301 lines
36 KiB
XML
301 lines
36 KiB
XML
<document id="33B867DBC5514ED5E34E2F9B128BEBDC" ID-DOI="10.5281/zenodo.5167725" ID-ISSN="1942-1354" ID-Zenodo-Dep="5167725" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270427885" checkinUser="felipe" docAuthor="Austin, George T. & Warren, Andrew D." docDate="2009" docId="03B687432D63854C3DDDFA5725821155" docLanguage="en" docName="InsectaMundi.0089.1-55.pdf" docOrigin="Insecta Mundi 2009 (89)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Neposa Austin and A. Warren 2009, new genus" docType="treatment" docVersion="3" lastPageNumber="40" masterDocId="FF8FFF3B2D4485653D1DFFF62651142B" masterDocTitle="New looks at and for Onespa, Buzyges, and Librita (Lepidoptera: Hesperiidae: Hesperiinae), with new combinations and descriptions of a new genus and six new species" masterLastPageNumber="55" masterPageNumber="1" pageNumber="38" updateTime="1699038508896" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="F64D8408F840790395A7DA7ACC650F6A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BA2FB0958B1398D09563FC229CB900EA">
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<mods:title id="DD44993AD09ABBAA622AB06AE47DF452">New looks at and for Onespa, Buzyges, and Librita (Lepidoptera: Hesperiidae: Hesperiinae), with new combinations and descriptions of a new genus and six new species</mods:title>
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</mods:titleInfo>
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<mods:name id="123ED8592F10CE131EA25C7F1B81B242" type="personal">
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<mods:namePart id="4AAD8BBBD3BB83688E8D3BC69152DE70">Austin, George T.</mods:namePart>
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</mods:name>
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<mods:name id="BF7DC6AB1E633964B02F7BD4AB9D0B1D" type="personal">
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<mods:role id="236DE8AC4569A1A0FB686A2AFF5E9C60">
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<mods:roleTerm id="C51050976972EB4FBBE99701CF806C7D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7730D5CEDD9CA56D3AA59C9CB7A6665D">Warren, Andrew D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="03221235322EFCF92CEB8B444B4DD554">text</mods:typeOfResource>
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<mods:titleInfo id="9EDB1EF670CAF8D7E39C5409DF95306B">
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<mods:title id="272DB1ABADB637F9A60D32B0DBACD3EA">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="B60AE41146B1758E5DBB0AD5A2E97164">
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<mods:date id="CBEE6BC1387962F513E320D8632CAAF9">2009</mods:date>
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<mods:detail id="5C248340F06E6746DD5E3D48BB34243A" type="pubDate">
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<mods:number id="DC0D9E6B7A84E4D99EA4F4651A6FCE5D">2009-07-24</mods:number>
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</mods:detail>
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<mods:detail id="210487153A609D3880DD130B73382950" type="volume">
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<mods:number id="5C9D56F3CD7845675A6EB43AA0FF968F">2009</mods:number>
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</mods:detail>
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<mods:detail id="F650A1FAD54E048070B1049CD913C6FB" type="issue">
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<mods:number id="18CE8AC8722206FB43BA439CDA048121">89</mods:number>
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<mods:extent id="889F68D227BF893A8850240552E17996" unit="page">
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<mods:start id="C0BF1E52BF22BC230F95483A28C5BB75">1</mods:start>
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<mods:classification id="B39088E78FE9F900A639DB941EC90175">journal article</mods:classification>
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<mods:identifier id="65F6C399943838D6D5442DA6B0ECBB1E" type="DOI">10.5281/zenodo.5167725</mods:identifier>
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<mods:identifier id="A9A4A6ECB1129968CD2FA8CC2884D94B" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="8ACED45C3C277C7A8D3498670BEE00AD" type="Zenodo-Dep">5167725</mods:identifier>
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</mods:mods>
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<treatment id="03B687432D63854C3DDDFA5725821155" LSID="urn:lsid:plazi:treatment:03B687432D63854C3DDDFA5725821155" httpUri="http://treatment.plazi.org/id/03B687432D63854C3DDDFA5725821155" lastPageId="41" lastPageNumber="40" pageId="39" pageNumber="38">
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<subSubSection id="C30565DE2D6385423DDDFA5724BF1193" box="[192,750,1440,1464]" pageId="39" pageNumber="38" type="nomenclature">
