335 lines
35 KiB
XML
335 lines
35 KiB
XML
<document id="0EF910CE2AAD70E347A6E9FAD8BDD79A" ID-DOI="10.11646/zootaxa.3985.2.5" ID-GBIF-Dataset="450ef146-033c-4fa5-b495-063daf4abb51" ID-ISSN="1175-5326" ID-Zenodo-Dep="239619" ID-ZooBank="211AB430-929E-4468-A68A-FEE77415908A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461257578087" checkinUser="plazi" docAuthor="Bolet, Arnau, Daza, Juan D., Augé, Marc & Bauer, Aaron M." docDate="2015" docId="03B68793FFE2FFAFC58A944CFA97FC70" docLanguage="en" docName="zt03985p274.pdf" docOrigin="Zootaxa 3985 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cadurcogekko verus Bolet, Daza, Augé & Bauer, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="268" masterDocId="FF8FFFEBFFE3FFACC51D9231FFDFFFF0" masterDocTitle="New genus and species names for the Eocene lizard Cadurcogekko rugosus Augé, 2005" masterLastPageNumber="274" masterPageNumber="265" pageNumber="266" updateTime="1698608421459" updateUser="plazi">
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<mods:mods id="E591C2D20DC4A1852A2AC85E8225AD4A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="65361DCC23D711E06B958B32D7637F3D">
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<mods:title id="4BEC6F04FBEB1F9555C8A031D19CD6F1">New genus and species names for the Eocene lizard Cadurcogekko rugosus Augé, 2005</mods:title>
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</mods:titleInfo>
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<mods:name id="787ADB5801A57D81BB39510BD5B272D8" type="personal">
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<mods:roleTerm id="CE3D1C0A5D9AAC5F84CA60405761D90F">Author</mods:roleTerm>
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<mods:namePart id="A54B1FFFA0A2A5040E9D6F6D58A0430F">Bolet, Arnau</mods:namePart>
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<mods:roleTerm id="8B260680FF36C5EBB8943DB8197C7B51">Author</mods:roleTerm>
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<mods:namePart id="DFB94E6FA4783D41AFB27B279A365631">Daza, Juan D.</mods:namePart>
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<mods:name id="FCA35A738120BD4798697E63BD11F124" type="personal">
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<mods:roleTerm id="67C3C62D3EA1819A2CA43B80B21D105A">Author</mods:roleTerm>
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<mods:namePart id="DEE0D9EA383B89DA9183CBD6D8FD60E9">Augé, Marc</mods:namePart>
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</mods:name>
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<mods:name id="42CD2409C32E0A1634690F89A95867A9" type="personal">
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<mods:roleTerm id="47D0DA268DA809046BA5BE00788F1777">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AAB0FD12B5F65E84D19209E9E7C27A28">Bauer, Aaron M.</mods:namePart>
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<mods:typeOfResource id="96387322821208C07261FF135C2E971F">text</mods:typeOfResource>
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<mods:title id="470DDCF1B9A9644EBAB26148D3D7E393">Zootaxa</mods:title>
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<mods:part id="66C98569B86C1CF99379D0D324CD4631">
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<mods:date id="7E9838DF86F8B23E5AFF1ADC201CD95E">2015</mods:date>
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<mods:detail id="500DF1A4E6BE6CAD215699C596064B4C" type="volume">
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<mods:number id="A87371611F063DD51A514DD5C472FED1">3985</mods:number>
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<mods:detail id="11B762583EC5F83460CEEF1B276F6E8B" type="issue">
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<mods:number id="C6CC31BBB28F871D54BC44962141A6CA">2</mods:number>
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<mods:classification id="52FA9E09CFD8E4AA6E4D988DF1C53485">journal article</mods:classification>
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<mods:identifier id="1C3F03ABC6A9E0DA1F31A6ED5C7777E3" type="DOI">10.11646/zootaxa.3985.2.5</mods:identifier>
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<mods:identifier id="C65596A3F14D36687A808FFB808F2866" type="GBIF-Dataset">450ef146-033c-4fa5-b495-063daf4abb51</mods:identifier>
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<mods:identifier id="F878C9E2F7F207F062AD1AB78B3A3934" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5CDC7860359BB548931695615D8A2819" type="Zenodo-Dep">239619</mods:identifier>
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<treatment id="03B68793FFE2FFAFC58A944CFA97FC70" ID-DOI="http://doi.org/10.5281/zenodo.5680474" ID-GBIF-Taxon="119636357" ID-Zenodo-Dep="5680474" LSID="urn:lsid:plazi:treatment:03B68793FFE2FFAFC58A944CFA97FC70" httpUri="http://treatment.plazi.org/id/03B68793FFE2FFAFC58A944CFA97FC70" lastPageId="3" lastPageNumber="268" pageId="1" pageNumber="266">
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<subSubSection id="C305650EFFE2FFADC58A944CFF34F948" pageId="1" pageNumber="266" type="nomenclature">
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<paragraph id="8BA03685FFE2FFADC58A944CFE2BF967" blockId="1.[151,500,1661,1720]" box="[151,500,1661,1687]" pageId="1" pageNumber="266">
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<heading id="D0E881E9FFE2FFADC58A944CFE2BF967" bold="true" box="[151,500,1661,1687]" fontSize="11" level="1" pageId="1" pageNumber="266" reason="1">
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<emphasis id="B96BEA97FFE2FFADC58A944CFE2BF967" bold="true" box="[151,500,1661,1687]" pageId="1" pageNumber="266">
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<taxonomicName id="4C1F4D06FFE2FFADC58A944CFE50F967" box="[151,399,1661,1687]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="verus" status="sp. nov.">
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<emphasis id="B96BEA97FFE2FFADC58A944CFE50F967" bold="true" box="[151,399,1661,1687]" italics="true" pageId="1" pageNumber="266">Cadurcogekko verus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25857ECFFE2FFADC48B944CFE2BF967" box="[406,500,1661,1687]" pageId="1" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA03685FFE2FFADC58A94AEFF34F948" blockId="1.[151,500,1661,1720]" box="[151,235,1695,1720]" pageId="1" pageNumber="266">
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(
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<figureCitation id="13242A00FFE2FFADC58294AEFF3FF948" box="[159,224,1695,1720]" captionStart="FIGURE 1" captionStartId="2.[151,250,901,923]" captionTargetBox="[169,1414,206,866]" captionTargetId="figure@2.[151,1436,195,877]" captionTargetPageId="2" captionText="FIGURE 1. Holotype (right maxilla) of Cadurcogekko verus (USTL, PRA 9) in labial (A) and lingual (B) views, from Les Pradigues (MP 17). Scale bar in mm." httpUri="https://zenodo.org/record/239620/files/figure.png" pageId="1" pageNumber="266">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C305650EFFE2FFADC58A94D6FA54F860" pageId="1" pageNumber="266" type="materials_examined">
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<paragraph id="8BA03685FFE2FFADC58A94D6FCFDF8F0" blockId="1.[151,1436,1767,2009]" box="[151,802,1767,1792]" pageId="1" pageNumber="266">
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<emphasis id="B96BEA97FFE2FFADC58A94D6FED4F8F0" bold="true" box="[151,267,1767,1792]" pageId="1" pageNumber="266">
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<typeStatus id="54A48827FFE2FFADC58A94D6FED9F8F0" box="[151,262,1767,1792]" pageId="1" pageNumber="266" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED0EAE40FFE2FFADC40F94D9FE84F8F0" box="[274,347,1768,1792]" pageId="1" pageNumber="266">USTL</collectionCode>
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, PRA 9 (almost complete right maxilla).
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</paragraph>
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<paragraph id="8BA03685FFE2FFADC5DA953DFC3CF8D4" blockId="1.[151,1436,1767,2009]" box="[199,995,1804,1829]" pageId="1" pageNumber="266">
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<emphasis id="B96BEA97FFE2FFADC5DA953DFEBAF8D5" bold="true" box="[199,357,1804,1829]" pageId="1" pageNumber="266">
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<typeStatus id="54A48827FFE2FFADC5DA953DFEDDF8D5" box="[199,258,1804,1829]" pageId="1" pageNumber="266">Type</typeStatus>
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locality.
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</emphasis>
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Les Pradigues (MP17), Phosphorites du Quercy,
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<collectingCountry id="F3087615FFE2FFADC692953CFC00F8D4" box="[911,991,1805,1828]" name="France" pageId="1" pageNumber="266">France</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BA03685FFE2FFADC5DA951EFBE8F89D" blockId="1.[151,1436,1767,2009]" pageId="1" pageNumber="266">
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<emphasis id="B96BEA97FFE2FFADC5DA951EFE7BF8B8" bold="true" box="[199,420,1839,1864]" pageId="1" pageNumber="266">Referred material.</emphasis>
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From Sindou D (MP18-19):
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<collectionCode id="ED0EAE40FFE2FFADC7F59501FCEEF8B8" box="[744,817,1840,1864]" pageId="1" pageNumber="266">USTL</collectionCode>
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,
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<collectionCode id="ED0EAE40FFE2FFADC6219501FCABF8B8" box="[828,884,1840,1864]" pageId="1" pageNumber="266">SND</collectionCode>
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2016 (frontal);
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<collectionCode id="ED0EAE40FFE2FFADC1399501FBB2F8B8" box="[1060,1133,1840,1864]" pageId="1" pageNumber="266">USTL</collectionCode>
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,
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<collectionCode id="ED0EAE40FFE2FFADC16A9501FB70F8B8" box="[1143,1199,1840,1864]" pageId="1" pageNumber="266">SND</collectionCode>
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626, 627 (two dorsal vertebrae, note that these specimens were reported as uncatalogued in
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<bibRefCitation id="EF8E4B74FFE2FFADC6B79565FBF4F89D" author="Auge" box="[938,1067,1876,1901]" pageId="1" pageNumber="266" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé 2005</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BA03685FFE2FFADC5DA9546FA54F860" blockId="1.[151,1436,1767,2009]" box="[199,1419,1911,1936]" pageId="1" pageNumber="266">
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<emphasis id="B96BEA97FFE2FFADC5DA9546FE1CF860" bold="true" box="[199,451,1911,1936]" pageId="1" pageNumber="266">Chronological range.</emphasis>
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From MP17 to MP19 (late Eocene). Les Pradigues (MP17) and Sindou D (MP18-19).
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</paragraph>
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</subSubSection>
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<subSubSection id="C305650EFFE2FFADC5DA95ADFB77F829" pageId="1" pageNumber="266" type="etymology">
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<paragraph id="8BA03685FFE2FFADC5DA95ADFB77F829" blockId="1.[151,1436,1767,2009]" pageId="1" pageNumber="266">
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<emphasis id="B96BEA97FFE2FFADC5DA95ADFE91F845" bold="true" box="[199,334,1948,1973]" pageId="1" pageNumber="266">Etymology.</emphasis>
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Latin ‘
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<taxonomicName id="4C1F4D06FFE2FFADC4B895ACFE3DF844" box="[421,482,1949,1972]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="verus">
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<emphasis id="B96BEA97FFE2FFADC4B895ACFE3DF844" box="[421,482,1949,1972]" italics="true" pageId="1" pageNumber="266">verus</emphasis>
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</taxonomicName>
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’, meaning true, referring to the fact that this species is being described after been separated from the
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<taxonomicName id="4C1F4D06FFE2FFADC47395F0FE37F828" box="[366,488,1985,2008]" class="Reptilia" family="Scincidae" genus="Gekkomimus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="rugosus">
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<emphasis id="B96BEA97FFE2FFADC47395F0FE37F828" box="[366,488,1985,2008]" italics="true" pageId="1" pageNumber="266">C. rugosus</emphasis>
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</taxonomicName>
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<typeStatus id="54A48827FFE2FFADC4F395F1FD8CF829" box="[494,595,1984,2009]" pageId="1" pageNumber="266" type="holotype">holotype</typeStatus>
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, which actually corresponds to a scincid (see below).
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</paragraph>
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</subSubSection>
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<caption id="DF60660DFFE1FFAEC58A91B4FDD6FC4A" httpUri="https://zenodo.org/record/239620/files/figure.png" pageId="2" pageNumber="267" targetBox="[169,1414,206,866]" targetPageId="2">
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<paragraph id="8BA03685FFE1FFAEC58A91B4FDD6FC4A" blockId="2.[151,1435,901,954]" pageId="2" pageNumber="267">
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<emphasis id="B96BEA97FFE1FFAEC58A91B4FECAFC6B" bold="true" box="[151,277,901,923]" pageId="2" pageNumber="267">FIGURE 1.</emphasis>
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Holotype (right maxilla) of
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<taxonomicName id="4C1F4D06FFE1FFAEC72391B4FCCFFC6C" box="[574,784,901,924]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="verus">
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<emphasis id="B96BEA97FFE1FFAEC72391B4FCCFFC6C" box="[574,784,901,924]" italics="true" pageId="2" pageNumber="267">Cadurcogekko verus</emphasis>
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</taxonomicName>
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(USTL, PRA 9) in labial (A) and lingual (B) views, from Les Pradigues (MP17). Scale bar in mm.
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</paragraph>
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</caption>
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<caption id="DF60660DFFE1FFAEC58A971CFDF8FA92" httpUri="https://zenodo.org/record/239621/files/figure.png" pageId="2" pageNumber="267" targetBox="[151,1436,1004,1303]" targetPageId="2">
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<paragraph id="8BA03685FFE1FFAEC58A971CFDF8FA92" blockId="2.[151,1436,1325,1378]" pageId="2" pageNumber="267">
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<emphasis id="B96BEA97FFE1FFAEC58A971CFEC9FAB2" bold="true" box="[151,278,1325,1347]" pageId="2" pageNumber="267">FIGURE 2.</emphasis>
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Partial left dentary of
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<emphasis id="B96BEA97FFE1FFAEC716971CFCCBFAB3" box="[523,788,1325,1347]" italics="true" pageId="2" pageNumber="267">Cadurciguana hoffstetteri</emphasis>
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(USTL, SND 625) in labial (A) and lingual (B) views, from Sindou D (MP18-19). Scale bar in mm.
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</paragraph>
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</caption>
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<subSubSection id="C305650EFFE1FFAEC5DA97B8FEA8FA1A" pageId="2" pageNumber="267" type="diagnosis">
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<paragraph id="8BA03685FFE1FFAEC5DA97B8FEA8FA1A" blockId="2.[151,1436,1417,2018]" pageId="2" pageNumber="267">
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||
<emphasis id="B96BEA97FFE1FFAEC5DA97B8FE9EFA52" bold="true" box="[199,321,1417,1442]" pageId="2" pageNumber="267">Diagnosis.</emphasis>
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A species of
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<taxonomicName id="4C1F4D06FFE1FFAEC4C297B8FD5EFA52" box="[479,641,1417,1442]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="genus">
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<emphasis id="B96BEA97FFE1FFAEC4C297B8FD5EFA52" box="[479,641,1417,1442]" italics="true" pageId="2" pageNumber="267">Cadurcogekko</emphasis>
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</taxonomicName>
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differing from
|
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<taxonomicName id="4C1F4D06FFE1FFAEC62F97BBFC65FA51" box="[818,954,1418,1441]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="piveteaui">
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<emphasis id="B96BEA97FFE1FFAEC62F97BBFC9AFA51" box="[818,837,1418,1441]" italics="true" pageId="2" pageNumber="267">C</emphasis>
|
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.
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<emphasis id="B96BEA97FFE1FFAEC64997BBFC65FA51" box="[852,954,1418,1441]" italics="true" pageId="2" pageNumber="267">piveteaui</emphasis>
|
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</taxonomicName>
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in its smaller size, a lower tooth count on the maxilla (ca. 30 vs.
|
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<quantity id="4CE79B60FFE1FFAEC4BC979CFE02FA36" box="[417,477,1453,1478]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="267" unit="in" value="50.0">50 in</quantity>
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<taxonomicName id="4C1F4D06FFE1FFAEC4FB979FFDAFFA35" box="[486,624,1454,1477]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="piveteaui">
|
||
<emphasis id="B96BEA97FFE1FFAEC4FB979FFE26FA35" box="[486,505,1454,1477]" italics="true" pageId="2" pageNumber="267">C</emphasis>
|
||
.
|
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<emphasis id="B96BEA97FFE1FFAEC714979FFDAFFA35" box="[521,624,1454,1477]" italics="true" pageId="2" pageNumber="267">piveteaui</emphasis>
|
||
</taxonomicName>
|
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), and a coarser ornamentation on the labial surface of the maxilla and possibly the frontal.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305650EFFE1FFAFC5DA97C4FA97FC70" lastPageId="3" lastPageNumber="268" pageId="2" pageNumber="267" type="discussion">
|
||
<paragraph id="8BA03685FFE1FFAEC5DA97C4FD84F8DE" blockId="2.[151,1436,1417,2018]" pageId="2" pageNumber="267">
|
||
<emphasis id="B96BEA97FFE1FFAEC5DA97C4FEE4F9FE" bold="true" box="[199,315,1525,1550]" pageId="2" pageNumber="267">Remarks.</emphasis>
|
||
The
|
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<typeStatus id="54A48827FFE1FFAEC46E97C4FE0AF9FE" box="[371,469,1525,1550]" pageId="2" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4D06FFE1FFAEC4E797C4FC5BF9FE" authority="Auge, 2005" authorityName="Auge" authorityYear="2005" box="[506,900,1525,1550]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE1FFAEC4E797C4FD24F9FD" box="[506,763,1525,1550]" italics="true" pageId="2" pageNumber="267">Cadurcogekko rugosus</emphasis>
|
||
<bibRefCitation id="EF8E4B74FFE1FFAEC61F97C7FC5BF9FE" author="Auge" box="[770,900,1525,1550]" pageId="2" pageNumber="267" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
is clearly a scincid dentary. Because the species name is linked to the
|
||
<typeStatus id="54A48827FFE1FFAEC4829428FDDBF9C2" box="[415,516,1561,1586]" pageId="2" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
, which exhibits significant differences with respect to both extant and extinct members of
|
||
<taxonomicName id="4C1F4D06FFE1FFAEC43A940CFE48F9A6" box="[295,407,1597,1622]" class="Reptilia" family="Scincidae" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="family">Scincidae</taxonomicName>
|
||
, we assign this species to a new genus (see below). The fragment of left dentary (USTL, SND 625;
|
||
<figureCitation id="13242A00FFE1FFAEC4099450FE86F98A" box="[276,345,1633,1658]" captionStart="FIGURE 2" captionStartId="2.[151,250,1325,1347]" captionTargetBox="[151,1436,1004,1303]" captionTargetId="figure@2.[151,1436,1004,1304]" captionTargetPageId="2" captionText="FIGURE 2. Partial left dentary of Cadurciguana hoffstetteri (USTL, SND 625) in labial (A) and lingual (B) views, from Sindou D (MP 18 - 19). Scale bar in mm." httpUri="https://zenodo.org/record/239621/files/figure.png" pageId="2" pageNumber="267">Fig. 2</figureCitation>
|
||
) tentatively referred to
|
||
<taxonomicName id="4C1F4D06FFE1FFAEC7709453FD36F989" box="[621,745,1634,1657]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE1FFAEC7709453FD5FF989" box="[621,640,1634,1657]" italics="true" pageId="2" pageNumber="267">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE1FFAEC78D9453FD36F989" box="[656,745,1634,1657]" italics="true" pageId="2" pageNumber="267">rugosus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8E4B74FFE1FFAEC6059453FC76F98A" author="Auge" box="[792,937,1633,1658]" pageId="2" pageNumber="267" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé (2005)</bibRefCitation>
|
||
presents a closed Meckelian canal, but the clearly tricuspid teeth, the way the Meckelian canal is fused, and the overall morphology of the dentary all allow a referral to the iguanid
|
||
<emphasis id="B96BEA97FFE1FFAEC48E9498FD6BF932" box="[403,692,1705,1730]" italics="true" pageId="2" pageNumber="267">Cadurciguana hoffstetteri</emphasis>
|
||
<bibRefCitation id="EF8E4B74FFE1FFAEC7A1949BFC9EF932" author="Auge" box="[700,833,1706,1730]" pageId="2" pageNumber="267" refString="Auge, M. (1987) Confirmation de la presence d'Iguanidae (Reptilia, Lacertilia) dans l'Eocene europeen. Comptes Rendus de l'Academie des Sciences Paris, 305, 633 - 636." type="journal article" year="1987">Augé, 1987</bibRefCitation>
|
||
. This taxon was previously known from Sindou D on the basis of vertebrae (
|
||
<bibRefCitation id="EF8E4B74FFE1FFAEC48894FFFDC8F916" author="Auge" box="[405,535,1741,1766]" pageId="2" pageNumber="267" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé 2005</bibRefCitation>
|
||
), and misidentification of
|
||
<emphasis id="B96BEA97FFE1FFAEC62394FCFC00F916" box="[830,991,1741,1766]" italics="true" pageId="2" pageNumber="267">Cadurciguana</emphasis>
|
||
as a gekkotan has been reported before (
|
||
<bibRefCitation id="EF8E4B74FFE1FFAEC58394C3FEF8F8FA" author="Auge" box="[158,295,1778,1802]" pageId="2" pageNumber="267" refString="Auge, M. (1987) Confirmation de la presence d'Iguanidae (Reptilia, Lacertilia) dans l'Eocene europeen. Comptes Rendus de l'Academie des Sciences Paris, 305, 633 - 636." type="journal article" year="1987">Augé 1987</bibRefCitation>
|
||
). The remaining material previously assigned to
|
||
<taxonomicName id="4C1F4D06FFE1FFAEC66594C3FC27F8F9" box="[888,1016,1778,1801]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE1FFAEC66594C3FC54F8F9" box="[888,907,1778,1801]" italics="true" pageId="2" pageNumber="267">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE1FFAEC68294C3FC27F8F9" box="[927,1016,1778,1801]" italics="true" pageId="2" pageNumber="267">rugosus</emphasis>
|
||
</taxonomicName>
|
||
(one maxilla, one frontal and two vertebrae) all possess gekkotan features.
|
||
</paragraph>
|
||
<paragraph id="8BA03685FFE1FFAFC5DA9508FDEFFDCD" blockId="2.[151,1436,1417,2018]" lastBlockId="3.[151,1436,151,896]" lastPageId="3" lastPageNumber="268" pageId="2" pageNumber="267">
|
||
The diagnostic characters of the genus
|
||
<taxonomicName id="4C1F4D06FFE1FFAEC7609508FCC0F8A2" box="[637,799,1849,1874]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA97FFE1FFAEC7609508FCC0F8A2" box="[637,799,1849,1874]" italics="true" pageId="2" pageNumber="267">Cadurcogekko</emphasis>
|
||
</taxonomicName>
|
||
are, according to
|
||
<bibRefCitation id="EF8E4B74FFE1FFAEC6F7950BFBA3F8A2" author="Auge" box="[1002,1148,1849,1874]" pageId="2" pageNumber="267" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé (2005)</bibRefCitation>
|
||
: the rugose surface of the facial process of the maxilla, an elongated groove associated with the most posterior neurovascular foramen of the maxilla, a postnarial anterodorsal depression of the maxilla, and presence of a supradental gutter. Thus, most diagnostic characters reported by
|
||
<bibRefCitation id="EF8E4B74FFE1FFAEC7019597FD72F84E" author="Auge" box="[540,685,1957,1982]" pageId="2" pageNumber="267" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé (2005)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="4C1F4D06FFE1FFAEC7FF9597FC81F84D" box="[738,862,1958,1981]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE1FFAEC7FF9597FD2AF84D" box="[738,757,1958,1981]" italics="true" pageId="2" pageNumber="267">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE1FFAEC6189597FC81F84D" box="[773,862,1958,1981]" italics="true" pageId="2" pageNumber="267">rugosus</emphasis>
|
||
</taxonomicName>
|
||
focus on the maxilla. USTL, PRA 9 presents the diagnostic characters of
|
||
<taxonomicName id="4C1F4D06FFE1FFAEC4AC95F8FD8CF812" box="[433,595,1993,2018]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="267" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA97FFE1FFAEC4AC95F8FD8CF812" box="[433,595,1993,2018]" italics="true" pageId="2" pageNumber="267">Cadurcogekko</emphasis>
|
||
</taxonomicName>
|
||
, and its smaller size, the lower tooth-count and a coarser ornamentation relative to the
|
||
<typeStatus id="54A48827FFE0FFAFC42092A9FEB2FF40" box="[317,365,152,176]" pageId="3" pageNumber="268">type</typeStatus>
|
||
species have been interpreted as characters of interspecific variation (
|
||
<bibRefCitation id="EF8E4B74FFE0FFAFC18D92A9FACDFF40" author="Auge" box="[1168,1298,151,176]" pageId="3" pageNumber="268" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B74FFE0FFAFC03D92A9FF0BFF24" author="Daza" pageId="3" pageNumber="268" refString="Daza, J. D., Bauer, A. M. & Snively, E. (2014) On the gekkotan fossil record. Anatomical Record, 297, 433 - 462. http: // dx. doi. org / 10.1002 / ar. 22856" type="journal article" year="2014">
|
||
Daza
|
||
<emphasis id="B96BEA97FFE0FFAFC07F92A8FA43FF40" box="[1378,1436,151,176]" italics="true" pageId="3" pageNumber="268">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EF8E4B74FFE0FFAFC5F4928DFEA5FF25" author="Auge" box="[233,378,188,213]" pageId="3" pageNumber="268" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé (2005)</bibRefCitation>
|
||
already noted that
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC746928CFD3AFF24" box="[603,741,189,212]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="piveteaui">
|
||
<emphasis id="B96BEA97FFE0FFAFC746928CFDB1FF24" box="[603,622,189,212]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC762928CFD3AFF24" box="[639,741,189,212]" italics="true" pageId="3" pageNumber="268">piveteaui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC63C928CFC42FF24" box="[801,925,189,212]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE0FFAFC63C928CFCEBFF24" box="[801,820,189,212]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC659928CFC42FF24" box="[836,925,189,212]" italics="true" pageId="3" pageNumber="268">rugosus</emphasis>
|
||
</taxonomicName>
|
||
were quite different and he expressed some doubts about the taxonomic position of
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC77792D1FD38FF08" box="[618,743,224,248]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE0FFAFC77792D1FDA2FF07" box="[618,637,224,247]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC79392D0FD38FF08" box="[654,743,225,248]" italics="true" pageId="3" pageNumber="268">rugosus</emphasis>
|
||
</taxonomicName>
|
||
. The similarities of the maxillae, shared only by these two species, were considered enough evidence for generic referral, but it is clear now that the contradiction between having a similar maxilla and exhibiting strong differences in the dentary was because the latter bone belonged to a different taxon, rendering
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC4DB937CFD9BFE94" box="[454,580,333,356]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE0FFAFC4DB937CFE06FE94" box="[454,473,333,356]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC4F6937CFD9BFE94" box="[491,580,333,356]" italics="true" pageId="3" pageNumber="268">rugosus</emphasis>
|
||
</taxonomicName>
|
||
a chimaera. A review of the maxillary characters supports the referral of USTL, PRA 9 to
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC446935EFE22FE78" box="[347,509,367,392]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA97FFE0FFAFC446935EFE22FE78" box="[347,509,367,392]" italics="true" pageId="3" pageNumber="268">Cadurcogekko</emphasis>
|
||
</taxonomicName>
|
||
, and its status as a different species from
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC6CE9341FB92FE78" box="[979,1101,368,392]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="pivetaui">
|
||
<emphasis id="B96BEA97FFE0FFAFC6CE9341FC39FE77" box="[979,998,368,391]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC6EE9340FB92FE78" box="[1011,1101,369,392]" italics="true" pageId="3" pageNumber="268">pivetaui</emphasis>
|
||
</taxonomicName>
|
||
, but the identity of the rest of material is more problematic.
|
||
<bibRefCitation id="EF8E4B74FFE0FFAFC4FA93A5FDABFE5D" author="Auge" box="[487,628,404,429]" pageId="3" pageNumber="268" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé (2005)</bibRefCitation>
|
||
provided no explicit reasons to refer the vertebrae and dentary fragment to
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC5AE9389FEF3FE20" box="[179,300,440,464]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B96BEA97FFE0FFAFC5AE9389FF19FE3F" box="[179,198,440,463]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC5CE9388FEF3FE20" box="[211,300,441,464]" italics="true" pageId="3" pageNumber="268">rugosus</emphasis>
|
||
</taxonomicName>
|
||
. The main reason for the referral was indeed that they represented supposedly gekkotan material from the same locality as the
|
||
<typeStatus id="54A48827FFE0FFAFC4BF93ECFE0DFE05" box="[418,466,477,501]" pageId="3" pageNumber="268">type</typeStatus>
|
||
specimen. Because
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC7A893ECFCE3FE04" box="[693,828,477,500]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="piveteaui">
|
||
<emphasis id="B96BEA97FFE0FFAFC7A893ECFD17FE04" box="[693,712,477,500]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC7CB93ECFCE3FE04" box="[726,828,477,500]" italics="true" pageId="3" pageNumber="268">piveteaui</emphasis>
|
||
</taxonomicName>
|
||
has been also identified at Sindou D (
|
||
<bibRefCitation id="EF8E4B74FFE0FFAFC1FB93EDFAB8FE05" author="Auge" box="[1254,1383,476,501]" pageId="3" pageNumber="268" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé 2005</bibRefCitation>
|
||
), the smaller size—comparable to that of the maxilla—of the additional material (vertebrae and frontal) was used to discriminate between the two forms.
|
||
</paragraph>
|
||
<paragraph id="8BA03685FFE0FFAFC5DA9079FA97FC70" blockId="3.[151,1436,151,896]" pageId="3" pageNumber="268">
|
||
<collectingCountry id="F3087615FFE0FFAFC5DA9079FF39FD90" box="[199,230,584,608]" name="American Samoa" pageId="3" pageNumber="268">As</collectingCountry>
|
||
explained above, the
|
||
<typeStatus id="54A48827FFE0FFAFC4C19079FDD3FD90" box="[476,524,584,608]" pageId="3" pageNumber="268">type</typeStatus>
|
||
dentary and the referred fragmentary dentary are no longer considered gekkotan, and identification of the rest of gekkotan material as part of
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC657905DFC33FD75" box="[842,1004,620,645]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA97FFE0FFAFC657905DFC33FD75" box="[842,1004,620,645]" italics="true" pageId="3" pageNumber="268">Cadurcogekko</emphasis>
|
||
</taxonomicName>
|
||
is difficult. The only exception is the frontal.
|
||
<bibRefCitation id="EF8E4B74FFE0FFAFC5EF90A1FE5DFD58" author="Auge" box="[242,386,655,680]" pageId="3" pageNumber="268" refString="Auge, M. (2005) Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Historie naturelle, 192, 1 - 369." type="journal article" year="2005">Augé (2005)</bibRefCitation>
|
||
noted that the size of this frontal was smaller to that of
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC11390A1FB48FD58" box="[1038,1175,656,680]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="piveteaui">
|
||
<emphasis id="B96BEA97FFE0FFAFC11390A1FBFEFD57" box="[1038,1057,656,679]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC12C90A0FB48FD58" box="[1073,1175,657,680]" italics="true" pageId="3" pageNumber="268">piveteaui</emphasis>
|
||
</taxonomicName>
|
||
. Ornamentation of the frontal from Sindou D closely matches that of the maxilla from Les Pradigues (here proposed as the
|
||
<typeStatus id="54A48827FFE0FFAFC1E69085FA82FD3D" box="[1275,1373,692,717]" pageId="3" pageNumber="268" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC09F9084FF0BFD00" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="verus">
|
||
<emphasis id="B96BEA97FFE0FFAFC09F9084FA4AFD3C" box="[1410,1429,693,716]" italics="true" pageId="3" pageNumber="268">C</emphasis>
|
||
.
|
||
<emphasis id="B96BEA97FFE0FFAFC58A90E8FF0BFD00" box="[151,212,729,752]" italics="true" pageId="3" pageNumber="268">verus</emphasis>
|
||
</taxonomicName>
|
||
). Likewise, some substantial differences of dermal ornamentation and frontal morphology serve to differentiate the currently recognized
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC75C90CDFD3CFCE5" box="[577,739,764,789]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA97FFE0FFAFC75C90CDFD3CFCE5" box="[577,739,764,789]" italics="true" pageId="3" pageNumber="268">Cadurcogekko</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EF8E4B74FFE0FFAFC65690CDFBD3FCE4" author="Daza" box="[843,1036,764,789]" pageId="3" pageNumber="268" refString="Daza, J. D., Bauer, A. M. & Snively, E. (2014) On the gekkotan fossil record. Anatomical Record, 297, 433 - 462. http: // dx. doi. org / 10.1002 / ar. 22856" type="journal article" year="2014">
|
||
Daza
|
||
<emphasis id="B96BEA97FFE0FFAFC69390CCFC60FCE5" box="[910,959,764,789]" italics="true" pageId="3" pageNumber="268">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). The referral of the non-diagnostic material (two dorsal vertebrae) to one of the two species of
|
||
<taxonomicName id="4C1F4D06FFE0FFAFC64B912EFC26FCC8" box="[854,1017,799,824]" class="Reptilia" family="Gekkonidae" genus="Cadurcogekko" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="268" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA97FFE0FFAFC64B912EFC26FCC8" box="[854,1017,799,824]" italics="true" pageId="3" pageNumber="268">Cadurcogekko</emphasis>
|
||
</taxonomicName>
|
||
present at the locality is even more difficult, and can only be justified in terms of size. Because there is no argument in favor or against this specific allocation, we accept Augé’s (2005) original referral as correct until new evidence sheds light on this issue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |