647 lines
98 KiB
XML
647 lines
98 KiB
XML
<document id="D1631982A79592871798AA79D03087F3" ID-DOI="10.5281/zenodo.199308" ID-GBIF-Dataset="fd9b98c1-6c36-412d-ba70-8193e43c034d" ID-ISSN="1175-5326" ID-Zenodo-Dep="199308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460333992737" checkinUser="plazi" docAuthor="Anker, Arthur & Komai, Tomoyuki" docDate="2010" docId="03B687DEFF928F55FF74A333FEC3FA70" docLanguage="en" docName="zt02680p054.pdf" docOrigin="Zootaxa 2680" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Athanas ahyongi Anker & Komai, 2010, n. sp." docType="treatment" docVersion="9" lastPageNumber="53" masterDocId="FF8FFFA6FF938F5DFFE3A254FF86FFBC" masterDocTitle="Description of a new species of Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae) from Madagascar and Japan" masterLastPageNumber="54" masterPageNumber="45" pageNumber="46" updateTime="1698259306118" updateUser="plazi">
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<mods:titleInfo id="2E9110454DDC9910B4DC1D42D0B73F20">
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<mods:title id="BDD9E3F4D3DC7ADC309BBFFCFEADE15E">Description of a new species of Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae) from Madagascar and Japan</mods:title>
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<mods:name id="CE4F8AD9F646B28BB17B3E9F9FCD621B" type="personal">
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<mods:roleTerm id="247568A9A5D21F3071B9E8E134DE1E80">Author</mods:roleTerm>
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<mods:namePart id="2A562015A2705086B51706AAA32AADBC">Anker, Arthur</mods:namePart>
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<mods:roleTerm id="9AD1E75FDC5FB1783630E7888A0683F1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B36310EE1D2E43FF67BEE7DBE53F0829">Komai, Tomoyuki</mods:namePart>
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<mods:typeOfResource id="C3981F3222063FEDD03857A50C1C66F5">text</mods:typeOfResource>
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<mods:title id="C5C8436C0537B6A15247B9B3ACF2BA25">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FAA3FFE030E24A671E966636079B6377">
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<mods:date id="ACE7C0526BAF00F2901CE14F8A24266F">2010</mods:date>
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<mods:detail id="C3CD849B0328879375E0BBD7D798A4C2" type="volume">
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<mods:number id="BA6D032F86C2DE97D82FB7E6BAA7ECD7">2680</mods:number>
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<mods:start id="2EE695969473DE5706C60866093B114D">45</mods:start>
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<mods:end id="F59669F6E2E04F39DE87616E92D3F5AC">54</mods:end>
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<mods:classification id="0FAEA1DF430A3B7BEBAA2E2E25AB05CF">journal article</mods:classification>
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<mods:identifier id="4C66E82CCB36340F35BD56CB5B7EB69E" type="DOI">10.5281/zenodo.199308</mods:identifier>
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<mods:identifier id="DB5C62755A7B9870FE1A87507048FDBE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="57B6CF7F973303FC47A4C3BAE7BF0CC4" type="Zenodo-Dep">199308</mods:identifier>
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<treatment id="03B687DEFF928F55FF74A333FEC3FA70" ID-DOI="http://doi.org/10.5281/zenodo.6196514" ID-GBIF-Taxon="119404885" ID-Zenodo-Dep="6196514" LSID="urn:lsid:plazi:treatment:03B687DEFF928F55FF74A333FEC3FA70" httpUri="http://treatment.plazi.org/id/03B687DEFF928F55FF74A333FEC3FA70" lastPageId="8" lastPageNumber="53" pageId="1" pageNumber="46">
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<subSubSection id="C3056543FF928F5CFF74A333FE91FE18" pageId="1" pageNumber="46" type="nomenclature">
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<paragraph id="8BA036C8FF928F5CFF74A333FE28FE3D" blockId="1.[151,430,359,420]" box="[151,430,359,385]" pageId="1" pageNumber="46">
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<heading id="D0E881A4FF928F5CFF74A333FE28FE3D" bold="true" box="[151,430,359,385]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
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<emphasis id="B96BEADAFF928F5CFF74A333FE28FE3D" bold="true" box="[151,430,359,385]" pageId="1" pageNumber="46">
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<taxonomicName id="4C1F4D4BFF928F5CFF74A333FEE3FE3D" ID-CoL="85MBH" box="[151,357,359,385]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
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<emphasis id="B96BEADAFF928F5CFF74A333FEE3FE3D" bold="true" box="[151,357,359,385]" italics="true" pageId="1" pageNumber="46">Athanas ahyongi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25857A1FF928F5CFE8FA333FE28FE3D" box="[364,430,359,385]" pageId="1" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA036C8FF928F5CFF74A3DEFE91FE18" blockId="1.[151,430,359,420]" box="[151,279,394,420]" pageId="1" pageNumber="46">
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(
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<figureCitation id="13242A4DFF928F5CFF7CA3DEFE88FE18" box="[159,270,394,420]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,256,1133,1157]" captionStartId-1="3.[151,256,1914,1938]" captionStartId-2="4.[151,257,1865,1889]" captionStartId-3="5.[151,258,1344,1368]" captionStartId-4="6.[151,255,1892,1916]" captionStartId-5="7.[151,259,1656,1680]" captionTargetBox-0="[156,1431,196,1106]" captionTargetBox-1="[160,1426,803,1885]" captionTargetBox-2="[170,1416,182,1830]" captionTargetBox-3="[152,1436,203,1309]" captionTargetBox-4="[152,1436,463,1834]" captionTargetBox-5="[151,1436,186,1635]" captionTargetId-0="figure@2.[151,1436,183,1112]" captionTargetId-1="figure@3.[151,1436,791,1892]" captionTargetId-2="figure@4.[163,1421,178,1832]" captionTargetId-3="figure@5.[151,1436,194,1323]" captionTargetId-4="figure@6.[151,1436,437,1871]" captionTargetId-5="figure@7.[151,1436,186,1635]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." captionText-1="FIGURE 2. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, third maxilliped, lateral; B, second pereiopod, lateral; C, third pereiopod, lateral; D, fifth pereiopod, lateral; paratype, male (CL 1.9 mm), same locality (FLMNH UF Arthropoda 14094): E, second pleopod, with close-up of appendix masculina and appendix interna." captionText-2="FIGURE 3. Athanas ahyongi n. sp., holotype, female, CL 2.7 mm, Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, right cheliped, lateral; B, same, carpus and chela, mesial (= flexor view); C, left cheliped, lateral; D, same, carpus and chela, mesial (= flexor view)." captionText-3="FIGURE 4. Athanas ahyongi sp. nov., female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, third pereiopod, lateral." captionText-4="FIGURE 5. Athanas ahyongi n. sp., female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, right major cheliped, mesial; B, same, lateral; C, same, carpus and chela and, mesial (= flexor view); D, same, lateral (= extensor view); E, left minor cheliped, lateral; F, same, mesial; G, carpus and chela, lateral (= extensor view); H, same, chela, mesial (flexor view)." captionText-5="FIGURE 6. Athanas ahyongi n. sp., habitus and colour pattern. A, B, holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): dorsal (A) and lateral (B) [photographs by Arthur Anker]; C, D, specimen from Okinawa (not collected) in situ [photographs by Yusuke Yamada]." httpUri-0="https://zenodo.org/record/199309/files/figure.png" httpUri-1="https://zenodo.org/record/199310/files/figure.png" httpUri-2="https://zenodo.org/record/199311/files/figure.png" httpUri-3="https://zenodo.org/record/199312/files/figure.png" httpUri-4="https://zenodo.org/record/199313/files/figure.png" httpUri-5="https://zenodo.org/record/199314/files/figure.png" pageId="1" pageNumber="46">Figs. 1–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3056543FF928F5CFF74A381FA90FD16" pageId="1" pageNumber="46" type="materials_examined">
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<paragraph id="8BA036C8FF928F5CFF74A381FA90FD16" blockId="1.[151,1437,469,2026]" pageId="1" pageNumber="46">
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<emphasis id="B96BEADAFF928F5CFF74A381FE09FE53" bold="true" box="[151,399,469,495]" pageId="1" pageNumber="46">Material examined.</emphasis>
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<typeStatus id="54A4886AFF928F5CFE7BA381FD8DFE53" box="[408,523,469,495]" pageId="1" pageNumber="46" type="holotype">Holotype</typeStatus>
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: female (CL
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<quantity id="4CE79B2DFF928F5CFD4BA381FC8FFE53" box="[680,777,469,495]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="46" unit="mm" value="2.7">2.7 mm</quantity>
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),
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<collectingCountry id="F3087658FF928F5CFCFDA381FC29FE53" box="[798,943,469,495]" name="Madagascar" pageId="1" pageNumber="46">Madagascar</collectingCountry>
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, Nosy-Bé, west of Hellville,
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<geoCoordinate id="EE2B500FFF928F5CFAEEA381FA11FE53" box="[1293,1431,469,495]" direction="south" orientation="latitude" pageId="1" pageNumber="46" precision="9" value="-13.692667">13°41.56’S</geoCoordinate>
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,
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<geoCoordinate id="EE2B500FFF928F5CFF74A3AEFEA1FDA8" box="[151,295,506,532]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="9" value="48.41083">48°24.65’E</geoCoordinate>
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, sand flat with seagrass, rubble, patch reefs, some mounds of
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<taxonomicName id="4C1F4D4BFF928F5CFBC8A3AEFB20FDAF" box="[1067,1190,506,531]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEADAFF928F5CFBC8A3AEFB20FDAF" box="[1067,1190,506,531]" italics="true" pageId="1" pageNumber="46">Glypturus</emphasis>
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</taxonomicName>
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sp., depth
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<quantity id="4CE79B2DFF928F5CFACCA3AEFA1EFDA8" box="[1327,1432,506,532]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="46" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 m</quantity>
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, suction (yabby) pump, from burrows or mounds, coll. A. Anker,
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<date id="FFA11008FF928F5CFC7BA074FBBEFD86" box="[920,1080,544,570]" pageId="1" pageNumber="46" value="2008-05-25">25 May 2008</date>
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, fcn
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<collectionCode id="ED0EAE0DFF928F5CFB90A074FB66FD86" box="[1139,1248,544,570]" pageId="1" pageNumber="46">BMADA</collectionCode>
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0 466 (
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<collectionCode id="ED0EAE0DFF928F5CFAD0A074FA1AFD86" box="[1331,1436,544,570]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="46">FLMNH</collectionCode>
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UF Arthropoda 14093).
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<typeStatus id="54A4886AFF928F5CFE51A011FD9BFDE3" box="[434,541,581,607]" pageId="1" pageNumber="46" type="paratype">Paratype</typeStatus>
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: male (CL
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<quantity id="4CE79B2DFF928F5CFD7DA011FD7DFDE3" box="[670,763,581,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="46" unit="mm" value="1.9">1.9 mm</quantity>
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), same collection data as for
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<typeStatus id="54A4886AFF928F5CFBAFA011FB30FDE3" box="[1100,1206,581,607]" pageId="1" pageNumber="46" type="holotype">holotype</typeStatus>
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, fcn
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<collectionCode id="ED0EAE0DFF928F5CFB0EA011FADCFDE3" box="[1261,1370,581,607]" pageId="1" pageNumber="46">BMADA</collectionCode>
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0 467 (
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<collectionCode id="ED0EAE0DFF928F5CFF7CA03EFE8FFD38" box="[159,265,618,644]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="46">FLMNH</collectionCode>
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UF Arthropoda 14094). Non-type:
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<specimenCount id="9D19FD41FF928F5CFD55A03EFC99FD38" box="[694,799,618,644]" pageId="1" pageNumber="46" type="female">1 female</specimenCount>
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(CL
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<quantity id="4CE79B2DFF928F5CFCBCA03EFC46FD38" box="[863,960,618,644]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="46" unit="mm" value="2.6">2.6 mm</quantity>
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),
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<collectingCountry id="F3087658FF928F5CFC36A03EFB9AFD38" box="[981,1052,618,644]" name="Japan" pageId="1" pageNumber="46">Japan</collectingCountry>
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, Ryukyu Archipelago, Okinawa Island, Red Beach, Kin, under rock, depth
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<quantity id="4CE79B2DFF928F5CFD68A0C4FD3DFD16" box="[651,699,656,682]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="46" unit="m" value="8.0">8 m</quantity>
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, coll. Y. Yamada,
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<date id="FFA11008FF928F5CFC6CA0C4FBB5FD16" box="[911,1075,656,682]" pageId="1" pageNumber="46" value="2009-03-04">4 March 2009</date>
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(CBM-ZC 10014).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3056543FF928F5EFF26A0E1FAF8FD5E" lastPageId="3" lastPageNumber="48" pageId="1" pageNumber="46" type="description">
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<paragraph id="8BA036C8FF928F5CFF26A0E1FC6FFC13" blockId="1.[151,1437,469,2026]" pageId="1" pageNumber="46">
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<emphasis id="B96BEADAFF928F5CFF26A0E1FED0FD73" bold="true" box="[197,342,693,719]" pageId="1" pageNumber="46">Description</emphasis>
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(based on
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<typeStatus id="54A4886AFF928F5CFE36A0E1FD81FD73" box="[469,519,693,719]" pageId="1" pageNumber="46">type</typeStatus>
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specimens from Nosy-Bé). Small-sized alpheid shrimp. Body not particularly elongate, moderately slender. Carapace glabrous, not setose. Rostrum well developed, laterally compressed, horizontal, triangular in dorsal view, longer than broad; lateral margin slightly concave from rostral base to about mid-length, with some short setae; tip acute, not reaching distal margin of first segment of antennular peduncle; rostral carina faint, not reaching base of rostrum (
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<figureCitation id="13242A4DFF928F5CFC87A11EFC21FCD8" box="[868,935,842,868]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
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A, B). Extracorneal teeth well developed, triangular, sharp, not reaching anterior margin of cornea; infracorneal and supracorneal teeth absent (
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<figureCitation id="13242A4DFF928F5CFADAA124FAFDFC36" box="[1337,1403,880,906]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
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A, B). Pterygostomial angle broadly rounded; cardiac notch deep (
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<figureCitation id="13242A4DFF928F5CFC9DA1C1FC46FC13" box="[894,960,917,943]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
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A).
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</paragraph>
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<paragraph id="8BA036C8FF928F5CFF26A1EEFC12FB33" blockId="1.[151,1437,469,2026]" pageId="1" pageNumber="46">
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Pleura of first to fourth abdominal somites rounded posteroventrally; fifth pleuron with more pronounced posteroventral angle; sixth somite with large articulated triangular flap (
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<figureCitation id="13242A4DFF928F5CFC10A1B4FBB1FC46" box="[1011,1079,992,1018]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
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C). Telson widest at proximal third, distally tapering; dorsal surface with two pairs of strong spiniform setae inserted in deep pits; posterior margin slightly rounded medially, with two pairs of stout spiniform setae at each lateral angle, mesial almost twice as long as lateral; mesial spiniform setae obscured by long plumose setae originating from median area of posterior margin, between mesial spiniform setae (
|
||
<figureCitation id="13242A4DFF928F5CFCEBA621FCCFFB33" box="[776,841,1141,1167]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
|
||
D, E).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF928F5CFF26A6CEFF3DFB66" blockId="1.[151,1437,469,2026]" pageId="1" pageNumber="46">
|
||
Eyes almost completely exposed in dorsal and lateral views; cornea large, normally pigmented (
|
||
<figureCitation id="13242A4DFF928F5CFADAA6CEFAFDFB08" box="[1337,1403,1178,1204]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF928F5CFF26A6B1FDC3FA63" blockId="1.[151,1437,469,2026]" pageId="1" pageNumber="46">
|
||
Antennular peduncle stout, with second segment squarish, about as long as wide, shorter than dorsally visible portion of first segment, equal to third segment; stylocerite sharp, reaching to about 0.7–0.8 length of second segment (
|
||
<figureCitation id="13242A4DFF928F5CFE82A764FE24FAF6" box="[353,418,1328,1354]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
|
||
A); ventromesial carina with very strong, anteriorly produced tooth, as illustrated (
|
||
<figureCitation id="13242A4DFF928F5CFA8FA764FF25FAD3" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
|
||
G); lateral flagellum biramous, fused portion composed of five segments. Antenna with basicerite stout, bearing small subacute tooth distoventrally; scaphocerite slightly exceeding end of antennular peduncle, ovate; blade broad, with convex anterior margin reaching beyond distolateral tooth; carpocerite distinctly exceeding scaphocerite (
|
||
<figureCitation id="13242A4DFF928F5CFE54A791FE7FFA63" box="[439,505,1477,1503]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF928F5CFF26A7BEFA1AF9C8" blockId="1.[151,1437,469,2026]" pageId="1" pageNumber="46">
|
||
Mouthparts not dissected, appearing typical for genus in external view. Third maxilliped with coxa bearing large, distally acute lateral plate above mastigobranch; antepenultimate segment slender, somewhat flattened; penultimate segment almost four times as long as wide proximally, distally slightly widening; ultimate segment furnished with transverse rows of thick serrulate setae, tip without spiniform setae (
|
||
<figureCitation id="13242A4DFF928F5CFAD2A40EFAF1F9C8" box="[1329,1399,1626,1652]" captionStart="FIGURE 2" captionStartId="3.[151,256,1914,1938]" captionTargetBox="[160,1426,803,1885]" captionTargetId="figure@3.[151,1436,791,1892]" captionTargetPageId="3" captionText="FIGURE 2. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, third maxilliped, lateral; B, second pereiopod, lateral; C, third pereiopod, lateral; D, fifth pereiopod, lateral; paratype, male (CL 1.9 mm), same locality (FLMNH UF Arthropoda 14094): E, second pleopod, with close-up of appendix masculina and appendix interna." httpUri="https://zenodo.org/record/199310/files/figure.png" pageId="1" pageNumber="46">Fig. 2</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF928F5CFF26A4D4FEF7F856" blockId="1.[151,1437,469,2026]" pageId="1" pageNumber="46">
|
||
Chelipeds symmetrical in shape, equal in size (
|
||
<figureCitation id="13242A4DFF928F5CFD0DA4D4FCB5F926" box="[750,819,1664,1690]" captionStart="FIGURE 3" captionStartId="4.[151,257,1865,1889]" captionTargetBox="[170,1416,182,1830]" captionTargetId="figure@4.[163,1421,178,1832]" captionTargetPageId="4" captionText="FIGURE 3. Athanas ahyongi n. sp., holotype, female, CL 2.7 mm, Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, right cheliped, lateral; B, same, carpus and chela, mesial (= flexor view); C, left cheliped, lateral; D, same, carpus and chela, mesial (= flexor view)." httpUri="https://zenodo.org/record/199311/files/figure.png" pageId="1" pageNumber="46">Fig. 3</figureCitation>
|
||
A, C), similar in both sexes, slightly more slender in male; ischium elongate, at least six times as long as wide, with ventrolateral margin strongly toothed, distal margin with three spiniform setae proximally; merus elongate, about as long as ischium, ventral surface, especially distally, flattened, ventrolateral margin strongly toothed; carpus vase-shaped, distally broadening, somewhat depressed laterally; chela laterally somewhat compressed, smooth except for areas of minute, barely visible granules on ventrolateral surface; fingers shorter than palm, feebly curved, with a moderately dense brush of elongate stiff setae originating from lateral surface of both fingers (
|
||
<figureCitation id="13242A4DFF928F5CFB6AA534FB54F8C6" box="[1161,1234,1888,1914]" captionStart="FIGURE 3" captionStartId="4.[151,257,1865,1889]" captionTargetBox="[170,1416,182,1830]" captionTargetId="figure@4.[163,1421,178,1832]" captionTargetPageId="4" captionText="FIGURE 3. Athanas ahyongi n. sp., holotype, female, CL 2.7 mm, Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, right cheliped, lateral; B, same, carpus and chela, mesial (= flexor view); C, left cheliped, lateral; D, same, carpus and chela, mesial (= flexor view)." httpUri="https://zenodo.org/record/199311/files/figure.png" pageId="1" pageNumber="46">Fig. 3</figureCitation>
|
||
A); dactylus and pollex each with large triangular tooth fitting in a deep gap on opposable margin, tooth on pollex being more proximal than tooth on dactylus; pollex with additional, much smaller tooth adjacent and more distal to large tooth (
|
||
<figureCitation id="13242A4DFF928F5CFF00A584FEAFF856" box="[227,297,2000,2026]" captionStart="FIGURE 3" captionStartId="4.[151,257,1865,1889]" captionTargetBox="[170,1416,182,1830]" captionTargetId="figure@4.[163,1421,178,1832]" captionTargetPageId="4" captionText="FIGURE 3. Athanas ahyongi n. sp., holotype, female, CL 2.7 mm, Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, right cheliped, lateral; B, same, carpus and chela, mesial (= flexor view); C, left cheliped, lateral; D, same, carpus and chela, mesial (= flexor view)." httpUri="https://zenodo.org/record/199311/files/figure.png" pageId="1" pageNumber="46">Fig. 3</figureCitation>
|
||
B, D).
|
||
</paragraph>
|
||
<caption id="DF606640FF918F5FFF74A639FBACFB59" httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="2" pageNumber="47" targetBox="[156,1431,196,1106]" targetPageId="2">
|
||
<paragraph id="8BA036C8FF918F5FFF74A639FBACFB59" blockId="2.[151,1437,1133,1253]" pageId="2" pageNumber="47">
|
||
<emphasis id="B96BEADAFF918F5FFF74A639FE9AFB39" bold="true" box="[151,284,1133,1157]" pageId="2" pageNumber="47">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C1F4D4BFF918F5FFEC0A63AFE5DFB39" box="[291,475,1134,1157]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF918F5FFEC0A63AFE5DFB39" box="[291,475,1134,1157]" italics="true" pageId="2" pageNumber="47">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF918F5FFE00A639FDA7FB39" bold="true" box="[483,545,1133,1157]" pageId="2" pageNumber="47">
|
||
<taxonomicNameLabel id="A25857A1FF918F5FFE00A639FDA7FB39" box="[483,545,1133,1157]" pageId="2" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA036C8FF918F5FFF26A747FD8EF974" blockId="2.[151,1437,1299,2035]" pageId="2" pageNumber="47">
|
||
Second pereiopod with ischium shorter than merus, both unarmed; carpus five-segmented, first segment longest, about the same length as remaining four segments, ratio of carpal segments (proximal to distal) approximately: 4: 1: 1: 1: 2; chela simple, longer than most-distal carpal article, with fingers slightly longer than palm (
|
||
<figureCitation id="13242A4DFF918F5FFEC1A7D7FEEBFA21" box="[290,365,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[151,256,1914,1938]" captionTargetBox="[160,1426,803,1885]" captionTargetId="figure@3.[151,1436,791,1892]" captionTargetPageId="3" captionText="FIGURE 2. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, third maxilliped, lateral; B, second pereiopod, lateral; C, third pereiopod, lateral; D, fifth pereiopod, lateral; paratype, male (CL 1.9 mm), same locality (FLMNH UF Arthropoda 14094): E, second pleopod, with close-up of appendix masculina and appendix interna." httpUri="https://zenodo.org/record/199310/files/figure.png" pageId="2" pageNumber="47">Fig. 2</figureCitation>
|
||
B). Third pereiopod slender; ischium with one or two spiniform seta(e) on ventrolateral surface, dorsal margin with small spiniform seta proximally; merus more than five times as long as wide, unarmed; carpus conspicuously more slender than merus, about half as long as merus, distoventral angle with small spiniform seta; propodus as wide as carpus, ventral margin with three small spiniform setae, including a longer distal spiniform seta adjacent to dactylus; dactylus about 0.6 propodus length, simple, extremely slender, gradually curving distally (
|
||
<figureCitation id="13242A4DFF918F5FFDA9A46AFD13F9E4" box="[586,661,1598,1624]" captionStart="FIGURE 2" captionStartId="3.[151,256,1914,1938]" captionTargetBox="[160,1426,803,1885]" captionTargetId="figure@3.[151,1436,791,1892]" captionTargetPageId="3" captionText="FIGURE 2. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, third maxilliped, lateral; B, second pereiopod, lateral; C, third pereiopod, lateral; D, fifth pereiopod, lateral; paratype, male (CL 1.9 mm), same locality (FLMNH UF Arthropoda 14094): E, second pleopod, with close-up of appendix masculina and appendix interna." httpUri="https://zenodo.org/record/199310/files/figure.png" pageId="2" pageNumber="47">Fig. 2</figureCitation>
|
||
C). Fourth pereiopod generally similar to third. Fifth pereiopod similar to third or fourth, but more slender; ischium with one small spiniform seta; propodus without spiniform setae, instead with three rows of thick serrulate setae on distolateral surface, most-distal row bearing longest setae (
|
||
<figureCitation id="13242A4DFF918F5FFE7EA4FAFE65F974" box="[413,483,1710,1736]" captionStart="FIGURE 2" captionStartId="3.[151,256,1914,1938]" captionTargetBox="[160,1426,803,1885]" captionTargetId="figure@3.[151,1436,791,1892]" captionTargetPageId="3" captionText="FIGURE 2. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, third maxilliped, lateral; B, second pereiopod, lateral; C, third pereiopod, lateral; D, fifth pereiopod, lateral; paratype, male (CL 1.9 mm), same locality (FLMNH UF Arthropoda 14094): E, second pleopod, with close-up of appendix masculina and appendix interna." httpUri="https://zenodo.org/record/199310/files/figure.png" pageId="2" pageNumber="47">Fig. 2</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF918F5FFF26A487FB33F8E1" blockId="2.[151,1437,1299,2035]" pageId="2" pageNumber="47">
|
||
First pleopod with simple endopod, furnished with some setae, reaching half-length of exopod. Male second pleopod with appendix masculina shorter than appendix interna, apex with three slender setae (
|
||
<figureCitation id="13242A4DFF918F5FFA8FA4ADFF21F884" captionStart="FIGURE 2" captionStartId="3.[151,256,1914,1938]" captionTargetBox="[160,1426,803,1885]" captionTargetId="figure@3.[151,1436,791,1892]" captionTargetPageId="3" captionText="FIGURE 2. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, third maxilliped, lateral; B, second pereiopod, lateral; C, third pereiopod, lateral; D, fifth pereiopod, lateral; paratype, male (CL 1.9 mm), same locality (FLMNH UF Arthropoda 14094): E, second pleopod, with close-up of appendix masculina and appendix interna." httpUri="https://zenodo.org/record/199310/files/figure.png" pageId="2" pageNumber="47">Fig. 2</figureCitation>
|
||
E). Uropod with lateral lobe of protopod obscurely bifid, with small subacute point; diaeresis straight, ending in small acute tooth distolaterally, adjacent to slender distolateral spiniform seta (
|
||
<figureCitation id="13242A4DFF918F5FFBAEA517FB09F8E1" box="[1101,1167,1859,1885]" captionStart="FIGURE 1" captionStartId="2.[151,256,1133,1157]" captionTargetBox="[156,1431,196,1106]" captionTargetId="figure@2.[151,1436,183,1112]" captionTargetPageId="2" captionText="FIGURE 1. Athanas ahyongi n. sp., holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, sixth abdominal somite and tail fan, lateral; D, telson, dorsal; E, same, posterior margin (plumose setae omitted); F, uropod, lateral; G, mesioventral tooth on first segment of antennular peduncle, lateral." httpUri="https://zenodo.org/record/199309/files/figure.png" pageId="2" pageNumber="47">Fig. 1</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF918F5FFF26A53DFF7CF814" blockId="2.[151,1437,1299,2035]" pageId="2" pageNumber="47">
|
||
Gill-exopod formula typical for genus (see
|
||
<bibRefCitation id="EF8E4B39FF918F5FFD21A53DFC2BF83F" author="Anker" box="[706,941,1897,1923]" pageId="2" pageNumber="47" refString="Anker, A & Jeng, M. - S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinition of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker & Jeng 2007</bibRefCitation>
|
||
); no rudimentary exopods visible on basis of P1–2.
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF918F5EFF26A5E7FAD5FEFB" blockId="2.[151,1437,1299,2035]" lastBlockId="3.[151,1437,152,738]" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
|
||
<emphasis id="B96BEADAFF918F5FFF26A5E7FE04F871" bold="true" box="[197,386,1971,1997]" pageId="2" pageNumber="47">Colour in life.</emphasis>
|
||
Ground colour semitransparent greyish; most of mediodorsal area of carapace and abdomen occupied by broad white to pale yellowish band, extending from rostrum to sixth abdominal somite and continuing onto telson; flanks of carapace and abdomen with broad red band extending from anterolateral margin of carapace below eyes to sixth abdominal somite and continuing onto uropods; chelipeds and second to fifth pereiopods semitransparent-whitish; antennular peduncle partly colourless, partly white-yellowish, stylocerite bright red, antennular flagellum pale-orange; antennal basicerite, carpocerite and scaphocerite conspicuously red, antennal flagellum red proximally, colourless distally; corneas greyish-blue (
|
||
<figureCitation id="13242A4DFF908F5EFB1EA379FAC3FEFB" box="[1277,1349,301,327]" captionStart="FIGURE 6" captionStartId="7.[151,259,1656,1680]" captionTargetBox="[151,1436,186,1635]" captionTargetId="figure@7.[151,1436,186,1635]" captionTargetPageId="7" captionText="FIGURE 6. Athanas ahyongi n. sp., habitus and colour pattern. A, B, holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): dorsal (A) and lateral (B) [photographs by Arthur Anker]; C, D, specimen from Okinawa (not collected) in situ [photographs by Yusuke Yamada]." httpUri="https://zenodo.org/record/199314/files/figure.png" pageId="3" pageNumber="48">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF908F5EFF26A306FAF8FD5E" blockId="3.[151,1437,152,738]" pageId="3" pageNumber="48">
|
||
<emphasis id="B96BEADAFF908F5EFF26A306FEC6FED0" bold="true" box="[197,320,338,364]" pageId="3" pageNumber="48">Variation.</emphasis>
|
||
The non-type female from Okinawa differs from the two
|
||
<typeStatus id="54A4886AFF908F5EFC12A306FBA2FED0" box="[1009,1060,338,364]" pageId="3" pageNumber="48">type</typeStatus>
|
||
specimens from
|
||
<collectingCountry id="F3087658FF908F5EFB0CA306FAFBFED0" box="[1263,1405,338,364]" name="Madagascar" pageId="3" pageNumber="48">Madagascar</collectingCountry>
|
||
in several features, including the slightly shorter stylocerite, reaching to only half the length of the second segment of antennular peduncle (
|
||
<figureCitation id="13242A4DFF908F5EFDFEA3C9FDE3FE0B" box="[541,613,413,439]" captionStart="FIGURE 4" captionStartId="5.[151,258,1344,1368]" captionTargetBox="[152,1436,203,1309]" captionTargetId="figure@5.[151,1436,194,1323]" captionTargetPageId="5" captionText="FIGURE 4. Athanas ahyongi sp. nov., female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, third pereiopod, lateral." httpUri="https://zenodo.org/record/199312/files/figure.png" pageId="3" pageNumber="48">Fig. 4</figureCitation>
|
||
A), and the degree of symmetry and proportions of the chelipeds (
|
||
<figureCitation id="13242A4DFF908F5EFA8FA3C9FF2EFE60" captionStart="FIGURE 5" captionStartId="6.[151,255,1892,1916]" captionTargetBox="[152,1436,463,1834]" captionTargetId="figure@6.[151,1436,437,1871]" captionTargetPageId="6" captionText="FIGURE 5. Athanas ahyongi n. sp., female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, right major cheliped, mesial; B, same, lateral; C, same, carpus and chela and, mesial (= flexor view); D, same, lateral (= extensor view); E, left minor cheliped, lateral; F, same, mesial; G, carpus and chela, lateral (= extensor view); H, same, chela, mesial (flexor view)." httpUri="https://zenodo.org/record/199313/files/figure.png" pageId="3" pageNumber="48">Fig. 5</figureCitation>
|
||
). While in the female
|
||
<typeStatus id="54A4886AFF908F5EFE57A396FD99FE60" box="[436,543,450,476]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
, the chelipeds are almost perfectly equal in size and symmetrical in shape, the chelipeds of the female from Okinawa are both unequal and asymmetrical, i.e. it has a major cheliped (
|
||
<figureCitation id="13242A4DFF908F5EFA8EA3BCFF22FD9B" captionStart="FIGURE 5" captionStartId="6.[151,255,1892,1916]" captionTargetBox="[152,1436,463,1834]" captionTargetId="figure@6.[151,1436,437,1871]" captionTargetPageId="6" captionText="FIGURE 5. Athanas ahyongi n. sp., female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, right major cheliped, mesial; B, same, lateral; C, same, carpus and chela and, mesial (= flexor view); D, same, lateral (= extensor view); E, left minor cheliped, lateral; F, same, mesial; G, carpus and chela, lateral (= extensor view); H, same, chela, mesial (flexor view)." httpUri="https://zenodo.org/record/199313/files/figure.png" pageId="3" pageNumber="48">Fig. 5</figureCitation>
|
||
A-D) and a minor cheliped (
|
||
<figureCitation id="13242A4DFF908F5EFE0DA059FDB4FD9B" box="[494,562,525,551]" captionStart="FIGURE 5" captionStartId="6.[151,255,1892,1916]" captionTargetBox="[152,1436,463,1834]" captionTargetId="figure@6.[151,1436,437,1871]" captionTargetPageId="6" captionText="FIGURE 5. Athanas ahyongi n. sp., female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, right major cheliped, mesial; B, same, lateral; C, same, carpus and chela and, mesial (= flexor view); D, same, lateral (= extensor view); E, left minor cheliped, lateral; F, same, mesial; G, carpus and chela, lateral (= extensor view); H, same, chela, mesial (flexor view)." httpUri="https://zenodo.org/record/199313/files/figure.png" pageId="3" pageNumber="48">Fig. 5</figureCitation>
|
||
E-H). The major cheliped of the Okinawa female is generally similar to the cheliped of the female
|
||
<typeStatus id="54A4886AFF908F5EFE52A066FD99FDF0" box="[433,543,562,588]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
, except for the much shorter carpus. The minor cheliped differs from the major cheliped by the distinctly longer carpus (longer than the carpus in the
|
||
<typeStatus id="54A4886AFF908F5EFBF3A00CFBFDFDCE" box="[1040,1147,600,626]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
), the much smaller chela (only about 30–40% of the volume of the large chela), and the absence of a strong armature on the fingers. These differences are here interpreted as intraspecific in view of the fact that the cheliped polymorphism is not uncommon in
|
||
<taxonomicName id="4C1F4D4BFF908F5EFEA3A09CFE26FD5D" box="[320,416,712,737]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEADAFF908F5EFEA3A09CFE26FD5D" box="[320,416,712,737]" italics="true" pageId="3" pageNumber="48">Athanas</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<bibRefCitation id="EF8E4B39FF908F5EFE00A09CFDF6FD5E" author="Kemp" box="[483,624,712,738]" pageId="3" pageNumber="48" refString="Kemp, S. (1915) The fauna of the Chilka Lake, Crustacea, Decapoda. Memoirs of the Indian Museum, 5, 199 - 326, pls. 12 - 13." type="journal article" year="1915">Kemp 1915</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF908F5EFD9FA09CFC25FD5E" author="Banner" box="[636,931,712,738]" pageId="3" pageNumber="48" refString="Banner, A. H. & Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner and Banner 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF908F5EFC4CA09CFBB8FD5E" author="Anker" box="[943,1086,712,738]" pageId="3" pageNumber="48" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF908F5EFBA9A09CFAF4FD5E" author="Anker" box="[1098,1394,712,738]" pageId="3" pageNumber="48" refString="Anker, A and Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007">Anker and Ahyong 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF606640FF908F5EFF74A52EFDDFF84D" httpUri="https://zenodo.org/record/199310/files/figure.png" pageId="3" pageNumber="48" targetBox="[160,1426,803,1885]" targetPageId="3">
|
||
<paragraph id="8BA036C8FF908F5EFF74A52EFDDFF84D" blockId="3.[151,1436,1914,2033]" pageId="3" pageNumber="48">
|
||
<emphasis id="B96BEADAFF908F5EFF74A52EFE9DF82E" bold="true" box="[151,283,1914,1938]" pageId="3" pageNumber="48">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C1F4D4BFF908F5EFEC0A52EFE5DF82D" box="[291,475,1914,1937]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF908F5EFEC0A52EFE5DF82D" box="[291,475,1914,1937]" italics="true" pageId="3" pageNumber="48">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25857A1FF908F5EFE00A52EFDA7F82E" box="[483,545,1914,1938]" pageId="3" pageNumber="48" rank="species">
|
||
<emphasis id="B96BEADAFF908F5EFE00A52EFDA7F82E" bold="true" box="[483,545,1914,1938]" pageId="3" pageNumber="48">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, third maxilliped, lateral; B, second pereiopod, lateral; C, third pereiopod, lateral; D, fifth pereiopod, lateral; paratype, male (CL 1.9 mm), same locality (FLMNH UF Arthropoda 14094): E, second pleopod, with close-up of appendix masculina and appendix interna.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF606640FF978F59FF74A51DFDCDF81D" httpUri="https://zenodo.org/record/199311/files/figure.png" pageId="4" pageNumber="49" targetBox="[170,1416,182,1830]" targetPageId="4">
|
||
<paragraph id="8BA036C8FF978F59FF74A51DFDCDF81D" blockId="4.[151,1436,1865,1953]" pageId="4" pageNumber="49">
|
||
<emphasis id="B96BEADAFF978F59FF74A51DFE9BF8DD" bold="true" box="[151,285,1865,1889]" pageId="4" pageNumber="49">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C1F4D4BFF978F59FEC7A51EFE59F8DD" box="[292,479,1866,1889]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF978F59FEC7A51EFE59F8DD" box="[292,479,1866,1889]" italics="true" pageId="4" pageNumber="49">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF978F59FE05A51DFDA2F8DD" bold="true" box="[486,548,1865,1889]" pageId="4" pageNumber="49">
|
||
<taxonomicNameLabel id="A25857A1FF978F59FE05A51DFDA2F8DD" box="[486,548,1865,1889]" pageId="4" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female, CL 2.7 mm, Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): A, right cheliped, lateral; B, same, carpus and chela, mesial (= flexor view); C, left cheliped, lateral; D, same, carpus and chela, mesial (= flexor view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF606640FF968F58FF74A714FA91FAC4" httpUri="https://zenodo.org/record/199312/files/figure.png" pageId="5" pageNumber="50" targetBox="[152,1436,203,1309]" targetPageId="5">
|
||
<paragraph id="8BA036C8FF968F58FF74A714FA91FAC4" blockId="5.[151,1437,1344,1400]" pageId="5" pageNumber="50">
|
||
<emphasis id="B96BEADAFF968F58FF74A714FE99FAE4" bold="true" box="[151,287,1344,1368]" pageId="5" pageNumber="50">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C1F4D4BFF968F58FEC4A715FE63FAE4" box="[295,485,1345,1368]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF968F58FEC4A715FE63FAE4" box="[295,485,1345,1368]" italics="true" pageId="5" pageNumber="50">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF968F58FE0FA714FDC1FAE4" bold="true" box="[492,583,1344,1368]" pageId="5" pageNumber="50">
|
||
<taxonomicNameLabel id="A25857A1FF968F58FE0FA714FDC1FAE4" box="[492,583,1344,1368]" pageId="5" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, carapace and frontal appendages, lateral; B, frontal region of carapace and frontal appendages, dorsal; C, third pereiopod, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3056543FF968F58FF26A7F2FBF9FA59" pageId="5" pageNumber="50" type="etymology">
|
||
<paragraph id="8BA036C8FF968F58FF26A7F2FBF9FA59" blockId="5.[151,1437,1446,2032]" pageId="5" pageNumber="50">
|
||
<emphasis id="B96BEADAFF968F58FF26A7F2FED5FA7C" bold="true" box="[197,339,1446,1472]" pageId="5" pageNumber="50">Etymology.</emphasis>
|
||
Named after our friend and colleague, Dr. Shane T. Ahyong (Australian Museum, Sydney) for his numerous contributions to decapod and stomatopod taxonomy and phylogeny.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3056543FF968F58FF26A7A4FDFAF9B7" box="[197,636,1520,1547]" pageId="5" pageNumber="50" type="materials_examined">
|
||
<paragraph id="8BA036C8FF968F58FF26A7A4FDFAF9B7" blockId="5.[151,1437,1446,2032]" box="[197,636,1520,1547]" pageId="5" pageNumber="50">
|
||
<emphasis id="B96BEADAFF968F58FF26A7A4FEECF9B6" bold="true" box="[197,362,1520,1546]" pageId="5" pageNumber="50">
|
||
<typeStatus id="54A4886AFF968F58FF26A7A4FE84F9B6" box="[197,258,1520,1546]" pageId="5" pageNumber="50">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Nosy-Bé,
|
||
<collectingCountry id="F3087658FF968F58FE0BA7A5FDFEF9B7" box="[488,632,1521,1547]" name="Madagascar" pageId="5" pageNumber="50">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3056543FF968F58FF26A442FD7DF83C" pageId="5" pageNumber="50" type="distribution">
|
||
<paragraph id="8BA036C8FF968F58FF26A442FDC1F9C7" blockId="5.[151,1437,1446,2032]" pageId="5" pageNumber="50">
|
||
<emphasis id="B96BEADAFF968F58FF26A442FEE8F98C" bold="true" box="[197,366,1558,1584]" pageId="5" pageNumber="50">Distribution.</emphasis>
|
||
Presently known only from northwestern
|
||
<collectingCountry id="F3087658FF968F58FC9DA442FB95F98C" box="[894,1043,1558,1584]" name="Madagascar" pageId="5" pageNumber="50">Madagascar</collectingCountry>
|
||
(Nosy-Bé) and Ryukyu Islands (Okinawa) in
|
||
<collectingCountry id="F3087658FF968F58FEDEA46FFE02F9E9" box="[317,388,1595,1621]" name="Japan" pageId="5" pageNumber="50">Japan</collectingCountry>
|
||
; these two geographically remote records suggest that the species should be much more widespread in the Indo-West Pacific.
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF968F58FF26A4D2FD7DF83C" blockId="5.[151,1437,1446,2032]" pageId="5" pageNumber="50">
|
||
<emphasis id="B96BEADAFF968F58FF26A4D2FEB5F91C" bold="true" box="[197,307,1670,1696]" pageId="5" pageNumber="50">Ecology.</emphasis>
|
||
The Nosy-Bé specimens were collected from burrows or mounds of an unknown host on a shallow seagrass-rubble flat, approximately
|
||
<quantity id="4CE79B2DFF968F58FD22A4FFFC96F979" box="[705,784,1707,1733]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="50" unit="m" value="500.0">500 m</quantity>
|
||
from the shore, at a low tide depth of
|
||
<quantity id="4CE79B2DFF968F58FB16A4FFFAE6F979" box="[1269,1376,1707,1733]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="50" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">
|
||
0.5–
|
||
<specimenCount id="9D19FD41FF968F58FAD2A4FFFAE6F979" box="[1329,1376,1707,1733]" pageId="5" pageNumber="50" type="generic">1 m</specimenCount>
|
||
</quantity>
|
||
. The Okinawa specimen was found on sand under a large rock, at a depth of
|
||
<quantity id="4CE79B2DFF968F58FC01A485FB94F957" box="[994,1042,1745,1771]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="50" unit="m" value="8.0">8 m</quantity>
|
||
, according to the collector’s field notes. However, turning over large rocks, especially those deeply embedded in the sediments, often exposes horizontal or oblique sub-surface burrows (A. Anker, pers. obs.); therefore, a possible association of this individual with a larger burrow cannot be excluded. The present evidence suggests that
|
||
<taxonomicName id="4C1F4D4BFF968F58FB7FA515FA9DF8E6" box="[1180,1307,1857,1882]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF968F58FB7FA515FA9DF8E6" box="[1180,1307,1857,1882]" italics="true" pageId="5" pageNumber="50">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF968F58FAC0A514FA07F8E6" bold="true" box="[1315,1409,1856,1882]" pageId="5" pageNumber="50">
|
||
<taxonomicNameLabel id="A25857A1FF968F58FAC0A514FA07F8E6" box="[1315,1409,1856,1882]" pageId="5" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is at least a facultative, burrow-associated commensal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3056543FF968F5AFF26A5DFFA1AF851" lastPageId="7" lastPageNumber="52" pageId="5" pageNumber="50" type="discussion">
|
||
<paragraph id="8BA036C8FF968F5BFF26A5DFFF65FED0" blockId="5.[151,1437,1446,2032]" lastBlockId="6.[151,1436,152,364]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
|
||
<emphasis id="B96BEADAFF968F58FF26A5DFFEC4F819" bold="true" box="[197,322,1931,1957]" pageId="5" pageNumber="50">Remarks.</emphasis>
|
||
Among the 28 species of
|
||
<taxonomicName id="4C1F4D4BFF968F58FD66A5D8FD6EF819" box="[645,744,1932,1957]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEADAFF968F58FD66A5D8FD6EF819" box="[645,744,1932,1957]" italics="true" pageId="5" pageNumber="50">Athanas</emphasis>
|
||
</taxonomicName>
|
||
present in the Indo-West Pacific (
|
||
<bibRefCitation id="EF8E4B39FF968F58FB6FA5DFFA1EF819" author="Anker" box="[1164,1432,1931,1957]" pageId="5" pageNumber="50" refString="Anker, A & Jeng, M. - S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinition of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker and Jeng 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF968F58FF74A5E5FE4CF877" author="Anker" box="[151,458,1969,1995]" pageId="5" pageNumber="50" refString="Anker, A and Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007">Anker and Ahyong 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF968F58FE3BA5E5FD73F877" author="Anker" box="[472,757,1969,1995]" pageId="5" pageNumber="50" refString="Anker, A. & Marin, I. N. (2007) Athanas anatidactylus, n. sp., a new alpheid shrimp associated with crinoids in the tropical western Pacific (Crustacea: Decapoda). Zoological Studies, 46, 162 - 167." type="journal article" year="2007">Anker and Marin 2007</bibRefCitation>
|
||
), only a few species share with
|
||
<taxonomicName id="4C1F4D4BFF968F58FB9DA5E5FA85F876" box="[1150,1283,1969,1994]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF968F58FB9DA5E5FA85F876" box="[1150,1283,1969,1994]" italics="true" pageId="5" pageNumber="50">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF968F58FAEFA5E4FAE8F876" bold="true" box="[1292,1390,1968,1994]" pageId="5" pageNumber="50">
|
||
<taxonomicNameLabel id="A25857A1FF968F58FAEFA5E4FAE8F876" box="[1292,1390,1968,1994]" pageId="5" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the combination of the following features: (1) the frontal margin of the carapace without infracorneal and supracorneal teeth; (2) the chelipeds more or less symmetrical in shape and subequal in size, with slender ischium and non-inflated merus and elongate, vase-shaped carpus; and (3) the third to fifth pereiopods with a simple, long, slender, sickle-shaped dactylus. These species are
|
||
<taxonomicName id="4C1F4D4BFF958F5BFCD6A2B6FA9CFF40" authority="Anker and Ahyong, 2007" authorityName="Anker and Ahyong" authorityYear="2007" box="[821,1306,226,252]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B96BEADAFF958F5BFCD6A2B6FC65FF47" box="[821,995,226,251]" italics="true" pageId="6" pageNumber="51">A. sydneyensis</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF958F5BFC08A2B6FA9CFF40" author="Anker" box="[1003,1306,226,252]" pageId="6" pageNumber="51" refString="Anker, A and Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007">Anker and Ahyong, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
from New South
|
||
<collectingCountry id="F3087658FF958F5BFF07A35CFEA8FE9E" box="[228,302,264,290]" name="United Kingdom" pageId="6" pageNumber="51">Wales</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3087658FF958F5BFED8A35CFE2DFE9E" box="[315,427,264,290]" name="Australia" pageId="6" pageNumber="51">Australia</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C1F4D4BFF958F5BFE54A35CFCA2FE9E" authority="Hayashi, 2002" authorityName="Hayashi" authorityYear="2002" box="[439,804,264,290]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B96BEADAFF958F5BFE54A35CFDF6FE9D" box="[439,624,264,289]" italics="true" pageId="6" pageNumber="51">A. squillophilus</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF958F5BFD9AA35CFCA2FE9E" author="Hayashi" box="[633,804,264,290]" pageId="6" pageNumber="51" refString="Hayashi, K. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3087658FF958F5BFC8EA35CFC32FE9E" box="[877,948,264,290]" name="Japan" pageId="6" pageNumber="51">Japan</collectingCountry>
|
||
; and
|
||
<taxonomicName id="4C1F4D4BFF958F5BFC17A35CFF52FEFB" authority="Banner and Banner, 1983" authorityName="Banner and Banner" authorityYear="1983" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="phyllocheles">
|
||
<emphasis id="B96BEADAFF958F5BFC17A35CFB2EFE9D" box="[1012,1192,264,289]" italics="true" pageId="6" pageNumber="51">A. phyllocheles</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF958F5BFB52A35CFF52FEFB" author="Banner" pageId="6" pageNumber="51" refString="Banner, A. H. & Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner and Banner, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
, a deep-water species from
|
||
<collectingCountry id="F3087658FF958F5BFDFBA379FDFDFEFB" box="[536,635,301,327]" name="Réunion" pageId="6" pageNumber="51">Réunion</collectingCountry>
|
||
Island (
|
||
<bibRefCitation id="EF8E4B39FF958F5BFD3BA379FB86FEFB" author="Banner" box="[728,1024,301,327]" pageId="6" pageNumber="51" refString="Banner, A. H. & Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner and Banner 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF958F5BFBEFA379FB37FEFB" author="Hayashi" box="[1036,1201,301,327]" pageId="6" pageNumber="51" refString="Hayashi, K. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF958F5BFB5FA379FF51FED0" author="Anker" pageId="6" pageNumber="51" refString="Anker, A and Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007">Anker and Ahyong 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF606640FF958F5BFF74A530FAFDF807" httpUri="https://zenodo.org/record/199313/files/figure.png" pageId="6" pageNumber="51" targetBox="[152,1436,463,1834]" targetPageId="6">
|
||
<paragraph id="8BA036C8FF958F5BFF74A530FAFDF807" blockId="6.[151,1436,1892,1979]" pageId="6" pageNumber="51">
|
||
<emphasis id="B96BEADAFF958F5BFF74A530FE9CF8C0" bold="true" box="[151,282,1892,1916]" pageId="6" pageNumber="51">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C1F4D4BFF958F5BFEC3A531FE53F8C0" box="[288,469,1893,1916]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF958F5BFEC3A531FE53F8C0" box="[288,469,1893,1916]" italics="true" pageId="6" pageNumber="51">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF958F5BFE3FA530FD9EF8C0" bold="true" box="[476,536,1892,1916]" pageId="6" pageNumber="51">
|
||
<taxonomicNameLabel id="A25857A1FF958F5BFE3FA530FD9EF8C0" box="[476,536,1892,1916]" pageId="6" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female (CL 2.6 mm), Okinawa (CBM-ZC 10014): A, right major cheliped, mesial; B, same, lateral; C, same, carpus and chela and, mesial (= flexor view); D, same, lateral (= extensor view); E, left minor cheliped, lateral; F, same, mesial; G, carpus and chela, lateral (= extensor view); H, same, chela, mesial (flexor view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF606640FF948F5AFF74A42CFC14F96C" httpUri="https://zenodo.org/record/199314/files/figure.png" pageId="7" pageNumber="52" targetBox="[151,1436,186,1635]" targetPageId="7">
|
||
<paragraph id="8BA036C8FF948F5AFF74A42CFC14F96C" blockId="7.[151,1436,1656,1744]" pageId="7" pageNumber="52">
|
||
<emphasis id="B96BEADAFF948F5AFF74A42CFEA7F92C" bold="true" box="[151,289,1656,1680]" pageId="7" pageNumber="52">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C1F4D4BFF948F5AFECAA42DFE6FF92C" box="[297,489,1657,1680]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF948F5AFECAA42DFE6FF92C" box="[297,489,1657,1680]" italics="true" pageId="7" pageNumber="52">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF948F5AFE12A42CFDB4F92C" bold="true" box="[497,562,1656,1680]" pageId="7" pageNumber="52">
|
||
<taxonomicNameLabel id="A25857A1FF948F5AFE12A42CFDB4F92C" box="[497,562,1656,1680]" pageId="7" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus and colour pattern. A, B, holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): dorsal (A) and lateral (B) [photographs by Arthur Anker]; C, D, specimen from Okinawa (not collected) in situ [photographs by Yusuke Yamada].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA036C8FF948F5AFF26A54DFA1AF851" blockId="7.[151,1437,1816,2029]" pageId="7" pageNumber="52">
|
||
<taxonomicName id="4C1F4D4BFF948F5AFF26A54DFE0DF88E" box="[197,395,1817,1842]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF948F5AFF26A54DFE0DF88E" box="[197,395,1817,1842]" italics="true" pageId="7" pageNumber="52">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF948F5AFE71A54CFE76F88E" bold="true" box="[402,496,1816,1842]" pageId="7" pageNumber="52">
|
||
<taxonomicNameLabel id="A25857A1FF948F5AFE71A54CFE76F88E" box="[402,496,1816,1842]" pageId="7" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C1F4D4BFF948F5AFD68A54DFCB0F88E" box="[651,822,1817,1842]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B96BEADAFF948F5AFD68A54DFCB0F88E" box="[651,822,1817,1842]" italics="true" pageId="7" pageNumber="52">A. sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
by the longer stylocerite, reaching to 0.7–0.8 length of the second segment of antennular peduncle (vs. reaching only to
|
||
<quantity id="4CE79B2DFF948F5AFBD5A56AFB04F8E4" box="[1078,1154,1854,1880]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="52" unit="in" value="0.5">0.5 in</quantity>
|
||
<taxonomicName id="4C1F4D4BFF948F5AFB6CA56AFAD5F8EB" box="[1167,1363,1854,1879]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B96BEADAFF948F5AFB6CA56AFAD5F8EB" box="[1167,1363,1854,1879]" italics="true" pageId="7" pageNumber="52">A. sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
); the proximolaterally non-convex, shorter rostrum, not reaching the distal margin of the first segment of antennular peduncle (vs. proximolaterally convex, reaching the distal margin in
|
||
<taxonomicName id="4C1F4D4BFF948F5AFBB5A5DDFA82F81E" box="[1110,1284,1929,1954]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B96BEADAFF948F5AFBB5A5DDFA82F81E" box="[1110,1284,1929,1954]" italics="true" pageId="7" pageNumber="52">A. sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
); the stouter antennular peduncle, with the second segment as long as wide (vs. longer than wide in
|
||
<taxonomicName id="4C1F4D4BFF948F5AFB4CA5FAFADBF87B" box="[1199,1373,1966,1991]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B96BEADAFF948F5AFB4CA5FAFADBF87B" box="[1199,1373,1966,1991]" italics="true" pageId="7" pageNumber="52">A. sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
); the much shorter cheliped carpus, which is only about 0.5 length of the merus (vs. almost equal to the merus in
|
||
<emphasis id="B96BEADAFF948F5AFA60A580FA1AF851" box="[1411,1436,2004,2029]" italics="true" pageId="7" pageNumber="52">A.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3056543FF9B8F55FF74A2CCFEC3FA70" pageId="8" pageNumber="53" type="reference_group">
|
||
<paragraph id="8BA036C8FF9B8F55FF74A2CCFC4AFE2E" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FF74A2CCFEA3FF0D" ID-CoL="85MD5" box="[151,293,152,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B96BEADAFF9B8F55FF74A2CCFEA3FF0D" box="[151,293,152,177]" italics="true" pageId="8" pageNumber="53">sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
); and the more gaping cheliped fingers, with a different configuration of teeth (see
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FAFDA2CCFEBBFF6B" author="Anker" pageId="8" pageNumber="53" refString="Anker, A and Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007">Anker and Ahyong 2007</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FEB3A2E9FD90FF6A" ID-CoL="85MBH" box="[336,534,189,214]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF9B8F55FEB3A2E9FD90FF6A" box="[336,534,189,214]" italics="true" pageId="8" pageNumber="53">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF9B8F55FDFDA2E9FDFAFF6B" bold="true" box="[542,636,189,215]" pageId="8" pageNumber="53">
|
||
<taxonomicNameLabel id="A25857A1FF9B8F55FDFDA2E9FDFAFF6B" box="[542,636,189,215]" pageId="8" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily separated from
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FC38A2E9FB12FF6A" ID-CoL="85MD2" box="[987,1172,189,214]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B96BEADAFF9B8F55FC38A2E9FB12FF6A" box="[987,1172,189,214]" italics="true" pageId="8" pageNumber="53">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
by the pterygostomial angle being rounded (vs. sharply produced in
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD28A2B6FC0DFF47" ID-CoL="85MD2" box="[715,907,226,251]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B96BEADAFF9B8F55FD28A2B6FC0DFF47" box="[715,907,226,251]" italics="true" pageId="8" pageNumber="53">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
); the cheliped ischium and merus having a dentate ventrolateral margin (vs. smooth in
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD71A35CFCCFFE9D" ID-CoL="85MD2" box="[658,841,264,289]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B96BEADAFF9B8F55FD71A35CFCCFFE9D" box="[658,841,264,289]" italics="true" pageId="8" pageNumber="53">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
); and the setal brush on the fingers of the chela not continuing onto the palm, as is the case in
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD7FA379FCDCFEFA" ID-CoL="85MD2" box="[668,858,301,326]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B96BEADAFF9B8F55FD7FA379FCDCFEFA" box="[668,858,301,326]" italics="true" pageId="8" pageNumber="53">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FC78A379FBC0FEFB" author="Hayashi" box="[923,1094,301,327]" pageId="8" pageNumber="53" refString="Hayashi, K. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
). The new species markedly differs from
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FED1A306FE6BFED7" ID-CoL="85MCS" box="[306,493,338,363]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="phyllocheles">
|
||
<emphasis id="B96BEADAFF9B8F55FED1A306FE6BFED7" box="[306,493,338,363]" italics="true" pageId="8" pageNumber="53">A. phyllocheles</emphasis>
|
||
</taxonomicName>
|
||
by the general shape of the chelipeds and the much more slender antennular peduncle, second and third pereiopods (see
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FD74A32CFC39FE2E" author="Banner" box="[663,959,376,402]" pageId="8" pageNumber="53" refString="Banner, A. H. & Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner and Banner 1983</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF9B8F55FF26A3C9FCD7FD00" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="53">
|
||
All the other Indo-West Pacific species of
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD22A3C9FCA4FE0A" ID-CoL="84LMQ" box="[705,802,413,438]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEADAFF9B8F55FD22A3C9FCA4FE0A" box="[705,802,413,438]" italics="true" pageId="8" pageNumber="53">Athanas</emphasis>
|
||
</taxonomicName>
|
||
either have infracorneal and/or supracorneal teeth or differently shaped chelipeds, and sometimes both (e.g.
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FCF9A396FC3AFE60" author="Pearson" box="[794,956,450,476]" pageId="8" pageNumber="53" refString="Pearson, J. (1905) Report on the Macrura collected by Professor Herdman, at Ceylon in 1902. In: Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Maanar by W. A. Herdman, 4. Royal Society, London, pp. 65 - 92, pls. 1 - 2." type="book chapter" year="1905">Pearson 1905</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FC2BA396FBECFE60" author="De" box="[968,1130,450,476]" pageId="8" pageNumber="53" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1, 133 - 465." type="journal article" year="1911">De Man 1911</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FB95A396FB35FE60" author="Man" box="[1142,1203,450,476]" pageId="8" pageNumber="53" refString="De Man, J. G. (1915) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Supplement. Explanations of plates of Alpheidae. Siboga-Expeditie, 39 a 1, pls. 1 - 23." type="journal article" year="1915">1915</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FB5CA396FB7BFE60" author="Man" box="[1215,1277,450,476]" pageId="8" pageNumber="53" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. Milne Edwards from the Indian Archipelago. Zoologische Mededelingen, 5, 103 - 109." type="journal article" year="1920">1920</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FAEAA396FA11FE60" author="Chace" box="[1289,1431,450,476]" pageId="8" pageNumber="53" refString="Chace, F. A., Jr. (1955) Notes on shrimps from the Marshall Islands. Proceedings of the United States National Museum, 105, 1 - 22." type="journal article" year="1955">Chace 1955</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FF74A3BCFE38FDBE" author="Banner" box="[151,446,488,514]" pageId="8" pageNumber="53" refString="Banner, A. H. & Banner, D. M. (1960) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part V: The Indo-Pacific members of the genus Athanas. Pacific Science, 14, 129 - 155." type="journal article" year="1960">Banner and Banner 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FE29A3BCFD81FDBE" author="Banner" box="[458,519,488,514]" pageId="8" pageNumber="53" refString="Banner, A. H. & Banner, D. M. (1966) The alpheid shrimp of Thailand: the alpheid shrimp of the Gulf of Thailand and adjacent waters. The Siam Society Monographs, 3, i - vi, 1 - 168." type="journal article" year="1966">1966</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FDF7A3BCFDD4FDBE" author="Banner" box="[532,594,488,514]" pageId="8" pageNumber="53" refString="Banner, D. M. & Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382." type="journal article" year="1973">1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FDBDA3BCFD6CFDBE" author="Bruce" box="[606,746,488,514]" pageId="8" pageNumber="53" refString="Bruce, A. J. (1990) Additions to the marine shrimp fauna of Hong Kong. In: Morton, B. (Ed.), Proceedings of the Second International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, 2. Hong Kong University Press, Hong Kong, pp. 611 - 648." type="journal article" year="1990">Bruce 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FD15A3BCFC4EFDBE" author="Anker" box="[758,968,488,514]" pageId="8" pageNumber="53" refString="Anker, A, Jeng, M. - S. & Chan, T. - Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061." type="journal article" year="2001">
|
||
Anker
|
||
<emphasis id="B96BEADAFF9B8F55FCA4A3BCFC04FDBD" box="[839,898,488,513]" italics="true" pageId="8" pageNumber="53">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FC37A3BCFBFFFDBE" author="Hayashi" box="[980,1145,488,514]" pageId="8" pageNumber="53" refString="Hayashi, K. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FB65A3BCFA93FDBE" author="Anker" box="[1158,1301,488,514]" pageId="8" pageNumber="53" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FAC2A3BCFEAEFD9B" author="Anker" pageId="8" pageNumber="53" refString="Anker, A. & Marin, I. N. (2007) Athanas anatidactylus, n. sp., a new alpheid shrimp associated with crinoids in the tropical western Pacific (Crustacea: Decapoda). Zoological Studies, 46, 162 - 167." type="journal article" year="2007">Anker and Marin 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FED6A059FD89FD9B" author="Anker" box="[309,527,525,551]" pageId="8" pageNumber="53" refString="Anker, A., Naderloo, R. & Marin, I. N. (2010) On a new species of the shrimp genus Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Iran. Zootaxa, 2372, 53 - 60." type="journal article" year="2010">
|
||
Anker
|
||
<emphasis id="B96BEADAFF9B8F55FE6BA059FE40FD9A" box="[392,454,525,550]" italics="true" pageId="8" pageNumber="53">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). The chelipeds of
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD15A059FBB5FD9B" ID-CoL="85MD6" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[758,1075,525,551]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="tenuipes">
|
||
<emphasis id="B96BEADAFF9B8F55FD15A059FCFAFD9A" box="[758,892,525,550]" italics="true" pageId="8" pageNumber="53">A. tenuipes</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FC66A059FBB5FD9B" author="De" box="[901,1075,525,551]" pageId="8" pageNumber="53" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous Decapod Crustacea from the " Siboga-Expedition ". Tijdschrift der Nederlandsche Dierkundige Vereeniging, (2), 11, 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FB8CA059FEBCFDF0" ID-CoL="85MBX" authority="Banner and Banner, 1978" authorityName="Banner and Banner" authorityYear="1978" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="B96BEADAFF9B8F55FB8CA059FA8FFD9A" box="[1135,1289,525,550]" italics="true" pageId="8" pageNumber="53">A. gracilipes</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FAF1A059FEBCFDF0" author="Banner" pageId="8" pageNumber="53" refString="Banner, D. M. & A. H. Banner (1978). Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner and Banner, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
are unknown, but these deep-water species differ from
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FC00A066FBE1FDF7" ID-CoL="85MBH" box="[995,1127,562,587]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF9B8F55FC00A066FBE1FDF7" box="[995,1127,562,587]" italics="true" pageId="8" pageNumber="53">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF9B8F55FB8CA066FB32FDF0" bold="true" box="[1135,1204,562,588]" pageId="8" pageNumber="53">
|
||
<taxonomicNameLabel id="A25857A1FF9B8F55FB8CA066FB32FDF0" box="[1135,1204,562,588]" pageId="8" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the presence of small blunt infra-corneal teeth and several other features. For instance
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FC04A00CFBEAFDCD" ID-CoL="85MD6" box="[999,1132,600,625]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="tenuipes">
|
||
<emphasis id="B96BEADAFF9B8F55FC04A00CFBEAFDCD" box="[999,1132,600,625]" italics="true" pageId="8" pageNumber="53">A. tenuipes</emphasis>
|
||
</taxonomicName>
|
||
differs by the longer and more slender rostrum, and
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FE32A029FDE0FD2A" ID-CoL="85MBX" box="[465,614,637,662]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="B96BEADAFF9B8F55FE32A029FDE0FD2A" box="[465,614,637,662]" italics="true" pageId="8" pageNumber="53">A. gracilipes</emphasis>
|
||
</taxonomicName>
|
||
by the much more slender second pereiopod and the distinctly shorter stylocerite (
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FEC3A0F6FE43FD00" author="De" box="[288,453,674,700]" pageId="8" pageNumber="53" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous Decapod Crustacea from the " Siboga-Expedition ". Tijdschrift der Nederlandsche Dierkundige Vereeniging, (2), 11, 287 - 319." type="journal article" year="1910">De Man 1910</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FE31A0F6FD96FD00" author="Man" box="[466,528,674,700]" pageId="8" pageNumber="53" refString="De Man, J. G. (1915) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Supplement. Explanations of plates of Alpheidae. Siboga-Expeditie, 39 a 1, pls. 1 - 23." type="journal article" year="1915">1915</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FDFFA0F6FCC2FD00" author="Banner" box="[540,836,674,700]" pageId="8" pageNumber="53" refString="Banner, D. M. & A. H. Banner (1978). Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner and Banner 1978</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF9B8F55FF26A09CFEE2FBEB" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="53">
|
||
Among the Eastern Atlantic species, only
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD3BA09CFBAAFD5E" ID-CoL="85MBL" authority="Holthuis, 1951" authorityName="Holthuis" authorityYear="1951" box="[728,1068,712,738]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis id="B96BEADAFF9B8F55FD3BA09CFCECFD5D" box="[728,874,712,737]" italics="true" pageId="8" pageNumber="53">A. amazone</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FC91A09CFBAAFD5E" author="Holthuis" box="[882,1068,712,738]" pageId="8" pageNumber="53" refString="Holthuis, L. B. (1951). The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FB88A09CFECCFCBB" ID-CoL="85MC6" authority="Anker and Ahyong, 2007" authorityName="Anker and Ahyong" authorityYear="2007" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ivoiriensis">
|
||
<emphasis id="B96BEADAFF9B8F55FB88A09CFA97FD5D" box="[1131,1297,712,737]" italics="true" pageId="8" pageNumber="53">A. ivoiriensis</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FAF9A09CFECCFCBB" author="Anker" pageId="8" pageNumber="53" refString="Anker, A and Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007">Anker and Ahyong, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
appear to be distantly related to the new species, from which they can be separated by numerous characters.
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FE7BA146FDFFFC97" ID-CoL="85MC6" box="[408,633,786,811]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ivoiriensis">
|
||
<emphasis id="B96BEADAFF9B8F55FE7BA146FDFFFC97" box="[408,633,786,811]" italics="true" pageId="8" pageNumber="53">Athanas ivoiriensis</emphasis>
|
||
</taxonomicName>
|
||
differs by the much shorter antennal carpocerite,
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FB20A146FACAFC97" ID-CoL="85MBL" box="[1219,1356,786,811]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis id="B96BEADAFF9B8F55FB20A146FACAFC97" box="[1219,1356,786,811]" italics="true" pageId="8" pageNumber="53">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a small infracorneal tooth, and both of them by the general shape and armature of the fingers of the chelipeds (cf.
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FE88A109FD91FCCB" author="Holthuis" box="[363,535,861,887]" pageId="8" pageNumber="53" refString="Holthuis, L. B. (1951). The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FDC6A109FCD7FCCB" author="Anker" box="[549,849,861,887]" pageId="8" pageNumber="53" refString="Anker, A and Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007">Anker and Ahyong 2007</bibRefCitation>
|
||
). The new species also differs from all species of
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FF74A1D6FF71FC27" ID-CoL="84LMQ" box="[151,247,898,923]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEADAFF9B8F55FF74A1D6FF71FC27" box="[151,247,898,923]" italics="true" pageId="8" pageNumber="53">Athanas</emphasis>
|
||
</taxonomicName>
|
||
with known colour patterns by its unique, longitudinally red-white striped colour pattern (
|
||
<figureCitation id="13242A4DFF9B8F55FAC6A1D6FAE8FC20" box="[1317,1390,898,924]" captionStart="FIGURE 6" captionStartId="7.[151,259,1656,1680]" captionTargetBox="[151,1436,186,1635]" captionTargetId="figure@7.[151,1436,186,1635]" captionTargetPageId="7" captionText="FIGURE 6. Athanas ahyongi n. sp., habitus and colour pattern. A, B, holotype, female (CL 2.7 mm), Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14093): dorsal (A) and lateral (B) [photographs by Arthur Anker]; C, D, specimen from Okinawa (not collected) in situ [photographs by Yusuke Yamada]." httpUri="https://zenodo.org/record/199314/files/figure.png" pageId="8" pageNumber="53">Fig. 6</figureCitation>
|
||
), in particular the presence of a well-delineated, broad dorsolateral red band running from the antenna and anterior margin of the carapace to the sixth abdominal somite. Most other species of
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FBECA199FBE9FC5A" ID-CoL="84LMQ" box="[1039,1135,973,998]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEADAFF9B8F55FBECA199FBE9FC5A" box="[1039,1135,973,998]" italics="true" pageId="8" pageNumber="53">Athanas</emphasis>
|
||
</taxonomicName>
|
||
have variously spotted or uniform reddish or purplish patterns, with or without a white mediodorsal band (
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FBDDA1A6FAAFFBB0" author="Anker" box="[1086,1321,1010,1036]" pageId="8" pageNumber="53" refString="Anker, A & Jeng, M. - S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinition of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker & Jeng 2007</bibRefCitation>
|
||
), whereas two echinoderm-associated species have a red-white transversely banded pattern (
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FB8DA64CFAC3FB8E" author="Marin" box="[1134,1349,1048,1074]" pageId="8" pageNumber="53" refString="Marin, I. N., Anker, A., Britayev, T. A. & Palmer, R. A. (2005) Symbiosis between the alpheid shrimp, Athanas ornithorhynchus Banner and Banner, 1973 (Crustacea: Decapoda), and the brittle star Macrophiothrix longipeda (Lamarck, 1816) (Echinodermata: Ophiuroidea). Zoological Studies, 44, 234 - 241." type="journal article" year="2005">
|
||
Marin
|
||
<emphasis id="B96BEADAFF9B8F55FB22A64CFB78FB8D" box="[1217,1278,1048,1073]" italics="true" pageId="8" pageNumber="53">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FAB1A64CFED1FBEB" author="Anker" pageId="8" pageNumber="53" refString="Anker, A. & Marin, I. N. (2007) Athanas anatidactylus, n. sp., a new alpheid shrimp associated with crinoids in the tropical western Pacific (Crustacea: Decapoda). Zoological Studies, 46, 162 - 167." type="journal article" year="2007">Anker and Marin 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA036C8FF9B8F55FF26A637FEC3FA70" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FF26A637FE15FBC0" ID-CoL="85MBH" box="[197,403,1123,1148]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF9B8F55FF26A637FE15FBC0" box="[197,403,1123,1148]" italics="true" pageId="8" pageNumber="53">Athanas ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF9B8F55FE78A636FE7BFBC0" bold="true" box="[411,509,1122,1148]" pageId="8" pageNumber="53">
|
||
<taxonomicNameLabel id="A25857A1FF9B8F55FE78A636FE7BFBC0" box="[411,509,1122,1148]" pageId="8" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the fifth known species of
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FC8FA637FC49FBC0" ID-CoL="84LMQ" box="[876,975,1123,1148]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEADAFF9B8F55FC8FA637FC49FBC0" box="[876,975,1123,1148]" italics="true" pageId="8" pageNumber="53">Athanas</emphasis>
|
||
</taxonomicName>
|
||
found in association with burrows of larger animals. The other species are
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD98A6DCFCDAFB1D" ID-CoL="85MBL" box="[635,860,1160,1185]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis id="B96BEADAFF9B8F55FD98A6DCFCDAFB1D" box="[635,860,1160,1185]" italics="true" pageId="8" pageNumber="53">Athanas amazone</emphasis>
|
||
</taxonomicName>
|
||
, found on a few occasions in burrows of the stomatopod
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FEC4A6F9FE56FB7A" box="[295,464,1197,1222]" class="Malacostraca" family="Squillidae" genus="Squilla" kingdom="Animalia" order="Stomatopoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="mantis">
|
||
<emphasis id="B96BEADAFF9B8F55FEC4A6F9FE56FB7A" box="[295,464,1197,1222]" italics="true" pageId="8" pageNumber="53">Squilla mantis</emphasis>
|
||
</taxonomicName>
|
||
(L) in the Mediterranean Sea (
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FCDEA6F9FB06FB7B" author="Froglia" box="[829,1152,1197,1223]" pageId="8" pageNumber="53" refString="Froglia, C. and Atkinson, R. J. A. (1998) Association between Athanas amazone (Decapoda: Alpheidae) and Squilla mantis (Stomatopoda: Squillidae). Journal of Crustacean Biology, 18, 529 - 532." type="journal article" year="1998">Froglia and Atkinson 1998</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FB76A6F9FA12FB7A" ID-CoL="85MD2" box="[1173,1428,1197,1222]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B96BEADAFF9B8F55FB76A6F9FA12FB7A" box="[1173,1428,1197,1222]" italics="true" pageId="8" pageNumber="53">Athanas squillophilus</emphasis>
|
||
</taxonomicName>
|
||
, associated with burrows of the stomatopod
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FD58A687FBDEFB50" authority="De Haan" authorityName="De Haan" box="[699,1112,1234,1260]" class="Malacostraca" family="Squillidae" genus="Oratosquilla" kingdom="Animalia" order="Stomatopoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="oratoria">
|
||
<emphasis id="B96BEADAFF9B8F55FD58A687FC4AFB50" box="[699,972,1235,1260]" italics="true" pageId="8" pageNumber="53">Oratosquilla oratoria</emphasis>
|
||
(De Haan)
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F3087658FF9B8F55FB61A686FB4FFB50" box="[1154,1225,1234,1260]" name="Japan" pageId="8" pageNumber="53">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FB39A686FA09FB50" author="Hayashi" box="[1242,1423,1234,1260]" pageId="8" pageNumber="53" refString="Hayashi, K. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FF74A6ACFD5EFAAE" ID-CoL="85MBQ" authority="Anker, Jeng and Chan, 2001" authorityName="Anker, Jeng and Chan" authorityYear="2001" box="[151,728,1272,1298]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="dentirostris">
|
||
<emphasis id="B96BEADAFF9B8F55FF74A6ACFE02FAAD" box="[151,388,1272,1297]" italics="true" pageId="8" pageNumber="53">Athanas dentirostris</emphasis>
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FE68A6ACFD5EFAAE" author="Anker" box="[395,728,1272,1298]" pageId="8" pageNumber="53" refString="Anker, A, Jeng, M. - S. & Chan, T. - Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061." type="journal article" year="2001">Anker, Jeng and Chan, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, of which the single known specimen was found in a burrow of the mud-shrimp
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FE67A749FDA3FA8A" ID-CoL="da4878fc-faef-4116-bf65-5ba9f84fbaa2" box="[388,549,1309,1334]" class="Malacostraca" family="Upogebiidae" genus="Austinogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEADAFF9B8F55FE67A749FDA3FA8A" box="[388,549,1309,1334]" italics="true" pageId="8" pageNumber="53">Austinogebia</emphasis>
|
||
</taxonomicName>
|
||
aff.
|
||
<emphasis id="B96BEADAFF9B8F55FD82A749FD64FA8A" box="[609,738,1309,1334]" italics="true" pageId="8" pageNumber="53">takaoensis</emphasis>
|
||
(Sakai and Türkay) in
|
||
<collectingCountry id="F3087658FF9B8F55FBE1A749FBEFFA8B" box="[1026,1129,1309,1335]" name="Vietnam" pageId="8" pageNumber="53">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FB99A749FADCFA8B" author="Anker" box="[1146,1370,1309,1335]" pageId="8" pageNumber="53" refString="Anker, A, Jeng, M. - S. & Chan, T. - Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061." type="journal article" year="2001">
|
||
Anker
|
||
<emphasis id="B96BEADAFF9B8F55FB32A749FA89FA8A" box="[1233,1295,1309,1334]" italics="true" pageId="8" pageNumber="53">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
); and
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FF74A717FE4AFAE0" ID-CoL="85MC7" authority="Kubo" authorityName="Kubo" box="[151,460,1346,1372]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="B96BEADAFF9B8F55FF74A717FEFBFAE0" box="[151,381,1347,1372]" italics="true" pageId="8" pageNumber="53">Athanas japonicus</emphasis>
|
||
<collectingCountry id="F3087658FF9B8F55FE66A716FE4AFAE0" box="[389,460,1346,1372]" name="Cuba" pageId="8" pageNumber="53">Kubo</collectingCountry>
|
||
</taxonomicName>
|
||
, occasionally associated with mudskippers,
|
||
<emphasis id="B96BEADAFF9B8F55FC16A717FB4BFAE0" box="[1013,1229,1347,1372]" italics="true" pageId="8" pageNumber="53">Periphthalmodon</emphasis>
|
||
sp., in
|
||
<collectingCountry id="F3087658FF9B8F55FAC8A716FA1BFAE0" box="[1323,1437,1346,1372]" name="Australia" pageId="8" pageNumber="53">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8E4B39FF9B8F55FF7CA73CFEBBFA3E" author="Anker" box="[159,317,1384,1410]" pageId="8" pageNumber="53" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker 2003</bibRefCitation>
|
||
) and other burrowing hosts (A. Anker, unpubl. data). The nature of these associations (facultative vs. obligate, mutualism vs. commensalism) as well as the hosts of
|
||
<taxonomicName id="4C1F4D4BFF9B8F55FBDFA7D9FB3AFA1A" ID-CoL="85MBH" box="[1084,1212,1421,1446]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ahyongi" status="sp. nov.">
|
||
<emphasis id="B96BEADAFF9B8F55FBDFA7D9FB3AFA1A" box="[1084,1212,1421,1446]" italics="true" pageId="8" pageNumber="53">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADAFF9B8F55FB27A7D9FAA5FA1B" bold="true" box="[1220,1315,1421,1447]" pageId="8" pageNumber="53">
|
||
<taxonomicNameLabel id="A25857A1FF9B8F55FB27A7D9FAA5FA1B" box="[1220,1315,1421,1447]" pageId="8" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
remain to be determined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |