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<document id="8EC4348DDBE7C16B8AF95F415946F62A" ID-DOI="10.5281/zenodo.198744" ID-GBIF-Dataset="e544272c-1895-485f-a9b0-29d623bd1034" ID-ISSN="1175-5326" ID-Zenodo-Dep="198744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460313028088" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2010" docId="BB5587C4A43CA626FF7A54415A3AF90E" docLanguage="en" docName="zt02649p051.pdf" docOrigin="Zootaxa 2649" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crotonia weiri Colloff, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="476CFFBCA424A63CFFED576C593BFFF6" masterDocTitle="The Gondwanan relict oribatid genus Crotonia (Acari: Oribatida: Crotoniidae) from rainforests in Queensland and Northern New South Wales: new species show a mixed pattern of short-range and long-range endemism" masterLastPageNumber="51" masterPageNumber="1" pageNumber="25" updateTime="1698254507029" updateUser="plazi">
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<mods:title id="5254CE9A04C62C58836F085446C176D5">The Gondwanan relict oribatid genus Crotonia (Acari: Oribatida: Crotoniidae) from rainforests in Queensland and Northern New South Wales: new species show a mixed pattern of short-range and long-range endemism</mods:title>
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<mods:namePart id="1FF40849277FAA7AD9F3B586A85F21A1">Colloff, Matthew J.</mods:namePart>
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<mods:date id="2E886F769C4D63782417E75ADB008A02">2010</mods:date>
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<treatment id="BB5587C4A43CA626FF7A54415A3AF90E" ID-DOI="http://doi.org/10.5281/zenodo.6204642" ID-GBIF-Taxon="119401095" ID-Zenodo-Dep="6204642" LSID="urn:lsid:plazi:treatment:BB5587C4A43CA626FF7A54415A3AF90E" httpUri="http://treatment.plazi.org/id/BB5587C4A43CA626FF7A54415A3AF90E" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
<subSubSection id="7BE66559A43CA624FF7A54415C33FC09" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="334336D2A43CA624FF7A54415890FCB0" blockId="24.[151,427,812,873]" box="[151,427,812,839]" pageId="24" pageNumber="25">
<heading id="680B81BEA43CA624FF7A54415890FCB0" bold="true" box="[151,427,812,839]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="0188EAC0A43CA624FF7A54415890FCB0" bold="true" box="[151,427,812,839]" pageId="24" pageNumber="25">
<taxonomicName id="F4FC4D51A43CA624FF7A5441587DFCB1" ID-CoL="ZRJ6" box="[151,326,813,839]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="weiri" status="sp. nov.">
<emphasis id="0188EAC0A43CA624FF7A5441587DFCB1" bold="true" box="[151,326,813,839]" italics="true" pageId="24" pageNumber="25">Crotonia weiri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ABB57BBA43CA624FEA054405890FCB0" box="[333,427,812,838]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="334336D2A43CA624FF7A54235861FC9F" blockId="24.[151,427,812,873]" box="[151,346,847,873]" pageId="24" pageNumber="25">
(
<figureCitation id="ABC72A57A43CA624FF72542359C8FC9F" box="[159,243,847,873]" captionStart="FIGURE 9" captionStartId="25.[151,255,2008,2032]" captionTargetBox="[199,1384,436,1985]" captionTargetId="figure@25.[198,1387,427,1994]" captionTargetPageId="25" captionText="FIGURE 9. Crotonia weiri sp. nov. holotype female a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/198753/files/figure.png" pageId="24" pageNumber="25">
Figs.
<date id="47421012A43CA624FF0F542359C8FC9F" box="[226,243,847,873]" pageId="24" pageNumber="25" value="1918-01-09">9</date>
</figureCitation>
,
<figureCitation id="ABC72A57A43CA624FF1254235825FC9F" box="[255,286,847,873]" captionStart="FIGURE 18" captionStartId="43.[151,257,1884,1908]" captionTargetBox="[230,1364,182,1858]" captionTargetId="figure@43.[223,1374,170,1865]" captionTargetPageId="43" captionText="FIGURE 18. Details of bothridia of a) Crotonia sterigma sp. nov.; b) Crotonia borbora Luxton, 1987; c) Crotonia brisbanensis sp. nov.; d) Crotonia maculata sp. nov.; e) Crotonia capistrata Luxton, 1987; f) Crotonia ardala Luxton, 1987; g) Crotonia monteithi sp. nov.; h) Crotonia daviesae sp. nov.; i) Crotonia weiri sp. nov.; j) Crotonia yeatesi sp. nov.; k) Crotonia cameroni sp. nov.; l) Crotonia eungella sp. nov. m) Crotonia queenslandiae sp. nov.; n) Crotonia seemani sp. nov.; o) Crotonia raveni sp. nov." httpUri="https://zenodo.org/record/198761/files/figure.png" pageId="24" pageNumber="25">18</figureCitation>
i, 21)
</paragraph>
<paragraph id="334336D2A43CA624FF7A54F65C33FC09" blockId="24.[151,1437,922,2018]" pageId="24" pageNumber="25">
<emphasis id="0188EAC0A43CA624FF7A54F6580AFC42" bold="true" box="[151,305,922,948]" pageId="24" pageNumber="25">Dimensions:</emphasis>
<typeStatus id="EC478870A43CA624FED554F658A4FC42" box="[312,415,922,948]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
female length 1137, breadth 561.
