357 lines
37 KiB
XML
357 lines
37 KiB
XML
<document id="32EAE1FC4BE5A168A6399D3AA7532420" ID-DOI="10.11646/zootaxa.3957.4.1" ID-GBIF-Dataset="9cdf029d-f4fe-415b-9d55-501bd3369f78" ID-ISSN="1175-5326" ID-Zenodo-Dep="239589" ID-ZooBank="90175CBC-5C93-44CD-B13C-B17516643317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461062920496" checkinUser="plazi" docAuthor="Pfingstl, Tobias" docDate="2015" docId="03B687D92832F477D3EEFD6FD1BB9B3B" docLanguage="en" docName="zt03957p382.pdf" docOrigin="Zootaxa 3957 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fortuynia maledivensis Pfingstl, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="357" masterDocId="FF8FFFA12830F471D379FF9DD3269D20" masterDocTitle="The intertidal Fortuyniidae (Acari: Oribatida): new species, morphological diversity, ecology and biogeography" masterLastPageNumber="382" masterPageNumber="351" pageNumber="353" updateTime="1698582922211" updateUser="plazi">
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<mods:mods id="144D4A53C03663251D22CE27684CDE8E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9445FCAF2FEE2C593627C99A18A2FC95">
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<mods:title id="349C60E6FAFEED8255DB93A14D60C483">The intertidal Fortuyniidae (Acari: Oribatida): new species, morphological diversity, ecology and biogeography</mods:title>
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</mods:titleInfo>
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<mods:name id="A3814153C0F4E862EC0EE65C7BF8CCC8" type="personal">
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<mods:roleTerm id="C21A23571703713AB9CF4C73C6D54688">Author</mods:roleTerm>
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<mods:namePart id="11B490F9CA2D953810CD5E748FBDDA51">Pfingstl, Tobias</mods:namePart>
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<mods:typeOfResource id="968E4C47C7807B0B2F34A1517D1FF3A0">text</mods:typeOfResource>
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<mods:title id="DDFFBEE3AF0A23BBA15461F78CACF049">Zootaxa</mods:title>
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<mods:part id="64709150C31AAE8C7232E23F5B21D20B">
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<mods:date id="4683794E13FABE7D71877EE4250E29E8">2015</mods:date>
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<mods:detail id="5129D570BDED3BD72FCAB139D5D3387F" type="volume">
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<mods:number id="9DD240B7C3AC0456B91F33DE07EF1FC4">3957</mods:number>
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</mods:detail>
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<mods:detail id="73444B98A85D375C267D3EA00C446425" type="issue">
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<mods:number id="FB67E0DDD2E85467CF122BF31F2DF165">4</mods:number>
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<mods:classification id="91D68DA5A8CEB25F490DD57C7794B054">journal article</mods:classification>
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<mods:identifier id="9EA3B886830A44F6B475B0C6C45FFAD8" type="DOI">10.11646/zootaxa.3957.4.1</mods:identifier>
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<mods:identifier id="2945F8217B479C139E917AA593C3389B" type="GBIF-Dataset">9cdf029d-f4fe-415b-9d55-501bd3369f78</mods:identifier>
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<mods:identifier id="48EC0A6585E6DC5FEF2ADD43D6C0ED31" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="246928E54422EC04FF7A1326316F7ADE" type="Zenodo-Dep">239589</mods:identifier>
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<mods:identifier id="931B1F2C2F9635A1E48D1FDF16AE478A" type="ZooBank">90175CBC-5C93-44CD-B13C-B17516643317</mods:identifier>
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</mods:mods>
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<treatment id="03B687D92832F477D3EEFD6FD1BB9B3B" ID-DOI="http://doi.org/10.5281/zenodo.6113654" ID-GBIF-Taxon="119614615" ID-Zenodo-Dep="6113654" LSID="urn:lsid:plazi:treatment:03B687D92832F477D3EEFD6FD1BB9B3B" httpUri="http://treatment.plazi.org/id/03B687D92832F477D3EEFD6FD1BB9B3B" lastPageId="6" lastPageNumber="357" pageId="2" pageNumber="353">
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<subSubSection id="C30565442832F473D3EEFD6FD13E9E2C" box="[151,536,754,780]" pageId="2" pageNumber="353" type="nomenclature">
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<paragraph id="8BA036CF2832F473D3EEFD6FD13E9E2C" blockId="2.[151,536,754,780]" box="[151,536,754,780]" pageId="2" pageNumber="353">
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<heading id="D0E881A32832F473D3EEFD6FD13E9E2C" bold="true" box="[151,536,754,780]" fontSize="11" level="1" pageId="2" pageNumber="353" reason="1">
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<emphasis id="B96BEADD2832F473D3EEFD6FD13E9E2C" bold="true" box="[151,536,754,780]" pageId="2" pageNumber="353">
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<taxonomicName id="4C1F4D4C2832F473D3EEFD6FD2959E2C" ID-CoL="8T7LK" box="[151,435,754,780]" class="Arachnida" family="Fortuyniidae" genus="Fortuynia" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="maledivensis" status="sp. nov.">
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<emphasis id="B96BEADD2832F473D3EEFD6FD2959E2C" bold="true" box="[151,435,754,780]" italics="true" pageId="2" pageNumber="353">Fortuynia maledivensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25857A62832F473D2C3FD6FD13E9E2C" box="[442,536,754,780]" pageId="2" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565442832F473D3EEFCA4D2EE9E9E" pageId="2" pageNumber="353" type="materials_examined">
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||
<paragraph id="8BA036CF2832F473D3EEFCA4D2EE9E9E" blockId="2.[151,1437,825,1942]" pageId="2" pageNumber="353">
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<emphasis id="B96BEADD2832F473D3EEFCA4D21B9E72" bold="true" box="[151,317,825,850]" pageId="2" pageNumber="353">
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<typeStatus id="54A4886D2832F473D3EEFCA4D3F49E72" box="[151,210,825,850]" pageId="2" pageNumber="353">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="54A4886D2832F473D232FCA4D29E9E72" box="[331,440,825,850]" pageId="2" pageNumber="353" type="holotype">Holotype</typeStatus>
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: female,
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<collectingCountry id="F308765F2832F473D162FCA4D1A09E72" box="[539,646,825,850]" name="Maldives" pageId="2" pageNumber="353">Maldives</collectingCountry>
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, Malé atoll, Island of Villingili;
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<date id="FFA1100F2832F473D094FCA7D7539E72" box="[1005,1141,825,850]" pageId="2" pageNumber="353" value="1983-02-08">8 Feb. 1983</date>
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; calcareous algae growing on sandstone; coll. W. Fiala, leg. R. Schuster.
