="234" value="2001-09-02">Sept 2, 2001</date>
(AL Wild;
<collectionCode id="ED0EAE3DFFFA7C62FAC7B32561A5DD05" box="[1351,1424,548,572]" pageId="3" pageNumber="234">RAJC</collectionCode>
); Sierra Bahoruco,
<quantity id="4CE79B1DFFFA7C62FEDDB3496581DD66" box="[349,436,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.7899999999999998" pageId="3" pageNumber="234" unit="m" value="1790.0">1790 m</quantity>
&amp;
<quantity id="4CE79B1DFFFA7C62FE59B3496604DD66" box="[473,561,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="234" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FFA11038FFFA7C62FDBEB34966EDDD59" box="[574,728,584,608]" pageId="3" pageNumber="234" value="2014-03-29">Mar 29, 2014</date>
(D Lubertazzi;
<collectionCode id="ED0EAE3DFFFA7C62FC0DB34967F9DD59" LSID="urn:lsid:biocol.org:col:33791" box="[909,972,584,608]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="234">MCZ</collectionCode>
,
<collectionCode id="ED0EAE3DFFFA7C62FC59B3496016DD59" box="[985,1059,584,608]" pageId="3" pageNumber="234">RAJC</collectionCode>
);
<quantity id="4CE79B1DFFFA7C62FBB7B346604ADD59" box="[1079,1151,583,608]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="234" unit="km" value="16.0">16 km</quantity>
ENE Pedernales,
<quantity id="4CE79B1DFFFA7C62FACDB34961A2DD66" box="[1357,1431,584,608]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="234" unit="m" value="800.0">800 m</quantity>
,
<date id="FFA11038FFFA7C62FF17B36D6505DDBD" box="[151,304,620,644]" pageId="3" pageNumber="234" value="1992-09-10">Sept 10, 1992</date>
(PS Ward;
<collectionCode id="ED0EAE3DFFFA7C62FE30B36D65DADDBD" LSID="urn:lsid:biocol.org:col:33791" box="[432,495,620,644]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="234">MCZ</collectionCode>
,
<collectionCode id="ED0EAE3DFFFA7C62FE7AB36D6678DDBD" box="[506,589,620,644]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="3" pageNumber="234">UCDC</collectionCode>
).
<collectingCountry id="F3087668FFFA7C62FDDEB36D6684DDBC" box="[606,689,620,645]" name="Haiti" pageId="3" pageNumber="234">
<emphasis id="B96BEAEAFFFA7C62FDDEB36D6684DDBC" bold="true" box="[606,689,620,645]" pageId="3" pageNumber="234">HAITI</emphasis>
</collectingCountry>
:
<emphasis id="B96BEAEAFFFA7C62FD3FB36A6737DDBD" bold="true" box="[703,770,619,644]" italics="true" pageId="3" pageNumber="234">Quest</emphasis>
: Furcy, no date (WM
<collectingCountry id="F3087668FFFA7C62FC77B36C6009DDBD" box="[1015,1084,621,644]" name="Isle of Man" pageId="3" pageNumber="234">Mann</collectingCountry>
;
<collectionCode id="ED0EAE3DFFFA7C62FBC7B36D60B3DDBD" LSID="urn:lsid:biocol.org:col:33791" box="[1095,1158,620,644]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="234">MCZ</collectionCode>
,
<collectionCode id="ED0EAE3DFFFA7C62FB11B36D60D3DDBD" LSID="urn:lsid:biocol.org:col:34871" box="[1169,1254,620,644]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="234">USNM</collectionCode>
), Petionville, no date (WM
<collectingCountry id="F3087668FFFA7C62FE90B3916560DD9E" box="[272,341,656,679]" name="Isle of Man" pageId="3" pageNumber="234">Mann</collectingCountry>
;
<collectionCode id="ED0EAE3DFFFA7C62FEE0B3916583DD91" LSID="urn:lsid:biocol.org:col:34871" box="[352,438,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="234">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3056573FFFA7C62FF47B3B560A7DDC9" pageId="3" pageNumber="234" type="etymology">
<paragraph id="8BA036F8FFFA7C62FF47B3B560A7DDC9" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="234">
<emphasis id="B96BEAEAFFFA7C62FF47B3B5657BDDF4" bold="true" box="[199,334,692,717]" pageId="3" pageNumber="234">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C1F4D7BFFFA7C62FDBEB3B4669EDDF5" box="[574,683,693,716]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FDBEB3B4669EDDF5" box="[574,683,693,716]" italics="true" pageId="3" pageNumber="234">aterrimus</emphasis>
</taxonomicName>
(Latin,
<emphasis id="B96BEAEAFFFA7C62FC84B3B46704DDF5" box="[772,817,693,716]" italics="true" pageId="3" pageNumber="234">ater</emphasis>
= black; -
<emphasis id="B96BEAEAFFFA7C62FC22B3B467D7DDF5" box="[930,994,693,716]" italics="true" pageId="3" pageNumber="234">rimus</emphasis>
= superlative suffix), was derived from the deep coal black coloration of this species, which Wheeler mentioned in his description.