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<paragraph id="8BA036552D6385423DDDFA5724BF1193" blockId="39.[192,750,1440,1464]" box="[192,750,1440,1464]" pageId="39" pageNumber="38">
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<heading id="D0E881392D6385423DDDFA5724BF1193" bold="true" box="[192,750,1440,1464]" fontSize="10" level="2" pageId="39" pageNumber="38" reason="3">
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<emphasis id="B96BEA472D6385423DDDFA5724BF1193" bold="true" box="[192,750,1440,1464]" pageId="39" pageNumber="38">
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<taxonomicName id="4C1F4DD62D6385423DDDFA5724001193" authority="Austin and A. Warren" authorityName="Austin and A. Warren" authorityYear="2009" box="[192,593,1440,1464]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B96BEA472D6385423DDDFA5727751193" bold="true" box="[192,292,1441,1464]" italics="true" pageId="39" pageNumber="38">Neposa</emphasis>
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Austin and A. Warren
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</taxonomicName>
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,
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<taxonomicNameLabel id="A258573C2D6385423F40FA5624BF1193" box="[605,750,1440,1464]" pageId="39" pageNumber="38" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565DE2D6385423DDDFA1724C21293" pageId="39" pageNumber="38" type="materials_examined">
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<paragraph id="8BA036552D6385423DDDFA1725CC11D3" blockId="39.[192,925,1504,1529]" box="[192,925,1504,1529]" pageId="39" pageNumber="38">
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<typeStatus id="54A488F72D6385423DDDFA1726AC11D2" box="[192,253,1505,1529]" pageId="39" pageNumber="38">Type</typeStatus>
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species:
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<taxonomicName id="4C1F4DD62D6385423C79FA17249D11D3" authority="(Godman, 1900)" authorityName="Austin & Warren" authorityYear="2009" baseAuthorityName="Godman" baseAuthorityYear="1900" box="[356,716,1504,1528]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="heras">
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<emphasis id="B96BEA472D6385423C79FA17245511D3" box="[356,516,1504,1528]" italics="true" pageId="39" pageNumber="38">Neposa heras</emphasis>
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(Godman, 1900)
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</taxonomicName>
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, here designated.
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</paragraph>
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<paragraph id="8BA036552D6385423DEDF9D624C21293" blockId="39.[192,1441,1568,1720]" pageId="39" pageNumber="38">
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The critical examination of
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<taxonomicName id="4C1F4DD62D6385423F25F9D624DF1213" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[568,654,1568,1592]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEA472D6385423F25F9D624DF1213" box="[568,654,1568,1592]" italics="true" pageId="39" pageNumber="38">Librita</emphasis>
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</taxonomicName>
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above confirmed that one species considered congeneric by
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<bibRefCitation id="EF8E4BA42D6385423849F9D727411273" author="Evans, W. H." pageId="39" pageNumber="38" refId="ref46128" refString="Evans, W. H. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV (groups H to P), Hesperiinae and Megathyminae. British Museum (Natural History): London. 499 p." type="book" year="1955">Evans (1955)</bibRefCitation>
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,
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<taxonomicName id="4C1F4DD62D6385423C07F9B627EA1273" authorityName="Evans" authorityYear="1955" box="[282,443,1600,1624]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="raspa">
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<emphasis id="B96BEA472D6385423C07F9B627EA1273" box="[282,443,1600,1624]" italics="true" pageId="39" pageNumber="38">Librita raspa</emphasis>
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</taxonomicName>
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, was indeed a
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<taxonomicName id="4C1F4DD62D6385423F74F9B724AB1273" authorityName="Godman" authorityYear="1900" box="[617,762,1601,1624]" class="Insecta" family="Hesperiidae" genus="Paratrytone" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEA472D6385423F74F9B724AB1273" box="[617,762,1601,1624]" italics="true" pageId="39" pageNumber="38">Paratrytone</emphasis>
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</taxonomicName>
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(see above), and revealed that a second species placed in