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<typeStatus id="54A4886D2832F473D1E4FCC3D0289E56" box="[669,782,862,886]" pageId="2" pageNumber="353" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D19FD462832F473D060FCC3D0AE9E56" box="[793,904,861,886]" pageId="2" pageNumber="353" type="female">2 females</specimenCount>
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, same locality as
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<typeStatus id="54A4886D2832F473D736FCC0D7939E56" box="[1103,1205,861,886]" pageId="2" pageNumber="353" type="holotype">holotype</typeStatus>
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. Deposition of holo- and
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<typeStatus id="54A4886D2832F473D3B1FC1FD21E9EBA" box="[200,312,898,922]" pageId="2" pageNumber="353" type="paratype">paratypes</typeStatus>
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: Naturhistorisches Museum Wien/NHM Vienna,
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<collectingCountry id="F308765F2832F473D010FC1CD0999EBA" box="[873,959,897,922]" name="Austria" pageId="2" pageNumber="353">Austria</collectingCountry>
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;
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<typeStatus id="54A4886D2832F473D0B5FC1CD7159EBA" box="[972,1075,897,922]" pageId="2" pageNumber="353" type="holotype">holotype</typeStatus>
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: collection Nr.
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<collectionCode id="ED0EAE0A2832F473D79AFC1FD6199EB9" box="[1251,1343,898,921]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="353">NHMW</collectionCode>
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27.590;
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<typeStatus id="54A4886D2832F473D3EEFC3BD2219E9E" box="[151,263,934,958]" pageId="2" pageNumber="353" type="paratype">paratypes</typeStatus>
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:
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<collectionCode id="ED0EAE0A2832F473D268FC3BD24B9E9D" box="[273,365,934,957]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="353">NHMW</collectionCode>
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27.591.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565442832F473D3BEFC54D7029EC2" box="[199,1060,969,994]" pageId="2" pageNumber="353" type="etymology">
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<paragraph id="8BA036CF2832F473D3BEFC54D7029EC2" blockId="2.[151,1437,825,1942]" box="[199,1060,969,994]" pageId="2" pageNumber="353">
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<emphasis id="B96BEADD2832F473D3BEFC54D2689EC2" bold="true" box="[199,334,969,994]" pageId="2" pageNumber="353">Etymology.</emphasis>
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The specific name refers to the collection locality, the
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<collectingCountry id="F308765F2832F473D0CDFC54D7069EC2" box="[948,1056,969,994]" name="Maldives" pageId="2" pageNumber="353">Maldives</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565442832F473D3BEFC70D0389971" pageId="2" pageNumber="353" type="diagnosis">
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<paragraph id="8BA036CF2832F473D3BEFC70D0389971" blockId="2.[151,1437,825,1942]" pageId="2" pageNumber="353">
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<emphasis id="B96BEADD2832F473D3BEFC70D2679926" bold="true" box="[199,321,1005,1030]" pageId="2" pageNumber="353">Diagnosis.</emphasis>
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Average length 400 µm, mean width 225 µm. Habitus typical for genus
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<taxonomicName id="4C1F4D4C2832F473D7E6FC73D6299925" box="[1183,1295,1006,1029]" class="Arachnida" family="Fortuyniidae" genus="Fortuynia" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="353" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEADD2832F473D7E6FC73D6299925" box="[1183,1295,1006,1029]" italics="true" pageId="2" pageNumber="353">Fortuynia</emphasis>
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</taxonomicName>
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. Notogaster slender, oval in shape. Sensilla short, clavate, bent inwards. Prodorsal canal
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<emphasis id="B96BEADD2832F473D090FB8FD0DB9909" box="[1001,1021,1042,1065]" italics="true" pageId="2" pageNumber="353">ci</emphasis>
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absent. Notogastral setae 14 pairs,
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<emphasis id="B96BEADD2832F473D6F1FB8FD6BA990E" box="[1416,1436,1042,1070]" italics="true" pageId="2" pageNumber="353">c3</emphasis>
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absent. Epimeral setation 3-1-3-2. Aggenital setae absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565442832F477D3BEFBC1D1BB9B3B" lastPageId="6" lastPageNumber="357" pageId="2" pageNumber="353" type="description">
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<paragraph id="8BA036CF2832F473D3BEFBC1D2AF99B9" blockId="2.[151,1437,825,1942]" pageId="2" pageNumber="353">
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<emphasis id="B96BEADD2832F473D3BEFBC1D2989954" bold="true" box="[199,446,1116,1141]" pageId="2" pageNumber="353">Description of adult.</emphasis>
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Females (N=15), length: 384–422 µm (mean 400 µm), width: 209–255 µm (mean 225 µm); no males found.
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</paragraph>
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<paragraph id="8BA036CF2832F473D3BEFB38D750999D" blockId="2.[151,1437,825,1942]" box="[199,1142,1188,1213]" pageId="2" pageNumber="353">Integument. Colour dark brown. Cuticle appears shiny under dissecting microscope.</paragraph>
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<paragraph id="8BA036CF2832F473D3BEFB55D6259851" blockId="2.[151,1437,825,1942]" pageId="2" pageNumber="353">
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Prodorsum. Cerotegument finely granular. Cuticle dark brown laterally, median interlamellar area lighter coloured. Rostrum triangular in dorsal view, slightly projecting anteroventrally in lateral view. Whole rostrum demarcated from remainder of prodorsum by transverse ridge. Rostral setae (
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<emphasis id="B96BEADD2832F473D76CFA8CD70A9808" box="[1045,1068,1297,1320]" italics="true" pageId="2" pageNumber="353">ro</emphasis>
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) simple, long (approx. 30 µm), lamellar setae (
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<emphasis id="B96BEADD2832F473D23BFAA9D270986C" box="[322,342,1332,1356]" italics="true" pageId="2" pageNumber="353">le</emphasis>
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) thin, shorter (approx. 20 µm). Interlamellar setae (
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<emphasis id="B96BEADD2832F473D0E3FAA8D096986C" box="[922,944,1333,1356]" italics="true" pageId="2" pageNumber="353">in</emphasis>
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) minute, short,exobothridial setae (
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<emphasis id="B96BEADD2832F473D639FAA8D67F986C" box="[1344,1369,1333,1356]" italics="true" pageId="2" pageNumber="353">ex</emphasis>
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) fine. Bothridia small cups, orifice narrow. Sensilla short, smooth, clavate with rounded head bent inwards.
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</paragraph>
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<paragraph id="8BA036CF2832F473D3BEFAE1D2419BB1" blockId="2.[151,1437,825,1942]" pageId="2" pageNumber="353">
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Gnathosoma. Pedipalps pentamerous 0-2-1-3-9 (including solenidion). Solenidion
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<emphasis id="B96BEADD2832F473D7EDFAE1D78198B3" bold="true" box="[1172,1191,1404,1427]" italics="true" pageId="2" pageNumber="353">ω</emphasis>
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on tarsus erect, not associated with eupathidium
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<emphasis id="B96BEADD2832F473D294FA3CD13D9898" box="[493,539,1441,1464]" italics="true" pageId="2" pageNumber="353">acm</emphasis>
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(
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<figureCitation id="13242A4A2832F473D149FA3DD1539898" box="[560,629,1440,1465]" captionStart="FIGURE 1" captionStartId="3.[151,250,879,901]" captionTargetBox="[234,1354,205,851]" captionTargetId="figure@3.[225,1362,193,858]" captionTargetPageId="3" captionText="FIGURE 1. Fortuynia maledivensis sp. nov. adult. A) right pedipalp, paraxial view. B) right rutellum, ventral view." httpUri="https://zenodo.org/record/239590/files/figure.png" pageId="2" pageNumber="353">Fig. 1</figureCitation>
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A). Large porose area on paraxial ventral side of femur. Chelicerae chelate, movable digit with 3 small, blunt but distinct teeth, whereas from frontal view most distal teeth split into 2 symmetrical teeth; fixed digit with 2 teeth, all teeth interlocking. Large lateral porose area from middle of chelicerae to joint of digits. Setae
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<emphasis id="B96BEADD2832F473D160F991D1679B04" box="[537,577,1548,1572]" italics="true" pageId="2" pageNumber="353">cha</emphasis>
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,
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<emphasis id="B96BEADD2832F473D129F991D15E9B04" box="[592,632,1548,1572]" italics="true" pageId="2" pageNumber="353">chb</emphasis>
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approximately the same length, both dorsally slightly pectinate. Genae well sclerotized, finely granular. Distal part of each rutellum developed as thin triangular slightly curved inward membrane (
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<figureCitation id="13242A4A2832F473D264F9C9D27B9B4C" box="[285,349,1620,1645]" captionStart="FIGURE 1" captionStartId="3.[151,250,879,901]" captionTargetBox="[234,1354,205,851]" captionTargetId="figure@3.[225,1362,193,858]" captionTargetPageId="3" captionText="FIGURE 1. Fortuynia maledivensis sp. nov. adult. A) right pedipalp, paraxial view. B) right rutellum, ventral view." httpUri="https://zenodo.org/record/239590/files/figure.png" pageId="2" pageNumber="353">Fig. 1</figureCitation>
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B) with longitudinal incision. Setae
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<emphasis id="B96BEADD2832F473D18EF9C8D0239B4C" box="[759,773,1621,1644]" italics="true" pageId="2" pageNumber="353">a</emphasis>
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and
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<emphasis id="B96BEADD2832F473D045F9C8D0769B4C" box="[828,848,1621,1644]" italics="true" pageId="2" pageNumber="353">m</emphasis>
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long, smooth. Mentum regular, finely granular, seta
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<emphasis id="B96BEADD2832F473D3EEF9E5D3839BB0" box="[151,165,1656,1680]" italics="true" pageId="2" pageNumber="353">h</emphasis>
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simple and long.
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</paragraph>
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<paragraph id="8BA036CF2832F473D3BEF901D6199AB1" blockId="2.[151,1437,825,1942]" pageId="2" pageNumber="353">
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Gastronotic region (
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<figureCitation id="13242A4A2832F473D2CAF901D2DA9B94" box="[435,508,1692,1717]" captionStart="FIGURE 2. F" captionStartId="4.[151,250,1905,1927]" captionTargetBox="[196,1381,201,1873]" captionTargetId="figure@4.[188,1398,193,1884]" captionTargetPageId="4" captionText="FIGURE 2. F. maledivensis sp. nov. adult. A) dorsal view. B) ventral view. C) lateral view." httpUri="https://zenodo.org/record/239591/files/figure.png" pageId="2" pageNumber="353">Fig. 2</figureCitation>
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A). Notogaster slender and oval in dorsal view. Cerotegument finely granular. Lenticulus more or less rectangular with irregular borders. Notogastral setae 14 pairs, thin, simple (length 16–50 µm),
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<emphasis id="B96BEADD2832F473D3AFF978D3D19A21" box="[214,247,1765,1793]" italics="true" pageId="2" pageNumber="353">c1-2</emphasis>
|
||
,
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||
<emphasis id="B96BEADD2832F473D27EF979D2059BDC" box="[263,291,1764,1788]" italics="true" pageId="2" pageNumber="353">da</emphasis>
|
||
,
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<emphasis id="B96BEADD2832F473D24DF979D2709BDC" box="[308,342,1764,1788]" italics="true" pageId="2" pageNumber="353">dm</emphasis>
|
||
,
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||
<emphasis id="B96BEADD2832F473D21FF979D2A49BDC" box="[358,386,1764,1788]" italics="true" pageId="2" pageNumber="353">dp</emphasis>
|
||
,
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||
<emphasis id="B96BEADD2832F473D2EAF979D28F9BDC" box="[403,425,1764,1788]" italics="true" pageId="2" pageNumber="353">la</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2832F473D2C0F979D2F39BDC" box="[441,469,1764,1788]" italics="true" pageId="2" pageNumber="353">lm</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2832F473D29CF979D2DD9BDC" box="[485,507,1764,1788]" italics="true" pageId="2" pageNumber="353">lp</emphasis>
|
||
,
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||
<emphasis id="B96BEADD2832F473D172F979D1089A21" box="[523,558,1764,1793]" italics="true" pageId="2" pageNumber="353">h1-3</emphasis>
|
||
,
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||
<emphasis id="B96BEADD2832F473D147F978D1469A21" box="[574,608,1765,1793]" italics="true" pageId="2" pageNumber="353">p1-3</emphasis>
|
||
;
|
||
<emphasis id="B96BEADD2832F473D108F978D1A09A21" box="[625,646,1765,1793]" italics="true" pageId="2" pageNumber="353">c3</emphasis>
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absent. Inconspicuous circular porose areas associated with bases of notogastral setae. Three pairs of single pores arranged in 2 median rows, first pair posterior to setae
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<emphasis id="B96BEADD2832F473D65BF896D6189A03" box="[1314,1342,1803,1827]" italics="true" pageId="2" pageNumber="353">da</emphasis>
|
||
, second between setae
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||
<emphasis id="B96BEADD2832F473D238F8B2D2459A67" box="[321,355,1839,1863]" italics="true" pageId="2" pageNumber="353">dm</emphasis>
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and
|
||
<emphasis id="B96BEADD2832F473D2D9F8B2D29A9A67" box="[416,444,1839,1863]" italics="true" pageId="2" pageNumber="353">dp</emphasis>
|
||
and third posterior to
|
||
<emphasis id="B96BEADD2832F473D1BEF8B2D1C59A67" box="[711,739,1839,1863]" italics="true" pageId="2" pageNumber="353">dp</emphasis>
|
||
, all hardly discernible. Five pairs of notogastral lyrifissures present;
|
||
<emphasis id="B96BEADD2832F473D383F8C9D2369A4B" box="[250,272,1876,1899]" italics="true" pageId="2" pageNumber="353">ia</emphasis>
|
||
next to anterior border of notogaster and seta
|
||
<emphasis id="B96BEADD2832F473D04AF8C9D0619A50" box="[819,839,1876,1904]" italics="true" pageId="2" pageNumber="353">c2</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2832F473D021F8C9D0529A4B" box="[856,884,1876,1899]" italics="true" pageId="2" pageNumber="353">im</emphasis>
|
||
posterior of seta
|
||
<emphasis id="B96BEADD2832F473D73DF8CED7469A4B" box="[1092,1120,1875,1899]" italics="true" pageId="2" pageNumber="353">lm</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2832F473D708F8CED7A19A4B" box="[1137,1159,1875,1899]" italics="true" pageId="2" pageNumber="353">ih</emphasis>
|
||
laterally,
|
||
<emphasis id="B96BEADD2832F473D784F8C9D6359A4B" box="[1277,1299,1876,1899]" italics="true" pageId="2" pageNumber="353">ip</emphasis>
|
||
laterad and anterior of seta
|
||
<emphasis id="B96BEADD2832F473D23FF8E7D27A9AB6" box="[326,348,1914,1942]" italics="true" pageId="2" pageNumber="353">p3</emphasis>
|
||
and ips next to
