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<mods:title id="2FA4B10AACD539C8BA04C5F9B3565372">Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae)</mods:title>
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<taxonomicName id="4C1FA387FFB30460FE4FFEBDFD6CFE4E" ID-CoL="L5J" authority="Hussey, 1929" authorityName="Hussey" authorityYear="1929" box="[454,748,379,404]" class="Insecta" family="Largidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="tribe" tribe="Physopeltini">
<emphasis id="B96B0416FFB30460FE4FFEBDFD6CFE4E" bold="true" box="[454,748,379,404]" pageId="5" pageNumber="510">
Physopeltini
<bibRefCitation id="EF8EA5F5FFB30460FDD0FEBAFD6CFE4E" author="Hussey" box="[601,748,379,404]" firstAuthor="Hussey" pageId="5" pageNumber="510" refId="ref65286" refString="HUSSEY F. 1929: Fascicle 3. Pyrrhocoridae. In: HORVATH G. &amp; PARSHLEY H. M. (eds.): General Catalogue of the Hemiptera. Smith College, Northampton, 144 pp." type="book" year="1929">Hussey, 1929</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<paragraph id="8BA0D804FFB30460FFF8FE67FEFBFE68" blockId="5.[113,805,417,437]" box="[113,379,417,437]" pageId="5" pageNumber="510">
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<taxonomicName id="4C1FA387FFB30460FFF8FE67FEF7FE68" ID-CoL="L5J" authority="Hussey, 1929: 28" authorityName="Hussey" authorityPageNumber="28" authorityYear="1929" box="[113,375,417,437]" class="Insecta" family="Largidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="tribe" tribe="Physopeltini">
Physopeltini
<treatmentCitation id="0ABEFE15FFB30460FF68FE64FEF7FE68" author="HUSSEY F." box="[225,375,418,437]" page="28" pageId="5" pageNumber="510" year="1929">
<bibRefCitation id="EF8EA5F5FFB30460FF68FE64FEF7FE68" author="Hussey" box="[225,375,418,437]" firstAuthor="Hussey" page="28" pageId="5" pageNumber="510" refId="ref65286" refString="HUSSEY F. 1929: Fascicle 3. Pyrrhocoridae. In: HORVATH G. &amp; PARSHLEY H. M. (eds.): General Catalogue of the Hemiptera. Smith College, Northampton, 144 pp." type="book" year="1929">Hussey, 1929: 28</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="C3058B8FFFB30460FEF6FE64FCA5FE68" box="[383,805,417,437]" pageId="5" pageNumber="510" type="type_taxon">
<paragraph id="8BA0D804FFB30460FEF6FE64FCA5FE68" blockId="5.[113,805,417,437]" box="[383,805,417,437]" pageId="5" pageNumber="510">
<typeStatus id="54A466A6FFB30460FEF6FE64FE29FE68" box="[383,425,418,437]" pageId="5" pageNumber="510">Type</typeStatus>
genus:
<taxonomicName id="4C1FA387FFB30460FE65FE64FCA0FE68" ID-CoL="6PZW" authority="Amyot &amp; Serville, 1843" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[492,800,417,437]" class="Insecta" family="Largidae" genus="Physopelta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FE65FE64FDCBFE69" box="[492,587,418,436]" italics="true" pageId="5" pageNumber="510">Physopelta</emphasis>
<bibRefCitation id="EF8EA5F5FFB30460FDD9FE64FCA0FE68" author="AMYOT C. J. B. &amp; SERVILLE J. G. A." bookContentInfo="xxvi + 675 + 6 pp." box="[592,800,417,437]" journalOrPublisher="Roret, Paris" pageId="5" pageNumber="510" refId="ref62473" refString="AMYOT C. J. B. &amp; SERVILLE J. G. A. 1843: Histoire naturelle des Insectes. Hemipteres. Roret, Paris, xxvi + 675 + 6 pp." title="Histoire naturelle des Insectes. Hemipteres" type="book" year="1843">Amyot &amp; Serville, 1843</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3058B8FFFB30460FFF8FE0BFE64FD78" pageId="5" pageNumber="510" type="diagnosis">
<paragraph id="8BA0D804FFB30460FFF8FE0BFE64FD78" blockId="5.[113,1089,461,965]" pageId="5" pageNumber="510">
<emphasis id="B96B0416FFB30460FFF8FE0BFF66FE38" bold="true" box="[113,230,461,485]" pageId="5" pageNumber="510">Diagnosis.</emphasis>
Labium reaching between meso- and metacoxae. Callar lobe gibbose or not. Ventral surface of profemora with smooth longitudinal furrow along its entire length, denticles forming two longitudinal rows, one on each side of the furrow. Parameres short, directed obliquely towards midline of the genital chamber, not crossing each other, not surpassing ventral rim of pygophore. Parameres basally wide, middle portion forming rounded lobe with sensory setae on its ventral side; apical part narrowed, straight, apically narrowly rounded (
<figureCitation id="1324C481FFB30460FFF0FD4BFF66FD78" box="[121,230,653,677]" captionStart="Figs 612" captionStartId="9.[780,817,1351,1370]" captionTargetBox="[112,1090,219,1557]" captionTargetId="figure-12@9.[112,1086,219,1553]" captionTargetPageId="9" captionText="Figs 612. Physopelta (Physopelta) robusta Stål, 1863. 67 pygophore (6 dorsal view, magnification 50×; 7 lateral view, 47×). 812 para- mere (different orientations, 150×). Scale bars: 0.2 mm (Figs 812), 0.5 mm (Figs 67). (SEM microgra- phs: P. Kment)." figureDoi="http://doi.org/10.5281/zenodo.5739965" httpUri="https://zenodo.org/record/5739965/files/figure.png" pageId="5" pageNumber="510">Figs 812</figureCitation>
,
<figureCitation id="1324C481FFB30460FF78FD4BFEB7FD78" box="[241,311,653,677]" captionStart="Figs 3034" captionStartId="17.[113,150,1205,1224]" captionTargetBox="[113,1087,219,1185]" captionTargetId="figure-159@17.[113,1087,218,1185]" captionTargetPageId="17" captionText="Figs 3034. Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), male, paramere (different orientations, magnification 190×). Scale bars: 0.2 mm. (SEM micrographs: P. Kment)." figureDoi="http://doi.org/10.5281/zenodo.5739979" httpUri="https://zenodo.org/record/5739979/files/figure.png" pageId="5" pageNumber="510">3034</figureCitation>
,
<figureCitation id="1324C481FFB30460FEC8FD4BFE07FD78" box="[321,391,653,677]" captionStart="Figs 5259" captionStartId="47.[113,149,1514,1533]" captionTargetBox="[113,1088,219,1433]" captionTargetId="figure-12@47.[113,1088,219,1433]" captionTargetPageId="47" captionText="Figs 5259.Physopelta (Physopeltoides) dentipes sp.nov., male.5254 pygophore:52 dorsal view (magnification 75×), 53 postero-dorsal view (75×), 54 lateral view (70×). 5559 paramere (different orientations; 230×).Scale bars: 0.2 mm. (SEM micrographs: P. Kment)." figureDoi="http://doi.org/10.5281/zenodo.5739995" httpUri="https://zenodo.org/record/5739995/files/figure.png" pageId="5" pageNumber="510">5559</figureCitation>
,
<figureCitation id="1324C481FFB30460FE18FD4BFE58FD78" box="[401,472,653,677]" captionStart="Figs 7579" captionStartId="58.[151,188,594,613]" captionTargetBox="[151,1126,220,578]" captionTargetId="figure-367@58.[151,1126,219,578]" captionTargetPageId="58" captionText="Figs 7579. Physopelta (Afrophysopelta) melanoptera Distant, 1904, male, paramere (different orientations; 230×). Scale bar: 0.2 mm. (SEM micrographs: P. Kment)." figureDoi="http://doi.org/10.5281/zenodo.5740007" httpUri="https://zenodo.org/record/5740007/files/figure.png" pageId="5" pageNumber="510">7579</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3058B8FFFB30460FF19FD6BFB85FC19" pageId="5" pageNumber="510" type="discussion">
<paragraph id="8BA0D804FFB30460FF19FD6BFCE9FCD8" blockId="5.[113,1089,461,965]" pageId="5" pageNumber="510">
These characters were examined in all known genera of the subfamily (except male characters of
<taxonomicName id="4C1FA387FFB30460FF36FD08FEB3FD38" authorityName="Schmidt" authorityYear="1931" box="[191,307,718,741]" class="Insecta" family="Largidae" genus="Wachsiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FF36FD08FEB3FD38" box="[191,307,718,741]" italics="true" pageId="5" pageNumber="510">Wachsiella</emphasis>
</taxonomicName>
). Parameres of various genera and species and their modifications were figured e.g. by
<bibRefCitation id="EF8EA5F5FFB30460FE9BFD2BFD9BFCD8" author="ZAMAL &amp; CHOPRA" box="[274,539,749,773]" firstAuthor="ZAMAL" pageId="5" pageNumber="510" pagination="1 - 15" refId="ref69003" refString="ZAMAL S. &amp; CHOPRA N. P. 1990: Aaxonomic [sic!] studies on Indian Largidae (Hemiptera: Heteroptera). Journal of Research, Assam Agricultural University 11: 1 - 15." type="journal article" year="1990">ZAMAL &amp; CHOPRA (1990)</bibRefCitation>
and
<bibRefCitation id="EF8EA5F5FFB30460FDC7FD2BFCE5FCD8" author="STEHLIK &amp; JINDRA" box="[590,869,749,773]" firstAuthor="STEHLIK" pageId="5" pageNumber="510" pagination="611 - 648" refId="ref67976" refString="STEHLIK J. L. &amp; JINDRA Z. 2008 a: New taxa of the Largidae and Pyrrhocoridae (Hemiptera: Heteroptera) from Oriental Region. Acta Entomologica Musei Nationalis Pragae 48: 611 - 648." type="journal article" year="2008" yearSuffix="a">STEHLÍK &amp; JINDRA (2008a)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA0D804FFB30460FFF8FCCBFCD0FC98" blockId="5.[113,1089,461,965]" pageId="5" pageNumber="510">
<emphasis id="B96B0416FFB30460FFF8FCCBFEB8FCF8" bold="true" box="[113,312,781,805]" pageId="5" pageNumber="510">Genera included.</emphasis>
<taxonomicName id="4C1FA387FFB30460FECBFCC8FDEEFCF8" authority="Distant, 1903" authorityName="Distant" authorityYear="1903" box="[322,622,781,805]" class="Insecta" family="Largidae" genus="Delacampius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FECBFCC8FE4FFCF8" box="[322,463,782,805]" italics="true" pageId="5" pageNumber="510">Delacampius</emphasis>
<bibRefCitation id="EF8EA5F5FFB30460FE51FCCBFDEEFCF8" author="DISTANT W. L." bookContentInfo="xvii + 503 pp." box="[472,622,781,805]" editor="BLANFORD W. T." pageId="5" pageNumber="510" pagination="243 - 503" refId="ref63619" refString="DISTANT W. L. 1903 a: Rhynchota. - Vol. II. (Heteroptera). In: BLANFORD W. T. (ed.): The fauna of British India including Ceylon and Burma. Taylor and Francis, London, xvii + 503 pp. [Published in two parts: pp. 1 - 242 (December 1903, incl. Pyrrhocoroidea), pp. 243 - 503 (April 1904) - see the Preface, p. iii]." title="Rhynchota. - Vol. II. (Heteroptera)" type="book chapter" volumeTitle="The fauna of British India including Ceylon and Burma. Taylor and Francis, London" year="1903">Distant, 1903</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FA387FFB30460FDF2FCC8FCB7FCF8" authority="Stal, 1870" authorityName="Stal" authorityYear="1870" box="[635,823,780,805]" class="Insecta" family="Largidae" genus="Iphita" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FDF2FCC8FD3BFCF8" box="[635,699,782,805]" italics="true" pageId="5" pageNumber="510">Iphita</emphasis>
<bibRefCitation id="EF8EA5F5FFB30460FD4CFCCAFCB7FCF8" author="Stal" box="[709,823,780,805]" firstAuthor="Stal" pageId="5" pageNumber="510" pagination="90 - 124" refId="ref67407" refString="STAL C. 1870: Enumeratio Hemipterorum I. Kongliga Svenska Vetenskap-Academiens Handlingar 9 (1): 90 - 124." type="journal article" year="1870">Stål, 1870</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FA387FFB30460FCCDFCC8FBBDFCF8" authority="Stehlik, 2006" authorityName="Stehlik" authorityYear="2006" box="[836,1085,781,805]" class="Insecta" family="Largidae" genus="Jindraia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FCCDFCC8FC1EFCF8" box="[836,926,782,805]" italics="true" pageId="5" pageNumber="510">Jindraia</emphasis>
<bibRefCitation id="EF8EA5F5FFB30460FC20FCCBFBBDFCF8" author="STEHLIK J. L." bookContentInfo="1 - 1184" box="[937,1085,781,805]" editor="RABITSCH W." journalOrPublisher="Denisia" pageId="5" pageNumber="510" pagination="653 - 680" part="19" refId="ref67644" refString="STEHLIK J. L. 2006 a: New taxa of Pyrrhocoroidea (Heteroptera) from the Oriental Region in the Natural History Museum in London. Pp. 653 - 680. In: RABITSCH W. (ed.): Hug the bug - For love of true bugs. Festschrift zum 70. Geburtstag von Ernst Heiss. Denisia 19: 1 - 1184." title="New taxa of Pyrrhocoroidea (Heteroptera) from the Oriental Region in the Natural History Museum in London." type="book chapter" volumeTitle="Hug the bug - For love of true bugs. Festschrift zum 70. Geburtstag von Ernst Heiss" year="2006">Stehlík, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FA387FFB30460FFF8FCE8FE71FC98" authority="Amyot &amp; Serville, 1843" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[113,497,813,837]" class="Insecta" family="Largidae" genus="Physopelta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FFF8FCE8FF68FC98" box="[113,232,814,837]" italics="true" pageId="5" pageNumber="510">Physopelta</emphasis>
<bibRefCitation id="EF8EA5F5FFB30460FF64FCEBFE71FC98" author="AMYOT C. J. B. &amp; SERVILLE J. G. A." bookContentInfo="xxvi + 675 + 6 pp." box="[237,497,813,837]" journalOrPublisher="Roret, Paris" pageId="5" pageNumber="510" refId="ref62473" refString="AMYOT C. J. B. &amp; SERVILLE J. G. A. 1843: Histoire naturelle des Insectes. Hemipteres. Roret, Paris, xxvi + 675 + 6 pp." title="Histoire naturelle des Insectes. Hemipteres" type="book" year="1843">Amyot &amp; Serville, 1843</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C1FA387FFB30460FDA0FCE8FCCDFC98" authority="Schmidt, 1932" authorityName="Schmidt" authorityYear="1932" box="[553,845,813,837]" class="Insecta" family="Largidae" genus="Taeuberella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FDA0FCE8FD26FC98" box="[553,678,814,837]" italics="true" pageId="5" pageNumber="510">Taeuberella</emphasis>
<bibRefCitation id="EF8EA5F5FFB30460FD24FCEBFCCDFC98" author="Schmidt" box="[685,845,813,837]" firstAuthor="Schmidt" pageId="5" pageNumber="510" pagination="236 - 281" refId="ref66829" refString="SCHMIDT E. 1932: Zur Kenntnis der Familie Pyrrhocoridae Fieber. (Hemiptera - Heteroptera). Teil II. Wiener Entomologische Zeitung 49: 236 - 281." type="journal article" year="1932">Schmidt, 1932</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA0D804FFB30460FF19FC8BFB85FC19" blockId="5.[113,1089,461,965]" pageId="5" pageNumber="510">
The systematic position of the genus
<taxonomicName id="4C1FA387FFB30460FDA8FC88FD15FCB8" authorityName="Schmidt" authorityYear="1931" box="[545,661,846,869]" class="Insecta" family="Largidae" genus="Wachsiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B96B0416FFB30460FDA8FC88FD15FCB8" box="[545,661,846,869]" italics="true" pageId="5" pageNumber="510">Wachsiella</emphasis>
</taxonomicName>
is doubtful. The profemur of this taxon lacks any longitudinal furrow ventrally, which is a diagnostic character of the tribe
<taxonomicName id="4C1FA387FFB30460FC5FFCABFF18FC78" authorityName="Hussey" authorityYear="1929" pageId="5" pageNumber="510" rank="tribe" tribe="Physopeltini">Physopeltini</taxonomicName>
. As the genus is known only by the female sex and male characters cannot be considered to elucidate its relationships, I classify the genus here as
<taxonomicName id="4C1FA387FFB30460FD47FC6BFCE4FC18" authorityName="Hussey" authorityYear="1929" box="[718,868,941,965]" class="Insecta" family="Largidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="subFamily" subFamily="Physopeltinae">Physopeltinae</taxonomicName>
<emphasis id="B96B0416FFB30460FCE2FC68FC7EFC18" box="[875,1022,942,965]" italics="true" pageId="5" pageNumber="510">incertae sedis</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>