</paragraph>
</subSubSection>
<subSubSection id="C3056573FFFA7C62FF47B3FD650AD8AC" pageId="3" pageNumber="234" type="discussion">
<paragraph id="8BA036F8FFFA7C62FF47B3FD65BCDB45" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="234">
<emphasis id="B96BEAEAFFFA7C62FF47B3FD65E8DC2C" bold="true" box="[199,477,764,789]" pageId="3" pageNumber="234">Discussion and biology.</emphasis>
Genetic data are not available for
<taxonomicName id="4C1F4D7BFFFA7C62FCDFB3FC67D2DC2D" box="[863,999,765,788]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FCDFB3FC67D2DC2D" box="[863,999,765,788]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
, but bivariate plots demonstrate strong morphological differences between
<emphasis id="B96BEAEAFFFA7C62FDA9B220669ADC0E" box="[553,687,800,824]" italics="true" pageId="3" pageNumber="234">P. at err imus</emphasis>
and
<taxonomicName id="4C1F4D7BFFFA7C62FD69B220676BDC01" box="[745,862,800,824]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="B96BEAEAFFFA7C62FD69B220676BDC01" box="[745,862,800,824]" italics="true" pageId="3" pageNumber="234">P. schmitti</emphasis>
</taxonomicName>
(
<figureCitation id="13242A7DFFFA7C62FCEFB21E67E6DC01" box="[879,979,799,824]" captionStart="FIGURE 4" captionStartId="7.[151,250,1921,1943]" captionTargetBox="[320,1264,193,1919]" captionTargetId="figure@7.[309,1277,180,1919]" captionTargetPageId="7" captionText="FIGURE 4. Bivariate plots for workers and queens of Pogonomyrmex aterrimus, Pogonomyrmex “ darlingtoni ”, and Pogonomyrmex schmitti: (A) head width versus head length, (B) head width versus maximum length of hind femur, and (C) head width versus the ratio of maximal petiolar node width / maximal postpetiole width. Sample size for each caste is in parentheses." httpUri="https://zenodo.org/record/239578/files/figure.png" pageId="3" pageNumber="234">
<emphasis id="B96BEAEAFFFA7C62FCEFB21E67E6DC01" bold="true" box="[879,979,799,824]" pageId="3" pageNumber="234">Figure 4</emphasis>
</figureCitation>
). These differences include: (1) workers of
<taxonomicName id="4C1F4D7BFFFA7C62FF35B244650BDC65" box="[181,318,837,860]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FF35B244650BDC65" box="[181,318,837,860]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
are larger (HW =
<quantity id="4CE79B1DFFFA7C62FD8DB2456685DC65" box="[525,688,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" metricValueMax="1.35" metricValueMin="1.11" pageId="3" pageNumber="234" unit="mm" value="1.23" valueMax="1.35" valueMin="1.11">1.111.35 mm</quantity>
) compared to workers of
<taxonomicName id="4C1F4D7BFFFA7C62FC52B2446073DC65" box="[978,1094,836,860]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="B96BEAEAFFFA7C62FC52B2446073DC65" box="[978,1094,836,860]" italics="true" pageId="3" pageNumber="234">P. schmitti</emphasis>
</taxonomicName>
(HW =
<quantity id="4CE79B1DFFFA7C62FB24B245617EDC65" box="[1188,1355,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="1.035" metricValueMax="1.17" metricValueMin="0.9" pageId="3" pageNumber="234" unit="mm" value="1.035" valueMax="1.17" valueMin="0.9">0.901.17 mm</quantity>
), (2) in dorsal view, the petiolar node of
<taxonomicName id="4C1F4D7BFFFA7C62FD95B2686694DC46" box="[533,673,872,896]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FD95B2686694DC46" box="[533,673,872,896]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
is elongate, not broadly fan-shaped as in
<taxonomicName id="4C1F4D7BFFFA7C62FB07B26860C8DCB9" box="[1159,1277,872,896]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="B96BEAEAFFFA7C62FB07B26860C8DCB9" box="[1159,1277,872,896]" italics="true" pageId="3" pageNumber="234">P. schmitti</emphasis>
</taxonomicName>
, (3) in dorsal view, the petiolar node width/postpetiole width is lower in
<taxonomicName id="4C1F4D7BFFFA7C62FCC1B28C67F9DC9D" box="[833,972,909,932]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FCC1B28C67F9DC9D" box="[833,972,909,932]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
(PW/PPW = 0.780.90) compared to
<taxonomicName id="4C1F4D7BFFFA7C62FA07B28C64C5DCF1" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="B96BEAEAFFFA7C62FA07B28C64C5DCF1" italics="true" pageId="3" pageNumber="234">P. schmitti</emphasis>
</taxonomicName>
(0.871.03), (4) in
<taxonomicName id="4C1F4D7BFFFA7C62FE4AB2B06666DCFE" box="[458,595,944,968]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FE4AB2B065EDDCF1" box="[458,472,945,968]" italics="true" pageId="3" pageNumber="234">P</emphasis>
.
<emphasis id="B96BEAEAFFFA7C62FE66B2B16666DCFE" box="[486,595,944,967]" italics="true" pageId="3" pageNumber="234">aterrimus</emphasis>
</taxonomicName>
, the anterior surface of the petiolar node is partly to completely sculptured, not strongly shining, and it forms a right to weakly obtuse angle with the peduncle of the petiole, whereas in
<taxonomicName id="4C1F4D7BFFFA7C62FA07B2D464C5DB29" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="B96BEAEAFFFA7C62FA07B2D464C5DB29" italics="true" pageId="3" pageNumber="234">P. schmitti</emphasis>
</taxonomicName>
, the anterior surface of petiolar node is mostly smooth and shining, and it forms a noticeably obtuse angle with the peduncle of the petiole, and (5) the dorsum of the postpetiole and first gastral tergum are strongly punctate, dull in
<taxonomicName id="4C1F4D7BFFFA7C62FF68B5406547DB6E" box="[232,370,1088,1112]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FF68B54064C3DB61" box="[232,246,1089,1112]" italics="true" pageId="3" pageNumber="234">P</emphasis>
.
<emphasis id="B96BEAEAFFFA7C62FE85B5416547DB6E" box="[261,370,1088,1111]" italics="true" pageId="3" pageNumber="234">aterrimus</emphasis>
</taxonomicName>
, whereas these structures are smooth and shining to partly or completely punctate and weakly shining in
<taxonomicName id="4C1F4D7BFFFA7C62FE8DB56465B7DB45" box="[269,386,1124,1148]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="B96BEAEAFFFA7C62FE8DB564652EDB45" box="[269,283,1125,1148]" italics="true" pageId="3" pageNumber="234">P</emphasis>
.
<emphasis id="B96BEAEAFFFA7C62FEA9B56565B7DB45" box="[297,386,1124,1148]" italics="true" pageId="3" pageNumber="234">schmitti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA036F8FFFA7C62FF47B5886743DBD1" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="234">
<taxonomicName id="4C1F4D7BFFFA7C62FF47B58865DADBA6" box="[199,495,1160,1184]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FF47B58865DADBA6" box="[199,495,1160,1184]" italics="true" pageId="3" pageNumber="234">Pogonomyrmex aterrimus</emphasis>
</taxonomicName>
is not known to be sympatric with any congener, but collections of
<taxonomicName id="4C1F4D7BFFFA7C62FAA3B58861A9DB99" box="[1315,1436,1160,1184]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="B96BEAEAFFFA7C62FAA3B58861A9DB99" box="[1315,1436,1160,1184]" italics="true" pageId="3" pageNumber="234">P. schmitti</emphasis>
</taxonomicName>
proximate to those of
<taxonomicName id="4C1F4D7BFFFA7C62FE18B5AC6611DBFD" box="[408,548,1197,1220]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FE18B5AC6611DBFD" box="[408,548,1197,1220]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
suggest that the two species may occur sympatrically. Workers of these two species are easily distinguished using characters described above.
</paragraph>
<paragraph id="8BA036F8FFFA7C62FF47B5F466FCD915" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="234">
<taxonomicName id="4C1F4D7BFFFA7C62FF47B5F465CBDA35" box="[199,510,1268,1292]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B96BEAEAFFFA7C62FF47B5F465CBDA35" box="[199,510,1268,1292]" italics="true" pageId="3" pageNumber="234">Pogonomyrmex darlingtoni</emphasis>
</taxonomicName>
is morphologically very similar to
<taxonomicName id="4C1F4D7BFFFA7C62FC18B5F46011DA35" box="[920,1060,1269,1292]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FC18B5F46793DA35" box="[920,934,1269,1292]" italics="true" pageId="3" pageNumber="234">P</emphasis>
.
<emphasis id="B96BEAEAFFFA7C62FC37B5F46011DA35" box="[951,1060,1269,1292]" italics="true" pageId="3" pageNumber="234">aterrimus</emphasis>
</taxonomicName>
. The two forms differ mainly in degree of granulation on the interrugae of the cephalic dorsum and dorsum of the mesosoma (weakly to moderately granulate-punctate in
<taxonomicName id="4C1F4D7BFFFA7C62FE0FB43C662CDA6D" box="[399,537,1341,1364]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FE0FB43C662CDA6D" box="[399,537,1341,1364]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
, strongly granulate-punctate in
<taxonomicName id="4C1F4D7BFFFA7C62FC06B43C6014DA6D" box="[902,1057,1340,1364]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B96BEAEAFFFA7C62FC06B43C67A1DA6D" box="[902,916,1341,1364]" italics="true" pageId="3" pageNumber="234">P</emphasis>
.
<emphasis id="B96BEAEAFFFA7C62FC24B43D6014DA6D" box="[932,1057,1340,1364]" italics="true" pageId="3" pageNumber="234">darlingtoni</emphasis>
</taxonomicName>
). These differences are based on very limited material, especially for
<taxonomicName id="4C1F4D7BFFFA7C62FDAEB46066FDDA41" box="[558,712,1376,1400]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B96BEAEAFFFA7C62FDAEB46066FDDA41" box="[558,712,1376,1400]" italics="true" pageId="3" pageNumber="234">P. darlingtoni</emphasis>
</taxonomicName>
, which is known only from the
<typeStatus id="54A4885AFFFA7C62FBADB4616068DA41" box="[1069,1117,1376,1400]" pageId="3" pageNumber="234">type</typeStatus>
series of six workers. These
<typeStatus id="54A4885AFFFA7C62FF17B48464E7DAA4" box="[151,210,1413,1437]" pageId="3" pageNumber="234">types</typeStatus>
were collected near the western end of the Tiburon peninsula (
<emphasis id="B96BEAEAFFFA7C62FC2BB4856013DAA4" bold="true" box="[939,1062,1412,1437]" pageId="3" pageNumber="234">
<figureCitation id="13242A7DFFFA7C62FC2BB4856024DAA4" box="[939,1041,1412,1437]" captionStart="FIGURE 5" captionStartId="8.[151,250,1398,1420]" captionTargetBox="[410,1172,193,1371]" captionTargetId="figure@8.[410,1177,193,1377]" captionTargetPageId="8" captionText="FIGURE 5. Geographic distribution for (A) Pogonomyrmex aterrimus (▲), P. “ darlingtoni ” (■), and P. saucius (●), and (B) P. schmitti (●). The bold line down the left-center of the map is the border between Haiti and the Dominican Republic; lighter lines are the first administrative divisions in each country. Both maps are drawn on the same geographic area so that distribution patterns are directly comparable." httpUri="https://zenodo.org/record/239579/files/figure.png" pageId="3" pageNumber="234">Figure 5</figureCitation>
A
</emphasis>
), about
<quantity id="4CE79B1DFFFA7C62FB06B48560E8DAA4" box="[1158,1245,1412,1437]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="234" unit="km" value="170.0">170 km</quantity>
distant from the nearest collection of
<taxonomicName id="4C1F4D7BFFFA7C62FE0FB4A86629DA86" box="[399,540,1448,1472]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FE0FB4A86629DA86" box="[399,540,1448,1472]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
. At present we believe the most conservative approach is to interpret these sculptural differences as variation between populations of a single species. Additional collecting will clarify whether or not this is correct. Because this is a judgement call, we provide
<typeStatus id="54A4885AFFFA7C62FBE2B4EE60FCD931" box="[1122,1225,1519,1544]" pageId="3" pageNumber="234" type="lectotype">lectotype</typeStatus>
photographs and morphometric data for both taxa (
<emphasis id="B96BEAEAFFFA7C62FD91B715668DD914" bold="true" box="[529,696,1556,1581]" pageId="3" pageNumber="234">
<figureCitation id="13242A7DFFFA7C62FD91B71566B5D914" box="[529,640,1556,1581]" captionStart="FIGURE 1" captionStartId="4.[151,250,1941,1963]" captionTargetBox="[382,1172,195,1896]" captionTargetId="figure@4.[382,1179,195,1896]" captionTargetPageId="4" captionText="FIGURE 1. Photograph of Pogonomyrmex aterrimus Wheeler lectotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217240). Photographs by Erin Prado from www. antweb. org." httpUri="https://zenodo.org/record/239575/files/figure.png" pageId="3" pageNumber="234">Figures 1</figureCitation>
,
<figureCitation id="13242A7DFFFA7C62FD0EB71566A8D914" box="[654,669,1556,1581]" captionStart="FIGURE 2" captionStartId="5.[151,250,1930,1952]" captionTargetBox="[410,1144,193,1893]" captionTargetId="figure@5.[410,1150,193,1894]" captionTargetPageId="5" captionText="FIGURE 2. Photograph of Pogonomyrmex “ darlingtoni ” Wheeler lectotype worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0217241). Photographs by Erin Prado from www. antweb. org." httpUri="https://zenodo.org/record/239576/files/figure.png" pageId="3" pageNumber="234">2</figureCitation>
,
<figureCitation id="13242A7DFFFA7C62FD2AB715668DD914" box="[682,696,1556,1581]" captionStart="FIGURE 4" captionStartId="7.[151,250,1921,1943]" captionTargetBox="[320,1264,193,1919]" captionTargetId="figure@7.[309,1277,180,1919]" captionTargetPageId="7" captionText="FIGURE 4. Bivariate plots for workers and queens of Pogonomyrmex aterrimus, Pogonomyrmex “ darlingtoni ”, and Pogonomyrmex schmitti: (A) head width versus head length, (B) head width versus maximum length of hind femur, and (C) head width versus the ratio of maximal petiolar node width / maximal postpetiole width. Sample size for each caste is in parentheses." httpUri="https://zenodo.org/record/239578/files/figure.png" pageId="3" pageNumber="234">4</figureCitation>
</emphasis>
).
</paragraph>
<paragraph id="8BA036F8FFFA7C62FF47B73660A8D83D" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="234">
Little is known about the biology or habitat affinities of
<taxonomicName id="4C1F4D7BFFFA7C62FCD4B73867EBD976" box="[852,990,1592,1616]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FCD4B73867EBD976" box="[852,990,1592,1616]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
. The geographic range of
<taxonomicName id="4C1F4D7BFFFA7C62FA91B73861A9D976" box="[1297,1436,1592,1616]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="aterrimus">
<emphasis id="B96BEAEAFFFA7C62FA91B73861A9D976" box="[1297,1436,1592,1616]" italics="true" pageId="3" pageNumber="234">P. aterrimus</emphasis>
</taxonomicName>
appears to be restricted to mid- to higher elevation habitats of southcentral Hispaniola (
<emphasis id="B96BEAEAFFFA7C62FB1CB75D612CD94C" bold="true" box="[1180,1305,1628,1653]" pageId="3" pageNumber="234">
<figureCitation id="13242A7DFFFA7C62FB1CB75D6131D94C" box="[1180,1284,1628,1653]" captionStart="FIGURE 5" captionStartId="8.[151,250,1398,1420]" captionTargetBox="[410,1172,193,1371]" captionTargetId="figure@8.[410,1177,193,1377]" captionTargetPageId="8" captionText="FIGURE 5. Geographic distribution for (A) Pogonomyrmex aterrimus (▲), P. “ darlingtoni ” (■), and P. saucius (●), and (B) P. schmitti (●). The bold line down the left-center of the map is the border between Haiti and the Dominican Republic; lighter lines are the first administrative divisions in each country. Both maps are drawn on the same geographic area so that distribution patterns are directly comparable." httpUri="https://zenodo.org/record/239579/files/figure.png" pageId="3" pageNumber="234">Figure 5</figureCitation>
A
</emphasis>
). The only available habitat data (from collection labels) indicate that workers were taken in secondary montane forest edge (AL Wild #1349), tropical moist forest edge (PS Ward #11744), and a grassy field (D Lubertazzi #DL03790:001, DL03786:001, DL03785:006). These sites range in elevation from
<quantity id="4CE79B1DFFFA7C62FC24B7C66000D9E6" box="[932,1077,1735,1760]" metricMagnitude="3" metricUnit="m" metricValue="1.28" metricValueMax="1.8" metricValueMin="0.76" pageId="3" pageNumber="234" unit="m" value="1280.0" valueMax="1800.0" valueMin="760.0">7601800 m</quantity>
(AL Wild &amp; D Lubertazzi) to
<quantity id="4CE79B1DFFFA7C62FF17B7ED6575D83D" box="[151,320,1772,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.8175" metricValueMax="2.12" metricValueMin="1.515" pageId="3" pageNumber="234" unit="m" value="1817.5" valueMax="2120.0" valueMin="1515.0">1,5152,120 m</quantity>
(
<bibRefCitation id="EF8E4B09FFFA7C62FECEB7ED65C7D83C" author="Wheeler" box="[334,498,1772,1797]" pageId="3" pageNumber="234" refString="Wheeler, W. M. (1936) Ants from Hispaniola and Mona Island. Bulletin of the Museum of Comparative Zoology, 80, 195 - 211." type="journal article" year="1936">Wheeler 1936</bibRefCitation>
). Nests in the grassy fields had their chambers under a stone.
</paragraph>
<paragraph id="8BA036F8FFFA7C62FF47B611650AD8AC" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="234">
No information is available on colony size, but it seems doubtful that colonies contain more than 200
<specimenCount id="9D19FD71FFFA7C62FAF3B61161A8D811" box="[1395,1437,1808,1832]" pageId="3" pageNumber="234" type="worker">300</specimenCount>
workers (D. Lubertazzi, pers. comm.). One dealate queen was collected by WM
<collectingCountry id="F3087668FFFA7C62FBAEB634605AD875" box="[1070,1135,1845,1868]" name="Isle of Man" pageId="3" pageNumber="234">Mann</collectingCountry>
(MCZ), but the collection date was not given. Additional collections are needed to determine microhabitat preference and other aspects of natural history.
</paragraph>
</subSubSection>
</treatment>
</document>