<typeStatus id="EC478870A43CA624FCC654F65A99FC42" box="[811,930,922,948]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
: females (
<emphasis id="0188EAC0A43CA624FBFA54F65D1DFC45" box="[1047,1062,922,947]" italics="true" pageId="24" pageNumber="25">n</emphasis>
= 14): mean length 1213 (range
<date id="47421012A43CA624FF7A54D35824FC2F" box="[151,287,959,985]" pageId="24" pageNumber="25" value="1074" valueMax="1372">10741372</date>
); mean breadth 621 (range 506718); males (
<emphasis id="0188EAC0A43CA624FCDE54AC5A79FC2F" box="[819,834,960,985]" italics="true" pageId="24" pageNumber="25">n</emphasis>
= 16): mean length 1076 (range
<date id="47421012A43CA624FB2C54D35C72FC2F" box="[1217,1353,959,985]" pageId="24" pageNumber="25" value="1003" valueMax="1130">10031130</date>
); mean breadth 532 (range 490587). Ratio of length of prodorsum to total body length: 0.35 (
<typeStatus id="EC478870A43CA624FB6354895DC7FC09" box="[1166,1276,997,1023]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="7BE66559A43CA626FF2853665A2AFCEF" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
<paragraph id="334336D2A43CA624FF2853665AB6FA49" blockId="24.[151,1437,922,2018]" pageId="24" pageNumber="25">
<emphasis id="0188EAC0A43CA624FF28536658D2FBD2" bold="true" box="[197,489,1034,1060]" pageId="24" pageNumber="25">Description of female.</emphasis>
<emphasis id="0188EAC0A43CA624FE1F53665B47FBD5" box="[498,636,1034,1059]" italics="true" pageId="24" pageNumber="25">Prodorsum</emphasis>
: ratio of length to breadth 1.0. Rostrum with squat naso, barely projecting beyond rostrum; lateral edges slightly concave; rostral seta 43, straight, spiniform, smooth (
<figureCitation id="ABC72A57A43CA624FA865343599CFB99" captionStart="FIGURE 9" captionStartId="25.[151,255,2008,2032]" captionTargetBox="[199,1384,436,1985]" captionTargetId="figure@25.[198,1387,427,1994]" captionTargetPageId="25" captionText="FIGURE 9. Crotonia weiri sp. nov. holotype female a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/198753/files/figure.png" pageId="24" pageNumber="25">Fig. 9</figureCitation>
a). Lamellar seta 243, curved, smooth, flagelliform apically. Lamellar apophyses 108, straight, thin, slightly diverging apically, shorter than their mutual distance by half; extending anteriorly beyond apices of rostral setae. Interlamellar apophyses twice as long as broad; interlamellar seta 335, flagelliform, smooth; extending anteriorly as far as apex of curve of lamellar setae. Prodorsal ridges well-developed, curved medially, extending anteriorly almost a third of the distance between bases of interlamellar and lamellar apophyses. Diameter of bothridium 42; auriculate ridge of bothridium in the form of a complex series of digitate or triangular spines, with a large, blunt, anterior projection; hexagonal reticulations of operculum lacking concentric ridges and with point of origin near lateral margin (
<figureCitation id="ABC72A57A43CA624FC8252365AFEFA82" box="[879,965,1370,1396]" captionStart="FIGURE 18" captionStartId="43.[151,257,1884,1908]" captionTargetBox="[230,1364,182,1858]" captionTargetId="figure@43.[223,1374,170,1865]" captionTargetPageId="43" captionText="FIGURE 18. Details of bothridia of a) Crotonia sterigma sp. nov.; b) Crotonia borbora Luxton, 1987; c) Crotonia brisbanensis sp. nov.; d) Crotonia maculata sp. nov.; e) Crotonia capistrata Luxton, 1987; f) Crotonia ardala Luxton, 1987; g) Crotonia monteithi sp. nov.; h) Crotonia daviesae sp. nov.; i) Crotonia weiri sp. nov.; j) Crotonia yeatesi sp. nov.; k) Crotonia cameroni sp. nov.; l) Crotonia eungella sp. nov. m) Crotonia queenslandiae sp. nov.; n) Crotonia seemani sp. nov.; o) Crotonia raveni sp. nov." httpUri="https://zenodo.org/record/198761/files/figure.png" pageId="24" pageNumber="25">
Fig.
<date id="47421012A43CA624FC4852365AFEFA82" box="[933,965,1370,1396]" pageId="24" pageNumber="25" value="1918-01">18</date>
</figureCitation>
i). Inter-bothridial ridge well-developed, straight, transverse, curved anteriolaterally at margins, with a slight median invagination. Median field of muscle sigilla present. Prodorsal microsculpture smooth, porose.
</paragraph>
<paragraph id="334336D2A43CA625FF2852A658D1FE8D" blockId="24.[151,1437,922,2018]" lastBlockId="25.[151,1437,152,379]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="0188EAC0A43CA624FF2852A6587CFA15" box="[197,327,1482,1507]" italics="true" pageId="24" pageNumber="25">Notogaster</emphasis>
: ratio of length of notogaster to breadth of notogastral shield 1.31; notogastral shield broadest between setae
<emphasis id="0188EAC0A43CA624FEA9529C585BF9FF" box="[324,352,1520,1545]" italics="true" pageId="24" pageNumber="25">cp</emphasis>
and
<emphasis id="0188EAC0A43CA624FE77529C589CF9FF" box="[410,423,1520,1545]" italics="true" pageId="24" pageNumber="25">e</emphasis>
2 (
<figureCitation id="ABC72A57A43CA624FE2D52835B33F9FF" box="[448,520,1519,1545]" captionStart="FIGURE 9" captionStartId="25.[151,255,2008,2032]" captionTargetBox="[199,1384,436,1985]" captionTargetId="figure@25.[198,1387,427,1994]" captionTargetPageId="25" captionText="FIGURE 9. Crotonia weiri sp. nov. holotype female a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/198753/files/figure.png" pageId="24" pageNumber="25">Fig. 9</figureCitation>
a); ratio of width of shield to entire notogastral width 0.82; lateral margins of shield greatly expanded, sub-circular. Dorsosejugal suture discrete, with single indented ridge. Region posterior of apophyses of setae
<emphasis id="0188EAC0A43CA624FDE351515B20F9A0" box="[526,539,1597,1622]" italics="true" pageId="24" pageNumber="25">c</emphasis>
3 slightly concave. Lyrifissure
<emphasis id="0188EAC0A43CA624FC6951515AA1F9A0" box="[900,922,1597,1622]" italics="true" pageId="24" pageNumber="25">ia</emphasis>
short (22), on shallow ridge posteriolateral of apophyses of setae
<emphasis id="0188EAC0A43CA624FE48510A5889F989" box="[421,434,1638,1663]" italics="true" pageId="24" pageNumber="25">c</emphasis>
3. With 14 pairs of smooth notogastral setae. Pre-notogastral shield separated from notogastral shield by narrow hyaline strip, and bearing well-developed cylindrical apophysis (27 long) of seta
<emphasis id="0188EAC0A43CA624FF7A51DF599FF93A" box="[151,164,1715,1740]" italics="true" pageId="24" pageNumber="25">c</emphasis>
3 and minute, subequal, setiform setae
<emphasis id="0188EAC0A43CA624FD8751DF5B4CF93A" box="[618,631,1715,1740]" italics="true" pageId="24" pageNumber="25">c</emphasis>
1 and
<emphasis id="0188EAC0A43CA624FD5751DF5BFCF93A" box="[698,711,1715,1740]" italics="true" pageId="24" pageNumber="25">c</emphasis>
2 (11)on posterior margin. Seta
<emphasis id="0188EAC0A43CA624FBD551DF5D7EF93A" box="[1080,1093,1715,1740]" italics="true" pageId="24" pageNumber="25">c</emphasis>
3 well-developed, long (390), flagelliform apically, extending anteriorly to point midway between bases of interlamellar and lamellar apophyses. Notogastral shield smooth, slightly convex, tapering to U-shape anterior of caudal apophyses; bordered laterally by two narrow strips of small tubercles extending to point between tubercles of setae
<emphasis id="0188EAC0A43CA624FAB7504A5C59F8C9" box="[1370,1378,1830,1855]" italics="true" pageId="24" pageNumber="25">f</emphasis>
2 and
<emphasis id="0188EAC0A43CA624FF7A502259A4F891" box="[151,159,1870,1895]" italics="true" pageId="24" pageNumber="25">f</emphasis>
1. Lateral hyaline strip (suprapleural scissure) wide, bearing tubercles of short, setiform, setae
<emphasis id="0188EAC0A43CA624FB1A50225C29F891" box="[1271,1298,1870,1895]" italics="true" pageId="24" pageNumber="25">cp</emphasis>
(94) and
<emphasis id="0188EAC0A43CA624FA6A50225CAFF891" box="[1415,1428,1870,1895]" italics="true" pageId="24" pageNumber="25">e</emphasis>
2 (70). Seta
<emphasis id="0188EAC0A43CA624FEFF501B581AF866" box="[274,289,1911,1936]" italics="true" pageId="24" pageNumber="25">d</emphasis>
2 thin, short (56), setiform; mutual distance (81) a third greater than that between setae
<emphasis id="0188EAC0A43CA624FAC1501B5C02F866" box="[1324,1337,1911,1936]" italics="true" pageId="24" pageNumber="25">c</emphasis>
2. Seta
<emphasis id="0188EAC0A43CA624FA61501B5CAFF866" box="[1420,1428,1911,1936]" italics="true" pageId="24" pageNumber="25">f</emphasis>
2 short (89), recurved, each on squat tubercle barely projecting from margin of notogastral shield. Opisthosomal gland opening
<emphasis id="0188EAC0A43CA624FEAB50A85850F82B" box="[326,363,1988,2013]" italics="true" pageId="24" pageNumber="25">gla</emphasis>
positioned anteriomedial of setae
<emphasis id="0188EAC0A43CA624FCEE50A85A30F82B" box="[771,779,1988,2013]" italics="true" pageId="24" pageNumber="25">f</emphasis>
2. Caudal apophyses short, clustered, projecting directly from posterior notogastral shield, not on stalk; those of
<emphasis id="0188EAC0A43DA625FCD357F45A76FF47" box="[830,845,152,177]" italics="true" pageId="25" pageNumber="26">h</emphasis>
2 short (44), more-or-less parallel, caudal margin between them transverse. Apophysis of
<emphasis id="0188EAC0A43DA625FD8757AC5B42FF2F" box="[618,633,192,217]" italics="true" pageId="25" pageNumber="26">h</emphasis>
1 (30) emerging dorsolaterally from base of that of
<emphasis id="0188EAC0A43DA625FB2757AC5DE2FF2F" box="[1226,1241,192,217]" italics="true" pageId="25" pageNumber="26">h</emphasis>
2; apophysis of
<emphasis id="0188EAC0A43DA625FA6157AC5CAFFF2F" box="[1420,1428,192,217]" italics="true" pageId="25" pageNumber="26">f</emphasis>
1 (16) projecting posteriolaterally; the base not closely associated with that of
<emphasis id="0188EAC0A43DA625FB8057855D47FEF4" box="[1133,1148,233,258]" italics="true" pageId="25" pageNumber="26">h</emphasis>
1. Apophysis of seta
<emphasis id="0188EAC0A43DA625FA6957855CA8FEF4" box="[1412,1427,233,258]" italics="true" pageId="25" pageNumber="26">h</emphasis>
3 positioned between
<emphasis id="0188EAC0A43DA625FE6E567D58A9FEDC" box="[387,402,273,298]" italics="true" pageId="25" pageNumber="26">h</emphasis>
1 and
<emphasis id="0188EAC0A43DA625FE39567D58E7FEDC" box="[468,476,273,298]" italics="true" pageId="25" pageNumber="26">f</emphasis>
1 ventrally; seta
<emphasis id="0188EAC0A43DA625FD77567D5B92FEDC" box="[666,681,273,298]" italics="true" pageId="25" pageNumber="26">h</emphasis>
3 thin, setiform, curved. Other caudal setae short, thick, straight, semi-spiniform;
<emphasis id="0188EAC0A43DA625FE8C56555852FEA4" box="[353,361,313,338]" italics="true" pageId="25" pageNumber="26">f</emphasis>
1 46,
<emphasis id="0188EAC0A43DA625FE4456555883FEA4" box="[425,440,313,338]" italics="true" pageId="25" pageNumber="26">h</emphasis>
1 67,
<emphasis id="0188EAC0A43DA625FE1556555B3CFEA4" box="[504,519,313,338]" italics="true" pageId="25" pageNumber="26">h</emphasis>
2 71. Caudal region of dorsal notogastral plate with small, sparse tubercles, extending onto lateral plates.
</paragraph>
<caption id="6783665AA43DA625FF7A50B45D2CF819" box="[151,1047,2008,2032]" httpUri="https://zenodo.org/record/198753/files/figure.png" pageId="25" pageNumber="26" targetBox="[199,1384,436,1985]" targetPageId="25">
<paragraph id="334336D2A43DA625FF7A50B45D2CF819" blockId="25.[151,1047,2008,2032]" box="[151,1047,2008,2032]" pageId="25" pageNumber="26">
<emphasis id="0188EAC0A43DA625FF7A50B45822F806" bold="true" box="[151,281,2008,2032]" pageId="25" pageNumber="26">FIGURE 9.</emphasis>
<taxonomicName id="F4FC4D51A43DA625FECD50B45886F819" box="[288,445,2008,2031]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="weiri" status="sp. nov.">
<emphasis id="0188EAC0A43DA625FECD50B45886F819" box="[288,445,2008,2031]" italics="true" pageId="25" pageNumber="26">Crotonia weiri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ABB57BBA43DA625FE2E50B45B23F819" box="[451,536,2008,2032]" pageId="25" pageNumber="26" rank="species">
<emphasis id="0188EAC0A43DA625FE2E50B45B29F806" bold="true" box="[451,530,2008,2032]" pageId="25" pageNumber="26">sp. nov</emphasis>
.
</taxonomicNameLabel>
holotype female a) dorsal; b) ventral; c) lateral.
</paragraph>
</caption>
<paragraph id="334336D2A43EA626FF2857F45BC4FDC6" blockId="26.[151,1438,152,1788]" pageId="26" pageNumber="27">
<emphasis id="0188EAC0A43EA626FF2857F45835FF47" box="[197,270,152,177]" italics="true" pageId="26" pageNumber="27">Ven ter</emphasis>
: epimeral microsculpture smooth, porose (
<figureCitation id="ABC72A57A43EA626FCFB57F45A64FF44" box="[790,863,152,178]" captionStart="FIGURE 9" captionStartId="25.[151,255,2008,2032]" captionTargetBox="[199,1384,436,1985]" captionTargetId="figure@25.[198,1387,427,1994]" captionTargetPageId="25" captionText="FIGURE 9. Crotonia weiri sp. nov. holotype female a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/198753/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
b); epimeral setae smooth, thin, spiniform, very short, ca. 815, formula 3-1-3-3; seta
<emphasis id="0188EAC0A43EA626FD8157D15BB1FF20" box="[620,650,189,214]" italics="true" pageId="26" pageNumber="27">3c</emphasis>
longer and stouter than others (24) on well-developed tubercle. Epimeral plates I with sharp, horn-like projections posteriolaterally. With
<emphasis id="0188EAC0A43EA626FBEF578E5D18FF0D" box="[1026,1059,226,251]" italics="true" pageId="26" pageNumber="27">lcs</emphasis>
broad, curved, forming a broad obtuse angle (ca. 140°) with
<emphasis id="0188EAC0A43EA626FE1556645B1BFED7" box="[504,544,264,289]" italics="true" pageId="26" pageNumber="27">pcs</emphasis>
. Anteriolateral margin of adanal plate with a broad, shallow indentation. Genital plates sub-circular; posterior margin transverse. Perigenital region strongly and densely tuberculate. Each genital plate 198 long, 118 broad, with eight setiform setae, subequal in length (30); two pairs of aggenital setae, subequal in length to epimeral setae. Anal plate narrow (41 broad), 251 long, with three very short (10) setae on posterior region of plate; three pairs of spiniform adanal setae,
<emphasis id="0188EAC0A43EA626FB9C56F15DB4FE40" box="[1137,1167,413,438]" italics="true" pageId="26" pageNumber="27">ad</emphasis>
2-3 slightly shorter than spiniform
<emphasis id="0188EAC0A43EA626FEFD56AA5816FE29" box="[272,301,454,479]" italics="true" pageId="26" pageNumber="27">ad</emphasis>
1 (29); distance between
<emphasis id="0188EAC0A43EA626FDA256AA5B57FE29" box="[591,620,454,479]" italics="true" pageId="26" pageNumber="27">ad</emphasis>
series subequal. Ventral margin of notogaster surrounding anal plates curved, U-shaped. Setae of
<emphasis id="0188EAC0A43EA626FE0A568258CDFDF1" box="[487,502,494,519]" italics="true" pageId="26" pageNumber="27">p</emphasis>
series curved, setiform,
<emphasis id="0188EAC0A43EA626FCCF56825A0AFDF1" box="[802,817,494,519]" italics="true" pageId="26" pageNumber="27">p</emphasis>
1 very short, 15,
<emphasis id="0188EAC0A43EA626FC1756825D32FDF1" box="[1018,1033,494,519]" italics="true" pageId="26" pageNumber="27">p</emphasis>
2 30,
<emphasis id="0188EAC0A43EA626FBA556825D6CFDF1" box="[1096,1111,494,519]" italics="true" pageId="26" pageNumber="27">p</emphasis>
3 43. Tubercles of
<emphasis id="0188EAC0A43EA626FAD856825C7FFDF1" box="[1333,1348,494,519]" italics="true" pageId="26" pageNumber="27">p</emphasis>
1 squat, separated by distance at least four times their length.
</paragraph>
<paragraph id="334336D2A43EA626FF2855575A2AFCEF" blockId="26.[151,1438,152,1788]" pageId="26" pageNumber="27">
<emphasis id="0188EAC0A43EA626FF2855575860FDA2" box="[197,347,571,596]" italics="true" pageId="26" pageNumber="27">Lateral view</emphasis>
: Caudal margin inflated, rounded, contiguous with notogastral shield (
<figureCitation id="ABC72A57A43EA626FB5D55575DC2FDA3" box="[1200,1273,571,597]" captionStart="FIGURE 9" captionStartId="25.[151,255,2008,2032]" captionTargetBox="[199,1384,436,1985]" captionTargetId="figure@25.[198,1387,427,1994]" captionTargetPageId="25" captionText="FIGURE 9. Crotonia weiri sp. nov. holotype female a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/198753/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
c); dorsal and ventral surfaces convex, not parallel, distance between them ca. 660 perpendicular of seta
<emphasis id="0188EAC0A43EA626FB15550D5C3EFD8C" box="[1272,1285,609,634]" italics="true" pageId="26" pageNumber="27">e</emphasis>
2, narrowing anteriorly. Apophysis of seta
<emphasis id="0188EAC0A43EA626FE0155E558CFFD54" box="[492,500,649,674]" italics="true" pageId="26" pageNumber="27">f</emphasis>
1 pointing dorsally, those of
<emphasis id="0188EAC0A43EA626FCD755E55A73FD54" box="[826,840,649,674]" italics="true" pageId="26" pageNumber="27">h</emphasis>
1 pointing posteriodorsally, those of
<emphasis id="0188EAC0A43EA626FB0155E55DC1FD54" box="[1260,1274,649,674]" italics="true" pageId="26" pageNumber="27">h</emphasis>
2 only slightly longer than those of
<emphasis id="0188EAC0A43EA626FE4755DD5882FD3C" box="[426,441,689,714]" italics="true" pageId="26" pageNumber="27">h</emphasis>
1, positioned ventral of them; those of
<emphasis id="0188EAC0A43EA626FC5755DD5AF2FD3C" box="[954,969,689,714]" italics="true" pageId="26" pageNumber="27">h</emphasis>
3 positioned ventral of
<emphasis id="0188EAC0A43EA626FB1755DD5C32FD3C" box="[1274,1289,689,714]" italics="true" pageId="26" pageNumber="27">h</emphasis>
2, pointing ventrolaterally. Caudal cluster positioned below level of notogastral shield. Apophyses of setae of
<emphasis id="0188EAC0A43EA626FAAD55B55C74FD04" box="[1344,1359,729,754]" italics="true" pageId="26" pageNumber="27">p</emphasis>
series equidistantly spaced. Pleuraspis with sparse tubercles.
</paragraph>
</subSubSection>
<subSubSection id="7BE66559A43EA626FF28544F5A7DFBB5" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="334336D2A43EA626FF28544F5A7DFBB5" blockId="26.[151,1438,152,1788]" pageId="26" pageNumber="27">
<emphasis id="0188EAC0A43EA626FF28544F58FAFCCB" bold="true" box="[197,449,803,829]" pageId="26" pageNumber="27">Material examined.</emphasis>
<typeStatus id="EC478870A43EA626FE2754485B01FCC8" box="[458,570,804,830]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
female, litter and moss under
<taxonomicName id="F4FC4D51A43EA626FC4254485827FC95" authority="Mount Bithongabel, Lamington" authorityName="Mount Bithongabel, Lamington" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="moorei">
<emphasis id="0188EAC0A43EA626FC4254485DACFCCB" box="[943,1175,804,829]" italics="true" pageId="26" pageNumber="27">Nothofagus moorei</emphasis>
, Mount Bithongabel, Lamington
</taxonomicName>
National Park, Queensland,
<collectionCode id="55EDAE17A43EA626FD9054255BFCFC95" LSID="urn:lsid:biocol.org:col:32981" box="[637,711,841,867]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="27">ANIC</collectionCode>
654,
<geoCoordinate id="56C85015A43EA626FCE354255AA0FC95" box="[782,923,841,867]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="-28.262777">28°15'46&quot;S</geoCoordinate>
,
<geoCoordinate id="56C85015A43EA626FC4554255D7DFC95" box="[936,1094,841,867]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="153.17139">153°10'17&quot;E</geoCoordinate>
,
<quantity id="F4049B37A43EA626FBBE54255D8FFC95" box="[1107,1204,841,867]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="26" pageNumber="27" unit="m" value="1190.0">1190 m</quantity>
., coll. T. Weir &amp; J. Lawrence,
<date id="47421012A43EA626FEF1540358ACFC7F" box="[284,407,879,905]" pageId="26" pageNumber="27" value="1978-10-23">23.x.1978</date>
.
<typeStatus id="EC478870A43EA626FE4E54035B30FC7F" box="[419,523,879,905]" pageId="26" pageNumber="27" type="paratype">Paratype</typeStatus>
female, QM
<accessionNumber id="2CAFAB31A43EA626FD4154035A31FC7F" box="[684,778,879,905]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S26049" pageId="26" pageNumber="27" type="EnaNcbi">S26049</accessionNumber>
, brushings from sticks, rainforest, Mount Bithongabel, Lamington National Park, Queensland,
<geoCoordinate id="56C85015A43EA626FD6A54F85BD3FC58" box="[647,744,916,942]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="-28.266666">28°16'S</geoCoordinate>
<geoCoordinate id="56C85015A43EA626FD1C54F85A5DFC58" box="[753,870,916,942]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="153.16667">153°10'E</geoCoordinate>
, coll. G.B. Monteith,
<date id="47421012A43EA626FB9B54F85DDDFC58" box="[1142,1254,916,942]" pageId="26" pageNumber="27" value="1979-10-08">8.x.1979</date>
.
<typeStatus id="EC478870A43EA626FB1954F85C49FC58" box="[1268,1394,916,942]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="25FAFD5BA43EA626FA9354F859C0FC25" pageId="26" pageNumber="27" type="female">13 females</specimenCount>
,
<specimenCount id="25FAFD5BA43EA626FEE554D55841FC25" box="[264,378,953,979]" pageId="26" pageNumber="27" type="male">16 males</specimenCount>
, QM
<accessionNumber id="2CAFAB31A43EA626FE2F54D55B18FC25" box="[450,547,953,979]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S26069" pageId="26" pageNumber="27" type="EnaNcbi">S26069</accessionNumber>
, litter, Mount Hobwee, Lamington National Park, Queensland,
<geoCoordinate id="56C85015A43EA626FAD154D55CA6FC25" box="[1340,1437,953,979]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="-28.2">28°12'S</geoCoordinate>
<geoCoordinate id="56C85015A43EA626FF7A54B35832FC0F" box="[151,265,991,1017]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="153.16667">153°10'E</geoCoordinate>
.
<typeStatus id="EC478870A43EA626FEF854B35845FC0F" box="[277,382,991,1017]" pageId="26" pageNumber="27" type="paratype">Paratype</typeStatus>
female, QM
<accessionNumber id="2CAFAB31A43EA626FDF254B35B46FC0F" box="[543,637,991,1017]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S26073" pageId="26" pageNumber="27" type="EnaNcbi">S26073</accessionNumber>
, pitfall trap, simple notophyll vine forest, Nagarigoon, Lamington National Park, Queensland,
<geoCoordinate id="56C85015A43EA626FE3253685B01FBE8" box="[479,570,1028,1054]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="-28.2">28°12'S</geoCoordinate>
<geoCoordinate id="56C85015A43EA626FDAC53685B8BFBE8" box="[577,688,1028,1054]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="153.16667">153°10'E</geoCoordinate>
, coll. R.J. Raven &amp; V.T. Davies,
<date id="47421012A43EA626FBDD53685DC9FBE8" box="[1072,1266,1028,1054]" pageId="26" pageNumber="27" value="1976-03-30" valueMax="1976-04-04" valueMin="1976-03-30">30.iii.4.iv.1976</date>
.
<typeStatus id="EC478870A43EA626FB1353685C50FBE8" box="[1278,1387,1028,1054]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
and
<typeStatus id="EC478870A43EA626FF7A5345583CFBB5" box="[151,263,1065,1091]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
deposited in the Queensland Museum, Brisbane.
</paragraph>
</subSubSection>
<subSubSection id="7BE66559A43EA626FF2853225C05FB78" pageId="26" pageNumber="27" type="etymology">
<paragraph id="334336D2A43EA626FF2853225C05FB78" blockId="26.[151,1438,152,1788]" pageId="26" pageNumber="27">
<emphasis id="0188EAC0A43EA626FF2853225863FB9E" bold="true" box="[197,344,1102,1128]" pageId="26" pageNumber="27">Etymology.</emphasis>
This species is named in honour of my colleague Tom Weir (Australian National Insect Collection, CSIRO Ecosystem Sciences), in recognition of his contribution to Australian entomology.
</paragraph>
</subSubSection>
<subSubSection id="7BE66559A43EA626FF2853F55A3AF90E" pageId="26" pageNumber="27" type="discussion">
<paragraph id="334336D2A43EA626FF2853F558F8FA81" blockId="26.[151,1438,152,1788]" pageId="26" pageNumber="27">
<emphasis id="0188EAC0A43EA626FF2853F55805FB45" bold="true" box="[197,318,1177,1203]" pageId="26" pageNumber="27">Remarks.</emphasis>
<taxonomicName id="F4FC4D51A43EA626FEA853F558CAFB44" box="[325,497,1177,1202]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="weiri">
<emphasis id="0188EAC0A43EA626FEA853F558CAFB44" box="[325,497,1177,1202]" italics="true" pageId="26" pageNumber="27">Crotonia weiri</emphasis>
</taxonomicName>
differs from all other
<taxonomicName id="F4FC4D51A43EA626FD1953F55A66FB44" box="[756,861,1177,1202]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="0188EAC0A43EA626FD1953F55A66FB44" box="[756,861,1177,1202]" italics="true" pageId="26" pageNumber="27">Crotonia</emphasis>
</taxonomicName>
spp. by the following combination of characters: 1) the expanded, sub-circular opisthosoma; 2) the short caudal apophyses with those of seta
<emphasis id="0188EAC0A43EA626FB0A53D35DCDFB2E" box="[1255,1270,1215,1240]" italics="true" pageId="26" pageNumber="27">h</emphasis>
2 only slightly longer than those of
<emphasis id="0188EAC0A43EA626FE63538B58A6FAF6" box="[398,413,1255,1280]" italics="true" pageId="26" pageNumber="27">h</emphasis>
1; 3) the short, smooth, spiniform caudal setae; 4) the very short epimeral, anal and
<emphasis id="0188EAC0A43EA626FA68538B5CAFFAF6" box="[1413,1428,1255,1280]" italics="true" pageId="26" pageNumber="27">p</emphasis>
1 setae; 5) the minute setae
<emphasis id="0188EAC0A43EA626FE24527C58EDFADF" box="[457,470,1296,1321]" italics="true" pageId="26" pageNumber="27">c</emphasis>
1 and
<emphasis id="0188EAC0A43EA626FDF4527C5B1DFADF" box="[537,550,1296,1321]" italics="true" pageId="26" pageNumber="27">c</emphasis>
2 positioned close together; 6) the considerable depth of the opisthosoma in lateral view; 7) the lateral ridge of the bothridium with a series of digitate and triangular spines and a large, blunt, anterior projection.
</paragraph>
<paragraph id="334336D2A43EA626FF2852EE5A3AF90E" blockId="26.[151,1438,152,1788]" pageId="26" pageNumber="27">
<taxonomicName id="F4FC4D51A43EA626FF2852EE5848FA6D" box="[197,371,1410,1435]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="weiri">
<emphasis id="0188EAC0A43EA626FF2852EE5848FA6D" box="[197,371,1410,1435]" italics="true" pageId="26" pageNumber="27">Crotonia weiri</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="F4FC4D51A43EA626FDBF52EE5BE6FA6D" box="[594,733,1410,1435]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="daviesae">
<emphasis id="0188EAC0A43EA626FDBF52EE5BE6FA6D" box="[594,733,1410,1435]" italics="true" pageId="26" pageNumber="27">C. daviesae</emphasis>
</taxonomicName>
in the morphology of the prodorsum and its structures; the inflated opisthosoma narrowing anteriorly in lateral view; the relative lengths of setae
<emphasis id="0188EAC0A43EA626FB7D52C45DA6FA37" box="[1168,1181,1448,1473]" italics="true" pageId="26" pageNumber="27">c</emphasis>
3,
<emphasis id="0188EAC0A43EA626FB5952C45DF4FA37" box="[1204,1231,1448,1473]" italics="true" pageId="26" pageNumber="27">cp</emphasis>
,
<emphasis id="0188EAC0A43EA626FB3252C45DD7FA37" box="[1247,1260,1448,1473]" italics="true" pageId="26" pageNumber="27">e</emphasis>
2 and
<emphasis id="0188EAC0A43EA626FAC352C45C0DFA37" box="[1326,1334,1448,1473]" italics="true" pageId="26" pageNumber="27">f</emphasis>
2 and the projections on epimeral plates I. It is also similar to
<taxonomicName id="F4FC4D51A43EA626FCF152BC5AADFA1F" box="[796,918,1488,1513]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="yeatesi" status="sp. nov.">
<emphasis id="0188EAC0A43EA626FCF152BC5AADFA1F" box="[796,918,1488,1513]" italics="true" pageId="26" pageNumber="27">C. yeatesi</emphasis>
</taxonomicName>
<emphasis id="0188EAC0A43EA626FC7352A35D3BFA1F" bold="true" box="[926,1024,1487,1513]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="1ABB57BBA43EA626FC7352A35D3BFA1F" box="[926,1024,1487,1513]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in these characters as well as the morphology and relative lengths of setae
<emphasis id="0188EAC0A43EA626FD4752995B82F9F8" box="[682,697,1525,1550]" italics="true" pageId="26" pageNumber="27">d</emphasis>
2 and the caudal apophyses and their setae. Other than the characters mentioned in the diagnosis,
<taxonomicName id="F4FC4D51A43EA626FD8F51725BFAF9C1" box="[610,705,1566,1591]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="weiri">
<emphasis id="0188EAC0A43EA626FD8F51725BFAF9C1" box="[610,705,1566,1591]" italics="true" pageId="26" pageNumber="27">C. weiri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4FC4D51A43EA626FCB351725ADCF9C1" box="[862,999,1566,1591]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="daviesae">
<emphasis id="0188EAC0A43EA626FCB351725ADCF9C1" box="[862,999,1566,1591]" italics="true" pageId="26" pageNumber="27">C. daviesae</emphasis>
</taxonomicName>
in having setae
<emphasis id="0188EAC0A43EA626FB4551725D8BF9C1" box="[1192,1200,1566,1591]" italics="true" pageId="26" pageNumber="27">f</emphasis>
2 on tubercles rather than on apophyses that project beyond the margin of the notogaster, in the more restricted tuberculate microsculpture, the positioning of the anal setae on the posterior half of the anal plate, in the short setae of the
<emphasis id="0188EAC0A43EA626FF7A51FD599DF95C" box="[151,166,1681,1706]" italics="true" pageId="26" pageNumber="27">p</emphasis>
series and the lateral regions of the notogastral plate bearing a narrower tuberculate strip.
<taxonomicName id="F4FC4D51A43EA626FB0151FD5CA7F95C" box="[1260,1436,1681,1706]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="weiri">
<emphasis id="0188EAC0A43EA626FB0151FD5CA7F95C" box="[1260,1436,1681,1706]" italics="true" pageId="26" pageNumber="27">Crotonia weiri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F4FC4D51A43EA626FEC651DA58A4F939" box="[299,415,1718,1743]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="yeatesi">
<emphasis id="0188EAC0A43EA626FEC651DA58A4F939" box="[299,415,1718,1743]" italics="true" pageId="26" pageNumber="27">C. yeatesi</emphasis>
</taxonomicName>
in having much longer setae
<emphasis id="0188EAC0A43EA626FD1751DA5A3CF939" box="[762,775,1718,1743]" italics="true" pageId="26" pageNumber="27">c</emphasis>
2, smooth reflexed setae
<emphasis id="0188EAC0A43EA626FBCB51DA5D15F939" box="[1062,1070,1718,1743]" italics="true" pageId="26" pageNumber="27">f</emphasis>
2 lacking projecting apophyses, short setae
<emphasis id="0188EAC0A43EA626FEF651B25811F901" box="[283,298,1758,1783]" italics="true" pageId="26" pageNumber="27">p</emphasis>
1 and a more rounded notogastral shield.
</paragraph>
</subSubSection>
</treatment>
</document>