|
||
<emphasis id="B96BEADD2832F473D174F8E7D1029AB6" box="[525,548,1914,1942]" italics="true" pageId="2" pageNumber="353">p2</emphasis>
|
||
. Orifice of opisthonotal gland
|
||
<emphasis id="B96BEADD2832F473D005F8E7D0869AB2" box="[892,928,1914,1938]" italics="true" pageId="2" pageNumber="353">gla</emphasis>
|
||
laterally and between seta
|
||
<emphasis id="B96BEADD2832F473D7A9F8E7D7CA9AB2" box="[1232,1260,1914,1938]" italics="true" pageId="2" pageNumber="353">lm</emphasis>
|
||
and
|
||
<emphasis id="B96BEADD2832F473D65AF8E7D61F9AB2" box="[1315,1337,1914,1938]" italics="true" pageId="2" pageNumber="353">lp</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF6066472833F472D3EEFCF2D6099EA5" box="[151,1327,879,902]" httpUri="https://zenodo.org/record/239590/files/figure.png" pageId="3" pageNumber="354" targetBox="[234,1354,205,851]" targetPageId="3">
|
||
<paragraph id="8BA036CF2833F472D3EEFCF2D6099EA5" blockId="3.[151,1327,879,902]" box="[151,1327,879,902]" pageId="3" pageNumber="354">
|
||
<emphasis id="B96BEADD2833F472D3EEFCF2D2359EA4" bold="true" box="[151,275,879,901]" pageId="3" pageNumber="354">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C1F4D4C2833F472D260FCECD1219EA5" box="[281,519,879,902]" class="Arachnida" family="Fortuyniidae" genus="Fortuynia" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="maledivensis" status="sp. nov.">
|
||
<emphasis id="B96BEADD2833F472D260FCECD1219EA5" box="[281,519,879,902]" italics="true" pageId="3" pageNumber="354">Fortuynia maledivensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADD2833F472D174FCEDD1789EA5" bold="true" box="[525,606,880,901]" pageId="3" pageNumber="354">
|
||
<taxonomicNameLabel id="A25857A62833F472D174FCEDD1789EA5" box="[525,606,880,901]" pageId="3" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
adult. A) right pedipalp, paraxial view. B) right rutellum, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA036CF2833F472D3BEFC31D12F9911" blockId="3.[151,1437,940,2018]" pageId="3" pageNumber="354">
|
||
Lateral aspect (
|
||
<figureCitation id="13242A4A2833F472D202FC31D2E59EE5" box="[379,451,940,965]" captionStart="FIGURE 2. F" captionStartId="4.[151,250,1905,1927]" captionTargetBox="[196,1381,201,1873]" captionTargetId="figure@4.[188,1398,193,1884]" captionTargetPageId="4" captionText="FIGURE 2. F. maledivensis sp. nov. adult. A) dorsal view. B) ventral view. C) lateral view." httpUri="https://zenodo.org/record/239591/files/figure.png" pageId="3" pageNumber="354">Fig. 2</figureCitation>
|
||
C). Cerotegument finely granular, larger granules on pedotectum I and in acetabular regions. Pedotectum I present, rounded but small. Cuticular canals of van der Hammen’s organ typical for genus, whereas
|
||
<emphasis id="B96BEADD2833F472D279FC6BD23F992D" box="[256,281,1014,1037]" italics="true" pageId="3" pageNumber="354">ce</emphasis>
|
||
very short, only reaching anterior border of bothridium and lateral canal with posterior internal enlargement close to bothridium.
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2833F472D3BEFBA0D10D9832" blockId="3.[151,1437,940,2018]" pageId="3" pageNumber="354">
|
||
Ventral region of idiosoma (
|
||
<figureCitation id="13242A4A2833F472D173FBA0D1769975" box="[522,592,1085,1110]" captionStart="FIGURE 2. F" captionStartId="4.[151,250,1905,1927]" captionTargetBox="[196,1381,201,1873]" captionTargetId="figure@4.[188,1398,193,1884]" captionTargetPageId="4" captionText="FIGURE 2. F. maledivensis sp. nov. adult. A) dorsal view. B) ventral view. C) lateral view." httpUri="https://zenodo.org/record/239591/files/figure.png" pageId="3" pageNumber="354">Fig. 2</figureCitation>
|
||
B). Cerotegument finely granular in sternal region; larger, densely packed granules laterally next to acetabula. Epimeral setation 3-1-3-2, all setae simple and short. Genital setae 5 pairs. Aggenital setae absent. Anal valves slightly triangular, 2 pairs of long anal setae
|
||
<emphasis id="B96BEADD2833F472D73BFB1BD7549982" box="[1090,1138,1158,1186]" italics="true" pageId="3" pageNumber="354">an1-2</emphasis>
|
||
. Preanal organ triangular. Adanal setae
|
||
<emphasis id="B96BEADD2833F472D254FB31D27899E8" box="[301,350,1196,1224]" italics="true" pageId="3" pageNumber="354">an1-3</emphasis>
|
||
3 pairs, simple, medium sized; rare variation with 2 setae unilaterally observed. Seta
|
||
<emphasis id="B96BEADD2833F472D661FB31D61A99E8" box="[1304,1340,1196,1224]" italics="true" pageId="3" pageNumber="354">ad3</emphasis>
|
||
on level with anterior border of anal opening,
|
||
<emphasis id="B96BEADD2833F472D117FB4FD1B499CF" box="[622,658,1234,1263]" italics="true" pageId="3" pageNumber="354">ad2</emphasis>
|
||
laterad,
|
||
<emphasis id="B96BEADD2833F472D07CFB4FD00F99CF" box="[773,809,1234,1263]" italics="true" pageId="3" pageNumber="354">ad1</emphasis>
|
||
posterior of anal valves. Lyrifissure
|
||
<emphasis id="B96BEADD2833F472D781FB4FD63A99CA" box="[1272,1308,1234,1258]" italics="true" pageId="3" pageNumber="354">iad</emphasis>
|
||
orientated longitudinally, flanking anal orifice.
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2833F472D3BEFA83D1A79B2E" blockId="3.[151,1437,940,2018]" pageId="3" pageNumber="354">
|
||
Legs (
|
||
<figureCitation id="13242A4A2833F472D276FA80D2709816" box="[271,342,1309,1334]" captionStart="FIGURE 3. F" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[378,1236,204,1921]" captionTargetId="figure@5.[341,1246,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. F. maledivensis sp. nov. adult left legs antiaxial view. A) leg I. B) leg II. C) leg III. D) leg IV." httpUri="https://zenodo.org/record/239592/files/figure.png" pageId="3" pageNumber="354">Fig. 3</figureCitation>
|
||
). Monodactylous. Long hook-like claws. Cerotegument finely granular. Cuticle heterogeneous, trochanters dark, proximal third of femur I and II light, remainder dark, femora III and IV dark, all genua dark, all tibiae light, proximal part of all tarsi slightly darker than distal part. Femora without ventral carinae. Large porose areas on paraxial side of femora I and II, paraxial porose areas on femora III and IV divided into dorsal and ventral part. Circular porose areas on dorsal paraxial part of trochanters III and IV. All tarsi with proximal lyrifissure. Dorsal seta
|
||
<emphasis id="B96BEADD2833F472D25BFA4CD21698C9" box="[290,304,1489,1513]" italics="true" pageId="3" pageNumber="354">d</emphasis>
|
||
on all femora slightly thickened, dorsally serrate. Ventral setae of tibiae and tarsi long, ventrally serrate. Chaetome and solenidia see table 1.
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2833F472D3BEF984D0359AC2" blockId="3.[151,1437,940,2018]" pageId="3" pageNumber="354">
|
||
<emphasis id="B96BEADD2833F472D3BEF984D1AA9B12" bold="true" box="[199,652,1561,1586]" pageId="3" pageNumber="354">Common features of juvenile stages.</emphasis>
|
||
Apheredermous. Colour dark brown. Integument plicate and soft, except for centrodorsal plate. Prodorsum triangular, rostrum rounded, cerotegument overall finely granular. Rostral setae (
|
||
<emphasis id="B96BEADD2833F472D398F9FFD3DE9B59" box="[225,248,1634,1657]" italics="true" pageId="3" pageNumber="354">ro</emphasis>
|
||
) thin, long, lamellar setae (
|
||
<emphasis id="B96BEADD2833F472D14EF9FCD16D9B59" box="[567,587,1633,1657]" italics="true" pageId="3" pageNumber="354">le</emphasis>
|
||
) also thin but slightly shorter. Minute exobothridial (
|
||
<emphasis id="B96BEADD2833F472D7C9F9FFD7EF9B59" box="[1200,1225,1634,1657]" italics="true" pageId="3" pageNumber="354">ex</emphasis>
|
||
) and interlamellar setae (
|
||
<emphasis id="B96BEADD2833F472D39AF91BD3DF9BBD" box="[227,249,1670,1693]" italics="true" pageId="3" pageNumber="354">in</emphasis>
|
||
). Bothridia small cup-like, laterally opened. Sensilla short, clavate. On posterior border of prodorsum groups of small pores; in dorsal view sometimes difficult to detect as they are covered by anterior folds of gastronotic region. Gnathosoma no obvious differences from adult stage. Hysterosoma oval in dorsal view, slightly convex in lateral view. Centrodorsal plate with faint reticulate pattern, except for median less sclerotized area resembling inverted Y. Large folds framing centrodorsal plate, showing fine granular surface. Within certain lateral folds series of pores aligned longitudinally, leading into tracheal tubes. Orifice of opisthonotal gland
|
||
<emphasis id="B96BEADD2833F472D67AF8A4D6019A71" box="[1283,1319,1849,1873]" italics="true" pageId="3" pageNumber="354">gla</emphasis>
|
||
located in posterior third of lateral folds. Ventral aspect showing specific pattern of ventral folds, typical for juveniles of this genus. Pores leading into tracheal tubes aligned along these folds. Cerotegument finely granular, slightly larger granules present in furrows and acetabular regions. Legs monodactylous with large hook-like claws. Large porose areas on the same leg segments and position as in adults.
|
||
</paragraph>
|
||
<caption id="DF6066472834F475D3EEF8ECD7119AA7" box="[151,1079,1905,1928]" httpUri="https://zenodo.org/record/239591/files/figure.png" pageId="4" pageNumber="355" targetBox="[196,1381,201,1873]" targetPageId="4">
|
||
<paragraph id="8BA036CF2834F475D3EEF8ECD7119AA7" blockId="4.[151,1079,1905,1928]" box="[151,1079,1905,1928]" pageId="4" pageNumber="355">
|
||
<emphasis id="B96BEADD2834F475D3EEF8ECD2349AA6" bold="true" box="[151,274,1905,1927]" pageId="4" pageNumber="355">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C1F4D4C2834F475D260F8EFD2939AA7" box="[281,437,1905,1927]" class="Arachnida" family="Fortuyniidae" genus="Fortuynia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="355" phylum="Arthropoda" rank="species" species="maledivensis" status="sp. nov.">
|
||
<emphasis id="B96BEADD2834F475D260F8EFD2939AA7" box="[281,437,1905,1927]" italics="true" pageId="4" pageNumber="355">F. maledivensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25857A62834F475D2C2F8EFD12D9AA7" box="[443,523,1906,1927]" pageId="4" pageNumber="355" rank="species">
|
||
<emphasis id="B96BEADD2834F475D2C2F8EFD12D9AA7" bold="true" box="[443,523,1906,1927]" pageId="4" pageNumber="355">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
adult. A) dorsal view. B) ventral view. C) lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6066472835F474D3EEF83CD7EF9A97" box="[151,1225,1953,1975]" httpUri="https://zenodo.org/record/239592/files/figure.png" pageId="5" pageNumber="356" targetBox="[378,1236,204,1921]" targetPageId="5">
|
||
<paragraph id="8BA036CF2835F474D3EEF83CD7EF9A97" blockId="5.[151,1225,1953,1975]" box="[151,1225,1953,1975]" pageId="5" pageNumber="356">
|
||
<emphasis id="B96BEADD2835F474D3EEF83CD2349A96" bold="true" box="[151,274,1953,1975]" pageId="5" pageNumber="356">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C1F4D4C2835F474D260F83FD2939A97" box="[281,437,1953,1975]" class="Arachnida" family="Fortuyniidae" genus="Fortuynia" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="maledivensis" status="sp. nov.">
|
||
<emphasis id="B96BEADD2835F474D260F83FD2939A97" box="[281,437,1953,1975]" italics="true" pageId="5" pageNumber="356">F. maledivensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25857A62835F474D2C2F83CD12D9A96" box="[443,523,1953,1974]" pageId="5" pageNumber="356" rank="species">
|
||
<emphasis id="B96BEADD2835F474D2C2F83CD12D9A96" bold="true" box="[443,523,1953,1974]" pageId="5" pageNumber="356">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
adult left legs antiaxial view. A) leg I. B) leg II. C) leg III. D) leg IV.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6066472836F477D3EEFC58D0319EFB" box="[151,791,965,987]" pageId="6" pageNumber="357">
|
||
<paragraph id="8BA036CF2836F477D3EEFC58D0319EFB" blockId="6.[151,791,965,987]" box="[151,791,965,987]" pageId="6" pageNumber="357">
|
||
<emphasis id="B96BEADD2836F477D3EEFC58D2349EFA" bold="true" box="[151,274,965,987]" pageId="6" pageNumber="357">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C1F4D4C2836F477D260FC5BD2939EFB" box="[281,437,965,987]" class="Arachnida" family="Fortuyniidae" genus="Fortuynia" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="maledivensis" status="sp. nov.">
|
||
<emphasis id="B96BEADD2836F477D260FC5BD2939EFB" box="[281,437,965,987]" italics="true" pageId="6" pageNumber="357">F. maledivensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96BEADD2836F477D2C2FC5BD12D9EFB" bold="true" box="[443,523,966,987]" pageId="6" pageNumber="357">
|
||
<taxonomicNameLabel id="A25857A62836F477D2C2FC5BD12D9EFB" box="[443,523,966,987]" pageId="6" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
protonymph, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA036CF2836F477D3BEFB9FD08C993B" blockId="6.[151,1436,1026,1564]" box="[199,938,1026,1051]" pageId="6" pageNumber="357">
|
||
<emphasis id="B96BEADD2836F477D3BEFB9FD27B993B" bold="true" box="[199,349,1026,1051]" pageId="6" pageNumber="357">Protonymph</emphasis>
|
||
(
|
||
<figureCitation id="13242A4A2836F477D214FB9FD294993B" box="[365,434,1026,1051]" captionStart="FIGURE 4. F" captionStartId="6.[151,250,965,987]" captionText="FIGURE 4. F. maledivensis sp. nov. protonymph, lateral view." pageId="6" pageNumber="357">Fig. 4</figureCitation>
|
||
). (N=6): length 228–277 µm (mean 260 µm)
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFBBBD788991E" blockId="6.[151,1436,1026,1564]" box="[199,1198,1062,1091]" pageId="6" pageNumber="357">
|
||
Gastronotic region. Notogastral setae simple, 15 pairs:
|
||
<emphasis id="B96BEADD2836F477D056FBBAD0769963" box="[815,848,1063,1091]" italics="true" pageId="6" pageNumber="357">c1-3</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D027FBBBD05C991E" box="[862,890,1062,1086]" italics="true" pageId="6" pageNumber="357">da</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D0F1FBBBD08C991E" box="[904,938,1062,1086]" italics="true" pageId="6" pageNumber="357">dm</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D0C0FBBBD0F3991E" box="[953,981,1062,1086]" italics="true" pageId="6" pageNumber="357">dp</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D09AFBBBD0DF991E" box="[995,1017,1062,1086]" italics="true" pageId="6" pageNumber="357">la</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D77FFBBBD704991E" box="[1030,1058,1062,1086]" italics="true" pageId="6" pageNumber="357">lm</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D749FBBBD760991E" box="[1072,1094,1062,1086]" italics="true" pageId="6" pageNumber="357">lp</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D72DFBBBD7519963" box="[1108,1143,1062,1091]" italics="true" pageId="6" pageNumber="357">h1-3</emphasis>
|
||
,
|
||
<emphasis id="B96BEADD2836F477D7FDFBBAD7819963" box="[1156,1191,1063,1091]" italics="true" pageId="6" pageNumber="357">p1-3</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFBD0D21A99AA" blockId="6.[151,1436,1026,1564]" pageId="6" pageNumber="357">Ventral region of idiosoma. Epimeral setation 3-1-2-1. One pair of short genital setae. Adanal and anal setae not developed.</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFB0BD1DE998E" blockId="6.[151,1436,1026,1564]" box="[199,760,1173,1198]" pageId="6" pageNumber="357">
|
||
Legs (
|
||
<figureCitation id="13242A4A2836F477D274FB08D277998E" box="[269,337,1173,1198]" captionStart="FIGURE 5. F" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[333,1255,207,1919]" captionTargetId="figure@7.[306,1281,193,1932]" captionTargetPageId="7" captionText="FIGURE 5. F. maledivensis sp. nov. protonymph left legs, antiaxial view. A) leg I. B) leg II. C) leg III. D) leg IV." httpUri="https://zenodo.org/record/239593/files/figure.png" pageId="6" pageNumber="357">Fig. 5</figureCitation>
|
||
). Chaetome and solenidia see table 1.
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFB24D0C999F2" blockId="6.[151,1436,1026,1564]" box="[199,1007,1209,1234]" pageId="6" pageNumber="357">
|
||
<emphasis id="B96BEADD2836F477D3BEFB24D24599F2" bold="true" box="[199,355,1209,1234]" pageId="6" pageNumber="357">Deutonymph</emphasis>
|
||
(
|
||
<figureCitation id="13242A4A2836F477D208FB24D29E99F2" box="[369,440,1209,1234]" captionStart="FIGURE 6. F" captionStartId="9.[151,250,1953,1975]" captionTargetBox="[266,1321,200,1925]" captionTargetId="figure@9.[253,1333,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. F. maledivensis sp. nov. A) deutonymph, dorsal view. B) deutonymph, ventral view. C) tritonymph, dorsal view. D) tritonymph, ventral view." httpUri="https://zenodo.org/record/239594/files/figure.png" pageId="6" pageNumber="357">Figs 6</figureCitation>
|
||
A–B). (N=4): length 316–335 µm (mean 327 µm).
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFB40D63D99D6" blockId="6.[151,1436,1026,1564]" box="[199,1307,1245,1270]" pageId="6" pageNumber="357">Gastronotic region. Fifteen pairs of notogastral setae, same positions and shapes as in protonymph.</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFA9CD66D981E" blockId="6.[151,1436,1026,1564]" pageId="6" pageNumber="357">
|
||
Ventral region of idiosoma. Epimeral setation 3-1-2-2, seta
|
||
<emphasis id="B96BEADD2836F477D019FA9CD05B9839" box="[864,893,1281,1305]" italics="true" pageId="6" pageNumber="357">4b</emphasis>
|
||
added in this stage. 2 pairs of short genital setae aligned in a longitudinal row. 3 pairs of adanal setae
|
||
<emphasis id="B96BEADD2836F477D19FFAB8D0319862" box="[742,791,1317,1346]" italics="true" pageId="6" pageNumber="357">ad1-3</emphasis>
|
||
flanking anal orifice. Anal setae vestigial, 2 pairs.
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFAD0D1BB9845" blockId="6.[151,1436,1026,1564]" box="[199,669,1356,1381]" pageId="6" pageNumber="357">Legs. Chaetome and solenidia see table 1.</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFAEDD0C298A8" blockId="6.[151,1436,1026,1564]" box="[199,996,1392,1417]" pageId="6" pageNumber="357">
|
||
<emphasis id="B96BEADD2836F477D3BEFAEDD27198A9" bold="true" box="[199,343,1392,1417]" pageId="6" pageNumber="357">Tritonymph</emphasis>
|
||
(
|
||
<figureCitation id="13242A4A2836F477D21FFAEDD28A98A9" box="[358,428,1392,1417]" captionStart="FIGURE 6. F" captionStartId="9.[151,250,1953,1975]" captionTargetBox="[266,1321,200,1925]" captionTargetId="figure@9.[253,1333,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. F. maledivensis sp. nov. A) deutonymph, dorsal view. B) deutonymph, ventral view. C) tritonymph, dorsal view. D) tritonymph, ventral view." httpUri="https://zenodo.org/record/239594/files/figure.png" pageId="6" pageNumber="357">Figs 6</figureCitation>
|
||
C–D). (N=5): length 372–397 µm (mean 381 µm).
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFA09D70F988D" blockId="6.[151,1436,1026,1564]" box="[199,1065,1428,1453]" pageId="6" pageNumber="357">Gastronotic region. Notogastral setae, 15 pairs, no difference to deutonymph.</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEFA25D76898D5" blockId="6.[151,1436,1026,1564]" pageId="6" pageNumber="357">
|
||
Ventral region of idiosoma. Epimeral setation 3-1-3-2, seta
|
||
<emphasis id="B96BEADD2836F477D0FBFA25D0BA98F0" box="[898,924,1464,1488]" italics="true" pageId="6" pageNumber="357">3c</emphasis>
|
||
added in this stage, close to trochanter III. Genital setae 4 pairs, adanal setae 3 pairs (
|
||
<emphasis id="B96BEADD2836F477D10AFA41D18298D9" box="[627,676,1500,1529]" italics="true" pageId="6" pageNumber="357">ad1-3</emphasis>
|
||
). Anal setae fully developed, 2 pairs..
|
||
</paragraph>
|
||
<paragraph id="8BA036CF2836F477D3BEF999D1BB9B3B" blockId="6.[151,1436,1026,1564]" box="[199,669,1539,1564]" pageId="6" pageNumber="357">Legs. Chaetome and solenidia see table 1.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |