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<document id="55803E81C15B31C98F86419A212BAC0C" ID-CLB-Dataset="32226" ID-DOI="10.5852/ejt.2013.54" ID-GBIF-Dataset="769a30e2-392e-4102-a382-f5f606a4c05a" ID-ISSN="2118-9773" ID-Zenodo-Dep="883983" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504510069928" checkinUser="plazi" docAuthor="Tanawat Chaowasku &amp; Paul J. A. Keßler" docDate="2013" docId="03B687D6747EFFCE63A53775FBFEFD9A" docLanguage="en" docName="EJT-54 Chaowasku.pdf" docOrigin="European Journal of Taxonomy 54" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miliusa butonensis Tanawat Chaowasku &amp; Paul J.A. Keßler 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="FF8FFFAE747AFFC96249341FFF9EFFDE" masterDocTitle="Phylogeny of Miliusa (Magnoliales: Annonaceae: Malmeoideae: Miliuseae), with descriptions of two new species from Malesia" masterLastPageNumber="21" masterPageNumber="1" pageNumber="5" updateTime="1698467800801" updateUser="ExternalLinkService">
<mods:mods id="3D7C371A60D2B27F64604AD67766C879" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CC2CC5E4E476C69045D1F4384D1647D3">
<mods:title id="D1F078949EA25EF70E51A31F4259E0B0">Phylogeny of Miliusa (Magnoliales: Annonaceae: Malmeoideae: Miliuseae), with descriptions of two new species from Malesia</mods:title>
</mods:titleInfo>
<mods:name id="23B3DAF4730C0B41518310CC059330F3" type="personal">
<mods:role id="A4E8E79B647024A3655362DFB88AAAC6">
<mods:roleTerm id="0DE488BE3B16DEAA117D4332195AE933">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32CE9E6FB0B13CD87AE43704E1CDC74D">Tanawat Chaowasku</mods:namePart>
</mods:name>
<mods:name id="23999ED5DC5EBDCE7834C885C21C3A29" type="personal">
<mods:role id="59AD2C186FB945D2DDB991505617FB23">
<mods:roleTerm id="4C44EB21B45D73D97666D75F76328520">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="309D7D691F431567507239735003C4CD">Paul J. A. Keßler</mods:namePart>
</mods:name>
<mods:typeOfResource id="5F7FBF5FBDC0DEDBBC10DB14C42C211A">text</mods:typeOfResource>
<mods:relatedItem id="D5B9543FB62FE712DB189B98A1B32E5C" type="host">
<mods:titleInfo id="899C17539487A65FE8B126D5794C73E1">
<mods:title id="DB9041EE215262547058196AED1B28E5">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="3C42F4B10FD700D483C405D093E1B329">
<mods:date id="3F1F59DEE67436659F89E28CC0AEFE99">2013</mods:date>
<mods:detail id="B403C5BD2264E4C956874D42C2817AD0" type="volume">
<mods:number id="0F54116BBE76483D0C711981796EA861">54</mods:number>
</mods:detail>
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<mods:start id="2F0601E7378BAFEBA5CC4A110B62141A">1</mods:start>
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<mods:classification id="DA8681DBB0AB0247D3A376171170A861">journal article</mods:classification>
<mods:identifier id="C1C8644B44D8D6589865A7B52A194BD2" type="CLB-Dataset">32226</mods:identifier>
<mods:identifier id="D99E4A7B3F19A3F316573DCBC59892EF" type="DOI">10.5852/ejt.2013.54</mods:identifier>
<mods:identifier id="576A10648D7E5A220A5A383FCC3130E2" type="GBIF-Dataset">769a30e2-392e-4102-a382-f5f606a4c05a</mods:identifier>
<mods:identifier id="87249E4B4F1222B4CF79E87BC295AE91" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="B6507E4F6630F7692920A95CA9A71947" type="Zenodo-Dep">883983</mods:identifier>
</mods:mods>
<treatment id="03B687D6747EFFCE63A53775FBFEFD9A" ID-DOI="http://doi.org/10.5281/zenodo.5680367" ID-GBIF-Taxon="133169757" ID-Zenodo-Dep="5680367" LSID="urn:lsid:plazi:treatment:03B687D6747EFFCE63A53775FBFEFD9A" httpUri="http://treatment.plazi.org/id/03B687D6747EFFCE63A53775FBFEFD9A" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
<subSubSection id="C305654B747EFFCD63A53775FC72FC76" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BA036C0747EFFCD63A53775FBD6FC5A" blockId="4.[378,1209,731,971]" box="[492,1096,874,901]" pageId="4" pageNumber="5">
<taxonomicName id="4C1F4D43747EFFCD63A53775FC7CFC5A" authority="Chaowasku &amp; Kessler" authorityName="Tanawat Chaowasku &amp; Paul J.A. Keßler" authorityYear="2013" box="[492,994,874,901]" class="Magnoliopsida" family="Annonaceae" genus="Miliusa" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="butonensis" status="sp. nov.">
<emphasis id="B96BEAD2747EFFCD63A53775FD4CFC5B" bold="true" box="[492,722,874,901]" italics="true" pageId="4" pageNumber="5">Miliusa butonensis</emphasis>
Chaowasku &amp; Kessler
</taxonomicName>
,
<taxonomicNameLabel id="A25857A9747EFFCD61A73775FBD6FC5A" box="[1006,1096,874,900]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA036C0747EFFCD600E3792FC72FC76" blockId="4.[378,1209,731,971]" box="[583,1004,909,936]" pageId="4" pageNumber="5">urn:lsid:ipni.org:names:77131155-1</paragraph>
</subSubSection>
<subSubSection id="C305654B747EFFCD60AE37AEFCD3FC15" box="[743,845,945,971]" pageId="4" pageNumber="5" type="description">
<paragraph id="8BA036C0747EFFCD60AE37AEFCD3FC15" blockId="4.[378,1209,731,971]" box="[743,845,945,971]" pageId="4" pageNumber="5">
<figureCitation id="13242A45747EFFCD60AE37AEFCD3FC15" box="[743,845,945,971]" captionStart-0="Fig. 3" captionStart-1="Fig. 4" captionStartId-0="5.[189,232,1965,1991]" captionStartId-1="6.[189,232,1965,1991]" captionTargetBox-0="[196,1394,311,1926]" captionTargetBox-1="[225,1363,1138,1922]" captionTargetId-0="figure@5.[189,1398,304,1931]" captionTargetId-1="figure@6.[225,1362,1139,1922]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 3. Miliusa butonensis sp. nov. a. Habit. b. Outside (abaxial surface) of a persistent inner petal. c. Inside (adaxial surface) of a persistent inner petal. d. Fruit. (All from Coode 6279). Drawn by E. Winkel." captionText-1="Fig. 4. Distribution of Miliusa butonensis sp. nov. (Flled square) and Miliusa viridiFora sp. nov. (Flled circle)." httpUri-0="https://zenodo.org/record/883989/files/figure.png" httpUri-1="https://zenodo.org/record/883991/files/figure.png" pageId="4" pageNumber="5">Figs 3–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C305654B747EFFCD62F437ECFE50FB5D" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BA036C0747EFFCD62F437ECFEA8FBD3" blockId="4.[189,310,1011,1037]" box="[189,310,1011,1037]" pageId="4" pageNumber="5">
<heading id="D0E881AC747EFFCD62F437ECFEA8FBD3" bold="true" box="[189,310,1011,1037]" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<emphasis id="B96BEAD2747EFFCD62F437ECFEA8FBD3" bold="true" box="[189,310,1011,1037]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA036C0747EFFCD62F4303DFE50FB5D" blockId="4.[189,1398,1058,1155]" pageId="4" pageNumber="5">
<taxonomicName id="4C1F4D43747EFFCD62F4303DFE02FBE2" authority="Tanawat Chaowasku &amp; Paul J.A. Keßler, 2013" authorityName="Tanawat Chaowasku &amp; Paul J.A. Keßler" authorityYear="2013" box="[189,412,1058,1084]" class="Magnoliopsida" family="Annonaceae" genus="Miliusa" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="butonensis" status="sp. nov.">
<emphasis id="B96BEAD2747EFFCD62F4303DFE02FBE2" box="[189,412,1058,1084]" italics="true" pageId="4" pageNumber="5">Miliusa butonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857A9747EFFCD63ED303CFD9EFBE3" box="[420,512,1059,1085]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
belongs to clade C. It is best characterized by (1) dense hairs on the apical half inside the inner petals and (2) glandular structures that cover almost the whole basal half of the inner side of the inner petals.
</paragraph>
</subSubSection>
<caption id="DF606648747EFFCD62F43394FF73F832" httpUri="https://zenodo.org/record/883987/files/figure.png" pageId="4" pageNumber="5" targetBox="[320,1266,1217,1875]" targetPageId="4">
<paragraph id="8BA036C0747EFFCD62F43394FF73F832" blockId="4.[189,1398,1931,2028]" pageId="4" pageNumber="5">
<emphasis id="B96BEAD2747EFFCD62F43394FE95F87B" bold="true" box="[189,267,1931,1957]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
50% Bayesian majority-rule consensus phylogram of combined seven plastid markers, showing branch length proportional to amount of lineage sequence divergence. Scale bar unit: substitution per site.
</paragraph>
</caption>
<caption id="DF606648747FFFCC62F433B2FAE9F834" httpUri="https://zenodo.org/record/883989/files/figure.png" pageId="5" pageNumber="6" targetBox="[196,1394,311,1926]" targetPageId="5">
<paragraph id="8BA036C0747FFFCC62F433B2FAE9F834" blockId="5.[189,1399,1965,2027]" pageId="5" pageNumber="6">
<emphasis id="B96BEAD2747FFFCC62F433B2FE92F819" bold="true" box="[189,268,1965,1991]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="4C1F4D43747FFFCC635D33B2FE6AF819" authority="Tanawat Chaowasku &amp; Paul J.A. Keßler, 2013" authorityName="Tanawat Chaowasku &amp; Paul J.A. Keßler" authorityYear="2013" box="[276,500,1965,1991]" class="Magnoliopsida" family="Annonaceae" genus="Miliusa" kingdom="Plantae" order="Magnoliales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="butonensis" status="sp. nov.">
<emphasis id="B96BEAD2747FFFCC635D33B2FE6AF819" box="[276,500,1965,1991]" italics="true" pageId="5" pageNumber="6">Miliusa butonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857A9747FFFCC63B433B1FDC7F816" box="[509,601,1966,1992]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96BEAD2747FFFCC602833B2FDE9F819" bold="true" box="[609,631,1965,1991]" pageId="5" pageNumber="6">a.</emphasis>
Habit.
<emphasis id="B96BEAD2747FFFCC609833B2FD77F819" bold="true" box="[721,745,1965,1991]" pageId="5" pageNumber="6">b.</emphasis>
Outside (abaxial surface) of a persistent inner petal.
<emphasis id="B96BEAD2747FFFCC672B33B2FAE8F819" bold="true" box="[1378,1398,1965,1991]" pageId="5" pageNumber="6">c.</emphasis>
Inside (adaxial surface) of a persistent inner petal.
<emphasis id="B96BEAD2747FFFCC614C33CFFC82F834" bold="true" box="[773,796,2000,2026]" pageId="5" pageNumber="6">d.</emphasis>
Fruit. (All from
<emphasis id="B96BEAD2747FFFCC619433CFFBF5F834" box="[989,1131,2000,2026]" italics="true" pageId="5" pageNumber="6">Coode 6279</emphasis>
). Drawn by E. Winkel.
</paragraph>
</caption>
<subSubSection id="C305654B747CFFCF62F43514FB91FE89" box="[189,1039,267,344]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BA036C0747CFFCF62F43514FEDAFEFB" blockId="6.[189,324,267,293]" box="[189,324,267,293]" pageId="6" pageNumber="7">
<heading id="D0E881AC747CFFCF62F43514FEDAFEFB" bold="true" box="[189,324,267,293]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<emphasis id="B96BEAD2747CFFCF62F43514FEDAFEFB" bold="true" box="[189,324,267,293]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA036C0747CFFCF62F43522FB91FE89" blockId="6.[189,1039,317,344]" box="[189,1039,317,344]" pageId="6" pageNumber="7">
Named after the Buton Island (
<collectingCountry id="F3087650747CFFCF606E3522FD03FE86" box="[551,669,317,344]" name="Indonesia" pageId="6" pageNumber="7">Indonesia</collectingCountry>
) where this species is endemic.
</paragraph>
</subSubSection>
<subSubSection id="C305654B747CFFCF62F4359CFB24FE29" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BA036C0747CFFCF62F4359CFF64FE43" blockId="6.[189,250,387,413]" box="[189,250,387,413]" pageId="6" pageNumber="7">
<heading id="D0E881AC747CFFCF62F4359CFF64FE43" bold="true" box="[189,250,387,413]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<emphasis id="B96BEAD2747CFFCF62F4359CFF64FE43" bold="true" box="[189,250,387,413]" pageId="6" pageNumber="7">Type</emphasis>
</heading>
</paragraph>
<paragraph id="8BA036C0747CFFCF62F435AAFB24FE29" blockId="6.[189,1399,437,503]" pageId="6" pageNumber="7">
<materialsCitation id="3B773C9D747CFFCF62F435AAFB24FE29" ID-GBIF-Occurrence="1598309672" collectingDate="1989-11" country="Indonesia" county="North Buton Is." location="Labuan Tobelo" municipality="Jismil" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Sulawesi">
<collectingCountry id="F3087650747CFFCF62F435AAFEFFFE11" box="[189,353,437,463]" name="Indonesia" pageId="6" pageNumber="7">INDONESIA</collectingCountry>
: Southeast of
<collectingRegion id="49DBF822747CFFCF604235AAFDE6FE11" box="[523,632,437,463]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="7">Sulawesi</collectingRegion>
,
<collectingCounty id="62C14E4C747CFFCF60CC35AAFCDFFE0E" box="[645,833,437,464]" pageId="6" pageNumber="7">North Buton Is.</collectingCounty>
,
<collectingMunicipality id="6BC4ACBA747CFFCF611935AAFC08FE11" box="[848,918,437,463]" pageId="6" pageNumber="7">Jismil</collectingMunicipality>
camp inland from
<location id="8EC0601B747CFFCF663535AAFAAFFE11" LSID="urn:lsid:plazi:treatment:03B687D6747EFFCE63A53775FBFEFD9A:8EC0601B747CFFCF663535AAFAAFFE11" box="[1148,1329,437,463]" country="Indonesia" county="North Buton Is." municipality="Jismil" name="Labuan Tobelo" pageId="6" pageNumber="7" stateProvince="Sulawesi">Labuan Tobelo</location>
,
<date id="FFA11000747CFFCF677735A9FF64FE29" pageId="6" pageNumber="7" value="1989-11">
<collectingDate id="EFE5E9E8747CFFCF677735A9FF64FE29" pageId="6" pageNumber="7" value="1989-11">Nov. 1989</collectingDate>
</date>
,
<emphasis id="B96BEAD2747CFFCF634F35C3FE0BFE28" box="[262,405,476,502]" italics="true" pageId="6" pageNumber="7">Coode 6279</emphasis>
[holo-: U; iso-: A, K, L], in fruit (with a few persistent inner petals).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C305654B747CFFCF62F4363DFC58FB9B" pageId="6" pageNumber="7" type="description">
<paragraph id="8BA036C0747CFFCF62F4363DFED0FDE2" blockId="6.[189,334,546,572]" box="[189,334,546,572]" pageId="6" pageNumber="7">
<heading id="D0E881AC747CFFCF62F4363DFED0FDE2" bold="true" box="[189,334,546,572]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<emphasis id="B96BEAD2747CFFCF62F4363DFED0FDE2" bold="true" box="[189,334,546,572]" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA036C0747CFFCF62F4364AFC58FB9B" blockId="6.[189,1399,596,1093]" pageId="6" pageNumber="7">
Trees, ca.
<quantity id="4CE79B25747CFFCF637A364BFEC1FDB1" box="[307,351,596,623]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="m" value="8.0">8 m</quantity>
tall, ca.
<quantity id="4CE79B25747CFFCF6388364BFD96FDB1" box="[449,520,596,623]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="cm" value="10.0">10 cm</quantity>
in diameter. Young twigs appressed-puberulous. Petioles 2.0–3.0 mm long, appressed-puberulous. Leaves elliptic to slightly ovate, 4.7–11.1 ×
<quantity id="4CE79B25747CFFCF619B3664FBC7FD48" box="[978,1113,635,662]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="4.5" metricValueMin="2.2" pageId="6" pageNumber="7" unit="cm" value="3.35" valueMax="4.5" valueMin="2.2">2.2–4.5 cm</quantity>
, base (broadly) cuneate, sometimes slightly unequal, apex acute; lamina glabrous both sides; midrib usually slightly raised to Fat above, sometimes slightly sunken, glabrous, raised below, appressed-puberulous; secondary veins 15–18 pairs, angle with midrib 42°–50°. Flowers partially known; fruiting peduncles or fruiting pedicels axillary; fruiting peduncles up to
<quantity id="4CE79B25747CFFCF60F03707FC92FCED" box="[697,780,792,819]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="cm" value="0.4">0.4 cm</quantity>
long; fruiting pedicels
<quantity id="4CE79B25747CFFCF666F3707FB32FCED" box="[1062,1196,792,819]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.1" metricValueMin="0.9" pageId="6" pageNumber="7" unit="cm" value="1.5" valueMax="2.1" valueMin="0.9">0.9–2.1 cm</quantity>
long; sepals and sometimes outer petals persistent in fruit, both ovate-triangular. Inner petals a few persistent ones fallen on herbarium sheet near the torus bearing monocarps seen, ovate-triangular, ca. 6.0 × 4.0 mm; outside appressed-puberulous, mostly on the middle part, margin puberulous, inside (curly-)velvety on the apical half; glandular structures observed on almost the whole basal half, slightly raised, between crescentshaped and semicircular, slightly Fnely warty; base not saccate. Monocarps 3–21, subglobose-ellipsoid, 0.7–0.9 ×
<quantity id="4CE79B25747CFFCF6371301CFE5EFBC0" box="[312,448,1027,1054]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="7" unit="cm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 cm</quantity>
; surface smooth, glabrous; apex not apiculate; stipe ca. 6.0 mm long, glabrous. Seed(s) 1 (rarely 2), subglobose(-ellipsoid), 0.6–0.8 ×
<quantity id="4CE79B25747CFFCF61723035FC5FFB9B" box="[827,961,1066,1093]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="6" pageNumber="7" unit="cm" value="0.55" valueMax="0.7" valueMin="0.4">0.4–0.7 cm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF606648747CFFCF62F433B2FE8EF832" httpUri="https://zenodo.org/record/883991/files/figure.png" pageId="6" pageNumber="7" targetBox="[225,1363,1138,1922]" targetPageId="6">
<paragraph id="8BA036C0747CFFCF62F433B2FE8EF832" blockId="6.[189,1398,1965,2028]" pageId="6" pageNumber="7">
<emphasis id="B96BEAD2747CFFCF62F433B2FE95F819" bold="true" box="[189,267,1965,1991]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
Distribution of
<taxonomicName id="4C1F4D43747CFFCF638133B2FD38F819" authority="Tanawat Chaowasku &amp; Paul J.A. Keßler, 2013" authorityName="Tanawat Chaowasku &amp; Paul J.A. Keßler" authorityYear="2013" box="[456,678,1965,1991]" class="Magnoliopsida" family="Annonaceae" genus="Miliusa" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="butonensis" status="sp. nov.">
<emphasis id="B96BEAD2747CFFCF638133B2FD38F819" box="[456,678,1965,1991]" italics="true" pageId="6" pageNumber="7">Miliusa butonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857A9747CFFCF60E733B1FC97F816" box="[686,777,1966,1992]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(Flled square) and
<emphasis id="B96BEAD2747CFFCF61A733B2FB59F819" box="[1006,1223,1965,1991]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C1F4D43747CFFCF61A733B2FBD6F819" box="[1006,1096,1965,1991]" class="Magnoliopsida" family="Annonaceae" genus="Miliusa" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="genus">Miliusa</taxonomicName>
viridiFora
</emphasis>
sp. nov. (Flled circle).
</paragraph>
</caption>
<subSubSection id="C305654B747DFFCE62F43514FB59FEA0" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BA036C0747DFFCE62F43514FDE3FEFB" blockId="7.[189,637,267,293]" box="[189,637,267,293]" pageId="7" pageNumber="8">
<heading id="D0E881AC747DFFCE62F43514FDE3FEFB" bold="true" box="[189,637,267,293]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<emphasis id="B96BEAD2747DFFCE62F43514FDE3FEFB" bold="true" box="[189,637,267,293]" pageId="7" pageNumber="8">Distribution, habitat and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA036C0747DFFCE62F43522FB59FEA0" blockId="7.[189,1398,317,383]" pageId="7" pageNumber="8">
<collectingCountry id="F3087650747DFFCE62F43522FEB1FE89" box="[189,303,317,343]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
(Buton Island,
<figureCitation id="13242A45747DFFCE63AD3522FDB4FE89" box="[484,554,317,343]" captionStart="Fig. 4" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[225,1363,1138,1922]" captionTargetId="figure@6.[225,1362,1139,1922]" captionTargetPageId="6" captionText="Fig. 4. Distribution of Miliusa butonensis sp. nov. (Flled square) and Miliusa viridiFora sp. nov. (Flled circle)." httpUri="https://zenodo.org/record/883991/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
), occurring in forests on Fat ridge-tops, with slightly broken canopy due to rocky terrain of raised coralline limestone. Elevation: ca.
<specimenCount id="9D19FD49747DFFCE6137357BFC51FEA1" box="[894,975,356,383]" pageId="7" pageNumber="8" type="generic">
<quantity id="4CE79B25747DFFCE6137357BFC55FEA1" box="[894,971,356,383]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="m" value="300.0">300 m</quantity>
.
</specimenCount>
Fruiting: November.
</paragraph>
</subSubSection>
<subSubSection id="C305654B747DFFCE62F435B6FBFEFD9A" pageId="7" pageNumber="8" type="description">
<paragraph id="8BA036C0747DFFCE62F435B6FEDBFE1A" blockId="7.[189,325,425,452]" box="[189,325,425,452]" pageId="7" pageNumber="8">
<heading id="D0E881AC747DFFCE62F435B6FEDBFE1A" bold="true" box="[189,325,425,452]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<emphasis id="B96BEAD2747DFFCE62F435B6FEDBFE1A" bold="true" box="[189,325,425,452]" pageId="7" pageNumber="8">Field notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA036C0747DFFCE62F435C3FBFEFD9A" blockId="7.[189,1399,476,581]" pageId="7" pageNumber="8">
Branches horizontal and foliage in Fat sprays. Bark grey-brown, ca.
<quantity id="4CE79B25747DFFCE664135C3FBCEFE29" box="[1032,1104,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="mm" value="4.0">4 mm</quantity>
thick overall, ± smooth with Fne vertical cracks and rows of low lenticels, wood straw-colored, cut bark and wood pleasantly aromatic. Young leaves ± yellow-green, mature leaves mid green on both sides.
</paragraph>
</subSubSection>
</treatment>
</document>
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