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<taxonomicName id="4C1F4DD62D6385423DDDF99627461253" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[192,279,1632,1656]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEA472D6385423DDDF99627461253" box="[192,279,1632,1656]" italics="true" pageId="39" pageNumber="38">Librita</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1F4DD62D6385423C39F996278E1253" authorityName="Godman" authorityYear="1900" box="[292,479,1632,1656]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="heras">
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<emphasis id="B96BEA472D6385423C39F996278E1253" box="[292,479,1632,1656]" italics="true" pageId="39" pageNumber="38">Augiades heras</emphasis>
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</taxonomicName>
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, was also not congeneric. Since no existing genus is available for this,
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<taxonomicName id="4C1F4DD62D6385423831F99723D71253" authorityName="Austin and A. Warren" authorityYear="2009" box="[1324,1414,1633,1656]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEA472D6385423831F99723D71253" box="[1324,1414,1633,1656]" italics="true" pageId="39" pageNumber="38">Neposa</emphasis>
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</taxonomicName>
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is here proposed and described to envelope
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<taxonomicName id="4C1F4DD62D6385423FBFF977255412B3" authorityName="Godman" authorityYear="1900" box="[674,773,1664,1688]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="heras">
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<emphasis id="B96BEA472D6385423FBFF977255412B3" box="[674,773,1664,1688]" italics="true" pageId="39" pageNumber="38">N. heras</emphasis>
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</taxonomicName>
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and three undescribed species, all known from southern
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<collectingCountry id="F30876C52D6385423DDDF95627471293" box="[192,278,1696,1720]" name="Mexico" pageId="39" pageNumber="38">Mexico</collectingCountry>
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and northern Central America.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565DE2D63854D3DDDF91627FB12B2" lastPageId="40" lastPageNumber="39" pageId="39" pageNumber="38" type="description">
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<paragraph id="8BA036552D63854D3DDDF916243A1792" blockId="39.[192,1440,1760,1881]" lastBlockId="40.[192,1441,897,1689]" lastPageId="40" lastPageNumber="39" pageId="39" pageNumber="38">
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<emphasis id="B96BEA472D6385423DDDF916273B12D3" bold="true" box="[192,362,1760,1784]" pageId="39" pageNumber="38">Description.</emphasis>
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Palpi quadrate, 3rd segment robust, nearly erect, not exceeding scales of second segment. Antennae more than 1/2 costa on males (51-53%), about 1/2 costa on females (48-49%); club stout, 1/3 or more of shaft (30-37% on males, 40-45% on females), bent to constricted apiculus beyond thickest part; nudum varies from 11 to 13 segments, subequally divided between club and apiculus (4-5 segments on club); shaft entirely dark or yellow checked with black above, black and yellow or yellow-orange on venter; nudum red-brown to yellow-orange.
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</paragraph>
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<caption id="DF6066DD2D6C854D3DDDFCF1240A177C" ID-DOI="http://doi.org/10.5281/zenodo.5167753" ID-Zenodo-Dep="5167753" httpUri="https://zenodo.org/record/5167753/files/figure.png" pageId="40" pageNumber="39" startId="40.[192,276,775,797]" targetBox="[283,1344,194,750]" targetPageId="40">
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<paragraph id="8BA036552D6C854D3DDDFCF1240A177C" blockId="40.[192,1440,775,855]" pageId="40" pageNumber="39">
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<emphasis id="B96BEA472D6C854D3DDDFCF127301737" bold="true" box="[192,353,775,797]" pageId="40" pageNumber="39">Figure 90-92.</emphasis>
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Female genitalia of
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<taxonomicName id="4C1F4DD62D6C854D3F5EFCFE24CF1736" authorityName="Godman" authorityYear="1900" box="[579,670,776,797]" class="Insecta" family="Hesperiidae" genus="Buzyges" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEA472D6C854D3F5EFCFE24CF1736" box="[579,670,776,797]" italics="true" pageId="40" pageNumber="39">Buzyges</emphasis>
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</taxonomicName>
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(full data in text; shown are left lateral and ventral views; scale bar = 1 mm).
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<emphasis id="B96BEA472D6C854D3C05FCD2276C1711" bold="true" box="[280,317,804,826]" pageId="40" pageNumber="39">90)</emphasis>
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<taxonomicName id="4C1F4DD62D6C854D3C58FCD327E81712" authorityName="Godman" authorityYear="1900" box="[325,441,804,826]" class="Insecta" family="Hesperiidae" genus="Buzyges" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="idothea">
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<emphasis id="B96BEA472D6C854D3C58FCD327E81712" box="[325,441,804,826]" italics="true" pageId="40" pageNumber="39">B. idothea</emphasis>
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</taxonomicName>
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, Costa Rica (SRS #3754);
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<emphasis id="B96BEA472D6C854D3FFBFCD2255A1711" bold="true" box="[742,779,804,826]" pageId="40" pageNumber="39">91)</emphasis>
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<taxonomicName id="4C1F4DD62D6C854D3E0EFCD325381712" authorityName="Austin & Warren" authorityYear="2009" baseAuthorityName="Mabille" baseAuthorityYear="1883" box="[787,873,804,826]" class="Insecta" family="Hesperiidae" genus="Buzyges" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="rolla">
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<emphasis id="B96BEA472D6C854D3E0EFCD325381712" box="[787,873,804,826]" italics="true" pageId="40" pageNumber="39">B. rolla</emphasis>
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</taxonomicName>
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, Costa Rica (GTA #14057);
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<emphasis id="B96BEA472D6C854D39B5FCD2229C1711" bold="true" box="[1192,1229,804,826]" pageId="40" pageNumber="39">92)</emphasis>
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<taxonomicName id="4C1F4DD62D6C854D39C8FCD323C81712" authorityName="Austin & Warren" authorityYear="2009" box="[1237,1433,804,826]" class="Insecta" family="Hesperiidae" genus="Buzyges" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="mellanaformis">
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<emphasis id="B96BEA472D6C854D39C8FCD323C81712" box="[1237,1433,804,826]" italics="true" pageId="40" pageNumber="39">B. mellanaformis</emphasis>
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</taxonomicName>
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, Costa Rica, paratype (GTA #14058).
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</paragraph>
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</caption>
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<paragraph id="8BA036552D6C854D3DEDFC37231E1072" blockId="40.[192,1441,897,1689]" pageId="40" pageNumber="39">
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Wings somewhat produced; forewing apex 1.4 times length along vein 2A; hindwing longest at CuA
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<subScript id="179B34102D6C854D388DFC2723C817F4" attach="left" box="[1424,1433,977,991]" fontSize="6" pageId="40" pageNumber="39">1</subScript>
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, nearly as long at 2A, about equally long at veins Sc+R
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<subScript id="179B34102D6C854D3E78FC07253F17D4" attach="left" box="[869,878,1009,1023]" fontSize="6" pageId="40" pageNumber="39">1</subScript>
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and 3A. Male with bipartite stigma (Fig. 62) flanked by a lower brush patch (
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<emphasis id="B96BEA472D6C854D3F58FBF424DB1032" box="[581,650,1026,1049]" italics="true" pageId="40" pageNumber="39">sensu</emphasis>
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<bibRefCitation id="EF8E4BA42D6C854D3F92FBF725151032" author="MacNeill, C. D." box="[655,836,1025,1049]" pageId="40" pageNumber="39" pagination="1 - 221" refId="ref46881" refString="MacNeill, C. D. 1964. The skippers of the genus Hesperia in western North America with special reference to California (Lepidoptera: Hesperiidae). University of California Publications in Entomology 35: 1 - 221." type="journal article" year="1964">MacNeill 1964</bibRefCitation>
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). Hindwing origin Rs nearer to cell end than base. Forewing origin CuA
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<subScript id="179B34102D6C854D3CDBFBC7279E1014" attach="left" box="[454,463,1073,1087]" fontSize="6" pageId="40" pageNumber="39">2</subScript>
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nearer to origin of CuA
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<subScript id="179B34102D6C854D3FE6FBC725551014" attach="left" box="[763,772,1073,1087]" fontSize="6" pageId="40" pageNumber="39">1</subScript>
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than to base of wing. Sexual dimorphism prominent. Wings dark brown with orange (male) or yellowish and/or orange and white (female) maculation.
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</paragraph>
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<paragraph id="8BA036552D6C854D3DEDFB94231F10B2" blockId="40.[192,1441,897,1689]" pageId="40" pageNumber="39">Meso- and metatibiae spined with long fringes of setiform scales on both tibia and femur; mesotibiae with one pair spurs, metatibiae two pairs; outer spur in each pair 1/2 to 2/3 length of inner spur.</paragraph>
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<paragraph id="8BA036552D6C854D3DEDFB5726A311D2" blockId="40.[192,1441,897,1689]" pageId="40" pageNumber="39">Male genitalia with gnathos bifid, arms slender and connivent, uncus short and narrow, weakly bilobed at end, narrower or about width of gnathos in dorsal view, longer or shorter than gnathos; separation of gnathos and uncus in lateral view shallow. Tegumen flaring cephalad, ventral arm with dorsal arm of saccus combining into a structure bent well below its middle, anterior arm of saccus short to moderately long, narrowing to a bluntly pointed or curved cephalic end in ventral view. Valvae simple and unarticulated, with costa/ampulla relatively straight and undifferentiated between themselves and harpe, ampulla with small dorsal triangular process caudad, harpe angled or curved, serrate on caudal edge before another sharply pointed process. Aedeagus stout, relatively short (1.1-1.4 times length of valva) one species with curved titillator on venter near caudal end, with cornuti consisting of two heavily sclerotized structures (narrowing abruptly caudad) and (on one species) a flexible, lightly sclerotized and spinulose pad.</paragraph>
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<paragraph id="8BA036552D6C854D3DEDF9F727FB12B2" blockId="40.[192,1441,897,1689]" pageId="40" pageNumber="39">
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Female genitalia characterized by a moderately long (
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<quantity id="4CE79BB02D6C854D3E6CF9F7259F1232" box="[881,974,1537,1561]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="40" pageNumber="39" unit="mm" value="2.3">2.3 mm</quantity>
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), sclerotized, and centrally constricted, straight, and slightly twisted ductus bursae that expands cephalad in lateral view. Lamella antevaginalis very broad, lobate, and shallowly excavated caudad; lamella postvaginalis also broad with V-shaped caudal indentation; ostium bursae broad and shallow, twice as broad as deep. Corpus bursae wrinkled longitudinally, no signa.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565DE2D6C854D3DDDF93723F11312" pageId="40" pageNumber="39" type="etymology">
|
|
<paragraph id="8BA036552D6C854D3DDDF93723F11312" blockId="40.[192,1440,1729,1849]" pageId="40" pageNumber="39">
|
|
<emphasis id="B96BEA472D6C854D3DDDF937270D12F2" bold="true" box="[192,348,1729,1753]" pageId="40" pageNumber="39">Etymology.</emphasis>
|
|
<taxonomicName id="4C1F4DD62D6C854D3C7CF93427EA12F2" authorityName="Austin and A. Warren" authorityYear="2009" box="[353,443,1730,1753]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6C854D3C7CF93427EA12F2" box="[353,443,1730,1753]" italics="true" pageId="40" pageNumber="39">Neposa</emphasis>
|
|
</taxonomicName>
|
|
is a meaningless anagram of
|
|
<taxonomicName id="4C1F4DD62D6C854D3E03F934252512F2" authorityName="Scudder" authorityYear="1872" box="[798,884,1730,1753]" class="Insecta" family="Hesperiidae" genus="Poanes" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6C854D3E03F934252512F2" box="[798,884,1730,1753]" italics="true" pageId="40" pageNumber="39">Poanes</emphasis>
|
|
</taxonomicName>
|
|
as was
|
|
<taxonomicName id="4C1F4DD62D6C854D3ECDF937234F12F2" authority="(Steinhauser 1974)" baseAuthorityName="Steinhauser" baseAuthorityYear="1974" box="[976,1310,1729,1753]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6C854D3ECDF937227D12F3" box="[976,1068,1729,1752]" italics="true" pageId="40" pageNumber="39">Onespa</emphasis>
|
|
(
|
|
<bibRefCitation id="EF8E4BA42D6C854D3927F937234412F2" author="Steinhauser, S. R." box="[1082,1301,1729,1753]" pageId="40" pageNumber="39" pagination="1 - 38" refId="ref47739" refString="Steinhauser, S. R. 1974. Notes on neotropical Nymphalidae and Hesperiidae with descriptions of new species and subspecies and a new genus. Bulletin of the Allyn Museum 22: 1 - 38." type="journal article" year="1974">Steinhauser 1974</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. The name was usurped from the notes of the late Steve Steinhauser (those now at MGCL). Steve apparently originally was to propose this name for a genus to embrace
|
|
<taxonomicName id="4C1F4DD62D6C854D3E5EF8F725A81332" authorityName="Mabille" authorityYear="1883" box="[835,1017,1793,1817]" class="Insecta" family="Hesperiidae" genus="Pamphila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="rolla">
|
|
<emphasis id="B96BEA472D6C854D3E5EF8F725A81332" box="[835,1017,1793,1817]" italics="true" pageId="40" pageNumber="39">Pamphila rolla</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1F4DD62D6C854D392FF8F422841332" authorityName="Freeman" authorityYear="1979" box="[1074,1237,1793,1817]" class="Insecta" family="Hesperiidae" genus="Poanes" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="species" species="benito">
|
|
<emphasis id="B96BEA472D6C854D392FF8F422841332" box="[1074,1237,1793,1817]" italics="true" pageId="40" pageNumber="39">Poanes benito</emphasis>
|
|
</taxonomicName>
|
|
(now included in
|
|
<taxonomicName id="4C1F4DD62D6C854D3DDDF8D427721312" authorityName="Godman" authorityYear="1900" box="[192,291,1826,1849]" class="Insecta" family="Hesperiidae" genus="Buzyges" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6C854D3DDDF8D427721312" box="[192,291,1826,1849]" italics="true" pageId="40" pageNumber="39">Buzyges</emphasis>
|
|
</taxonomicName>
|
|
), which he, along with
|
|
<bibRefCitation id="EF8E4BA42D6C854D3F2AF8D424B71312" author="Burns, J. M." box="[567,742,1825,1849]" pageId="40" pageNumber="39" pagination="1 - 23" refId="ref45324" refString="Burns, J. M. 1992 a. Genitalic recasting of Poanes and Paratrytone (Hesperiidae). Journal of the Lepidopterists' Society 46: 1 - 23." type="journal article" year="1992">Burns (1992a)</bibRefCitation>
|
|
, had recognized as not being species of
|
|
<taxonomicName id="4C1F4DD62D6C854D39A4F8D4235E1312" authorityName="Scudder" authorityYear="1872" box="[1209,1295,1826,1849]" class="Insecta" family="Hesperiidae" genus="Poanes" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="39" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6C854D39A4F8D4235E1312" box="[1209,1295,1826,1849]" italics="true" pageId="40" pageNumber="39">Poanes</emphasis>
|
|
</taxonomicName>
|
|
(see above).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption id="DF6066DD2D6D854C3DDEFD0E257B1763" ID-DOI="http://doi.org/10.5281/zenodo.5167755" ID-Zenodo-Dep="5167755" httpUri="https://zenodo.org/record/5167755/files/figure.png" pageId="41" pageNumber="40" startId="41.[195,279,760,782]" targetBox="[290,1345,194,744]" targetPageId="41">
|
|
<paragraph id="8BA036552D6D854C3DDEFD0E257B1763" blockId="41.[195,1440,760,840]" pageId="41" pageNumber="40">
|
|
<emphasis id="B96BEA472D6D854C3DDEFD0E27351726" bold="true" box="[195,356,760,782]" pageId="41" pageNumber="40">Figure 93-95.</emphasis>
|
|
Female genitalia of
|
|
<taxonomicName id="4C1F4DD62D6D854C3F58FD0F24C41725" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[581,661,761,782]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3F58FD0F24C41725" box="[581,661,761,782]" italics="true" pageId="41" pageNumber="40">Librita</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1F4DD62D6D854C3FD7FD0F254A1725" authorityName="Austin and A. Warren" authorityYear="2009" box="[714,795,761,782]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3FD7FD0F254A1725" box="[714,795,761,782]" italics="true" pageId="41" pageNumber="40">Neposa</emphasis>
|
|
</taxonomicName>
|
|
(full data in text; shown are left lateral and ventral views; scale bar = 1 mm).
|
|
<emphasis id="B96BEA472D6D854C3C89FCE327EB1700" bold="true" box="[404,442,789,811]" pageId="41" pageNumber="40">93)</emphasis>
|
|
<taxonomicName id="4C1F4DD62D6D854C3CDDFCE024751701" authorityName="librita (Plotz" authorityYear="1886" box="[448,548,789,811]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="librita">
|
|
<emphasis id="B96BEA472D6D854C3CDDFCE024751701" box="[448,548,789,811]" italics="true" pageId="41" pageNumber="40">L. librita</emphasis>
|
|
</taxonomicName>
|
|
, Oaxaca, Mexico (GTA #14137);
|
|
<emphasis id="B96BEA472D6D854C3E92FCE325E51700" bold="true" box="[911,948,789,811]" pageId="41" pageNumber="40">94)</emphasis>
|
|
<taxonomicName id="4C1F4DD62D6D854C3EA7FCE022491701" authorityName="Godman" authorityYear="1900" box="[954,1048,789,811]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="heras">
|
|
<emphasis id="B96BEA472D6D854C3EA7FCE022491701" box="[954,1048,789,811]" italics="true" pageId="41" pageNumber="40">N. heras</emphasis>
|
|
</taxonomicName>
|
|
, Guerrero, Mexico (ADW #97-111);
|
|
<emphasis id="B96BEA472D6D854C3DDEFCC426B91763" bold="true" box="[195,232,818,840]" pageId="41" pageNumber="40">95)</emphasis>
|
|
<taxonomicName id="4C1F4DD62D6D854C3DF2FCC52702176C" authorityName="Austin and A. Warren" authorityYear="2009" box="[239,339,818,840]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="hestia">
|
|
<emphasis id="B96BEA472D6D854C3DF2FCC52702176C" box="[239,339,818,840]" italics="true" pageId="41" pageNumber="40">N. hestia</emphasis>
|
|
</taxonomicName>
|
|
, Veracruz, Mexico, holotype (SRS #3738).
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="C30565DE2D6D854C3DDDFC90244E17B5" pageId="41" pageNumber="40" type="distribution">
|
|
<paragraph id="8BA036552D6D854C3DDDFC90244E17B5" blockId="41.[192,1440,870,926]" pageId="41" pageNumber="40">
|
|
<emphasis id="B96BEA472D6D854C3DDDFC90247C1755" bold="true" box="[192,557,870,894]" pageId="41" pageNumber="40">Distribution and richness.</emphasis>
|
|
<taxonomicName id="4C1F4DD62D6D854C3F29FC9124DF1755" authorityName="Austin and A. Warren" authorityYear="2009" box="[564,654,871,894]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3F29FC9124DF1755" box="[564,654,871,894]" italics="true" pageId="41" pageNumber="40">Neposa</emphasis>
|
|
</taxonomicName>
|
|
embraces
|
|
<taxonomicName id="4C1F4DD62D6D854C3E0DFC90259C1755" authorityName="Godman" authorityYear="1900" box="[784,973,870,894]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="heras">
|
|
<emphasis id="B96BEA472D6D854C3E0DFC90259C1755" box="[784,973,870,894]" italics="true" pageId="41" pageNumber="40">Augiades heras</emphasis>
|
|
</taxonomicName>
|
|
and three undescribed species known from Guatelama and
|
|
<collectingCountry id="F30876C52D6D854C3CDCFC70244A17B5" box="[449,539,902,926]" name="Mexico" pageId="41" pageNumber="40">Mexico</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30565DE2D6D854C3DDDFC3025821155" pageId="41" pageNumber="40" type="diagnosis">
|
|
<paragraph id="8BA036552D6D854C3DDDFC30230D10F5" blockId="41.[192,1440,966,1406]" pageId="41" pageNumber="40">
|
|
<emphasis id="B96BEA472D6D854C3DDDFC30244B17F5" bold="true" box="[192,538,966,990]" pageId="41" pageNumber="40">Diagnosis and discussion.</emphasis>
|
|
Species of
|
|
<taxonomicName id="4C1F4DD62D6D854C3F85FC3124A117F5" authorityName="Austin and A. Warren" authorityYear="2009" box="[664,752,967,990]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3F85FC3124A117F5" box="[664,752,967,990]" italics="true" pageId="41" pageNumber="40">Neposa</emphasis>
|
|
</taxonomicName>
|
|
, although initially appearing similar to other hesperiines, including
|
|
<taxonomicName id="4C1F4DD62D6D854C3C21FC1127C917D5" baseAuthorityName="Steinhauser" baseAuthorityYear="1974" box="[316,408,999,1022]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3C21FC1127C917D5" box="[316,408,999,1022]" italics="true" pageId="41" pageNumber="40">Onespa</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1F4DD62D6D854C3CC8FC11246817D5" authorityName="Godman" authorityYear="1900" box="[469,569,999,1022]" class="Insecta" family="Hesperiidae" genus="Buzyges" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3CC8FC11246817D5" box="[469,569,999,1022]" italics="true" pageId="41" pageNumber="40">Buzyges</emphasis>
|
|
</taxonomicName>
|
|
, have a unique combination of superficial and genital traits.
|
|
<taxonomicName id="4C1F4DD62D6D854C3830FC1027521035" authorityName="Godman" authorityYear="1900" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="species" species="heras">
|
|
<emphasis id="B96BEA472D6D854C3830FC1027521035" italics="true" pageId="41" pageNumber="40">Augiades heras</emphasis>
|
|
</taxonomicName>
|
|
was considered congeneric with
|
|
<taxonomicName id="4C1F4DD62D6D854C3F94FBF0248E1035" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[649,735,1030,1054]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3F94FBF0248E1035" box="[649,735,1030,1054]" italics="true" pageId="41" pageNumber="40">Librita</emphasis>
|
|
</taxonomicName>
|
|
based largely upon their conspicuous stigmas and perhaps their somewhat similar valvae (
|
|
<bibRefCitation id="EF8E4BA42D6D854C3F56FBD025D81015" author="Godman, F. D. & O. Salvin" box="[587,905,1062,1086]" pageId="41" pageNumber="40" refId="ref46243" refString="Godman, F. D., and O. Salvin. 1879 - 1901. Biologia Centrali-Americana. Insecta. Lepidoptera- Rhopalocera. London: Delau. 1229 p." type="book" year="1879">Godman and Salvin 1879</bibRefCitation>
|
|
-1901,
|
|
<bibRefCitation id="EF8E4BA42D6D854C3EC7FBD1223E1015" author="Evans, W. H." box="[986,1135,1062,1086]" pageId="41" pageNumber="40" refId="ref46128" refString="Evans, W. H. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV (groups H to P), Hesperiinae and Megathyminae. British Museum (Natural History): London. 499 p." type="book" year="1955">Evans 1955</bibRefCitation>
|
|
). Notwithstanding those similarities,
|
|
<taxonomicName id="4C1F4DD62D6D854C3C43FBB127E91075" authorityName="Austin and A. Warren" authorityYear="2009" box="[350,440,1095,1118]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3C43FBB127E91075" box="[350,440,1095,1118]" italics="true" pageId="41" pageNumber="40">Neposa</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="4C1F4DD62D6D854C3F41FBB024E51075" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[604,692,1094,1118]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3F41FBB024E51075" box="[604,692,1094,1118]" italics="true" pageId="41" pageNumber="40">Librita</emphasis>
|
|
</taxonomicName>
|
|
by its bipartite stigma, presence of a lower brush patch, its overall brown color on the dorsal wings (
|
|
<taxonomicName id="4C1F4DD62D6D854C3FA9FB90255D1055" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[692,780,1126,1150]" class="Insecta" family="Hesperiidae" genus="Librita" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3FA9FB90255D1055" box="[692,780,1126,1150]" italics="true" pageId="41" pageNumber="40">Librita</emphasis>
|
|
</taxonomicName>
|
|
is a more obviously orange insect), spined tibiae, and numerous genital differences. Those latter include a shorter saccus, a more compact tegumen-uncus, a broad and often angled harpe, spike-like cornuti that expand cephalad on males, a broader sterigma, and a relatively longer, thinner, and twisted antrum with complex plate-like sclerotization on females.
|
|
</paragraph>
|
|
<paragraph id="8BA036552D6D854C3DEDFB1125821155" blockId="41.[192,1440,966,1406]" pageId="41" pageNumber="40">
|
|
<taxonomicName id="4C1F4DD62D6D854C3DEDFB11271910D5" authorityName="Austin and A. Warren" authorityYear="2009" box="[240,328,1255,1278]" class="Insecta" family="Hesperiidae" genus="Neposa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3DEDFB11271910D5" box="[240,328,1255,1278]" italics="true" pageId="41" pageNumber="40">Neposa</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="4C1F4DD62D6D854C3CC0FB11246610D5" baseAuthorityName="Steinhauser" baseAuthorityYear="1974" box="[477,567,1255,1278]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C3CC0FB11246610D5" box="[477,567,1255,1278]" italics="true" pageId="41" pageNumber="40">Onespa</emphasis>
|
|
</taxonomicName>
|
|
by its stigma, spined tibiae (note, however, that one species of
|
|
<taxonomicName id="4C1F4DD62D6D854C380BFB11232110D5" baseAuthorityName="Steinhauser" baseAuthorityYear="1974" box="[1302,1392,1255,1278]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C380BFB11232110D5" box="[1302,1392,1255,1278]" italics="true" pageId="41" pageNumber="40">Onespa</emphasis>
|
|
</taxonomicName>
|
|
has spined tibiae), shorter 2nd palpal segment, a broad and usually angled harpe, shorter and stouter aedeagus, and relatively shorter and twisted ductus bursae without a lateral pouch. It differs from
|
|
<taxonomicName id="4C1F4DD62D6D854C39F1FAD1231E1115" authorityName="Godman" authorityYear="1900" box="[1260,1359,1319,1342]" class="Insecta" family="Hesperiidae" genus="Buzyges" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="40" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEA472D6D854C39F1FAD1231E1115" box="[1260,1359,1319,1342]" italics="true" pageId="41" pageNumber="40">Buzyges</emphasis>
|
|
</taxonomicName>
|
|
by the presence of a stigma, a shorter 2nd palpal segment, a broad and usually angled harpe, differently shaped cornuti, and a twisted ductus bursae with internal sclerotization.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |