359 lines
33 KiB
XML
359 lines
33 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.51909" ID-GBIF-Dataset="185fc6f3-e76d-42ef-911f-ad0e5cdd2315" ID-PMC="PMC7305247" ID-Pensoft-Pub="1314-2003-151-1" ID-Pensoft-UUID="73AED83DDB7D5E8395DB6FC61FA3502A" ID-PubMed="32587460" ModsDocID="1314-2003-151-1" checkinTime="1592065774662" checkinUser="pensoft" docAuthor="Wege, Juliet A." docDate="2020" docId="32CF3AD78F9054E58921BB3752C8136F" docLanguage="en" docName="PhytoKeys 151: 1-47" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.51909" docTitle="Levenhookia stipitata (Benth.) F. Muell. ex Benth., Fl. Austral. 4: 36. 1868" docType="treatment" docVersion="4" id="73AED83DDB7D5E8395DB6FC61FA3502A" lastPageNumber="1" masterDocId="73AED83DDB7D5E8395DB6FC61FA3502A" masterDocTitle="Styleworts under the microscope: a taxonomic account of Levenhookia (Stylidiaceae)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="1" updateTime="1668139842397" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Styleworts under the microscope: a taxonomic account of Levenhookia (Stylidiaceae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wege, Juliet A.</mods:namePart>
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<mods:affiliation>Western Australian Herbarium, Biodiversity and Conservation Science, Department of Biodiversity, Conservation and Attractions, 17 Dick Perry Ave Kensington, Western Australia 6151, Perth, Australia</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7312-3840</mods:nameIdentifier>
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<mods:nameIdentifier type="email">juliet.wege@dbca.wa.gov.au</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>151</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>47</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.151.51909</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.151.51909</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-151-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">73AED83DDB7D5E8395DB6FC61FA3502A</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164596955" LSID="urn:lsid:plazi:treatment:32CF3AD78F9054E58921BB3752C8136F" httpUri="http://treatment.plazi.org/id/32CF3AD78F9054E58921BB3752C8136F" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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10.
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<taxonomicName LSID="32CF3AD7-8F90-54E5-8921-BB3752C8136F" authority="(Benth.) F. Muell. ex Benth., Fl. Austral. 4: 36. 1868" authorityName="(Benth.) F. Muell. ex Benth., Fl. Austral. 4: 36." authorityYear="1868" class="Magnoliopsida" family="Stylidiaceae" genus="Levenhookia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Levenhookia stipitata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stipitata">Levenhookia stipitata (Benth.) F.Muell. ex Benth., Fl. Austral. 4: 36. 1868</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Labellum and column movement in Levenhookia. Black arrow = labellum hooded over column; white arrow = labellum ' triggered' to release column; yellow arrow = labellum repositioned following column release; blue arrow = column position immediately following release from labellum A L. stipitata (unvouchered, Augusta area, Western Australia) B L. aestiva (J. A. Wege 2090) C-E L. leptantha (J. A. Wege 2063), with labellum enclosing the column (C), triggered to release the column which moves to the opposite side of the flower (D) and subsequently repositioned (E) with the stigmatic lobes developed F L. aestiva (J. A. Wege 2090), labellum triggered with column momentum stopped by the sheath at the base of the corolla lobes G, H L. pauciflora (at J. A. Wege 1071 & C. Wilkins): note the unusual, distally-angled column and brush-tipped labellum I L. murfetii (J. A. Wege 2060): note the dorsal position of the labellum; J, K L. dubia, with the labellum opening (but not otherwise moving) to release the column (K). Photos by R. W. Davis (A, B) and J. A. Wege (C-K)." figureDoi="10.3897/phytokeys.151.51909.figure2" httpUri="https://binary.pensoft.net/fig/420774" pageId="0" pageNumber="1">Figs 2A</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Comparative distributions and floral morphologies A, B L. stipitata, with a disjunct distribution in Western Australia and South Australia and flowers on long pedicels and with a prominent column sheath (J. A. Wege 1874). Inset showing coloured markings at the base of the corolla lobes that have been recorded for many populations (J. A. Wege 1873) C, D L. octomaculata, a Western Australian endemic with prominent corolla markings and a densely papillate labellum (J. A. Wege 2074) E, F L. chippendalei, widespread in arid areas of Western Australia and the Northern Territory, with the incised apical labellum appendage visible in the lower right flower (N. Gibson 6559, S. van Leeuwen, M. A. Langley & K. Brown). Photos by J. A. Wege (B, D) and K. Brown (F). Scale bar on maps 1000 km." figureDoi="10.3897/phytokeys.151.51909.figure7" httpUri="https://binary.pensoft.net/fig/420779" pageId="0" pageNumber="1">, 7B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benth" authorityYear="1837" class="Magnoliopsida" family="Stylidiaceae" genus="Stylidium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stylidium stipitatum" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stipitatum">Stylidium stipitatum</taxonomicName>
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Benth., in S.F.L. Endlicher, E. Fenzl, G. Bentham & H.W. Schott,
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<emphasis italics="true" pageId="0" pageNumber="1">Enum. Pl.</emphasis>
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: 72. 1837.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Stylidiaceae" genus="Coleostylis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleostylis umbellata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umbellata">Coleostylis umbellata</taxonomicName>
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Sond. in J.G.C. Lehmann, Pl. Preiss. 1(3): 391. 1845,
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<emphasis italics="true" pageId="0" pageNumber="1">nom. illeg.</emphasis>
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[
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<taxonomicName authorityName="Benth" authorityYear="1837" class="Magnoliopsida" family="Stylidiaceae" genus="Stylidium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stylidium stipitatum" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stipitatum">Stylidium stipitatum</taxonomicName>
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cited in synonymy]
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName family="Stylidiaceae" genus="Leewenhoekia" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Leewenhoekia stipitata" order="Asterales" pageId="0" pageNumber="1" rank="species" species="stipitata">Leewenhoekia stipitata</taxonomicName>
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(Benth.) F.Muell., Fragm. 4(27): 94. 1864,
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<emphasis italics="true" pageId="0" pageNumber="1">nom. inval.</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="1">nom. prov.</emphasis>
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName family="Stylidiaceae" genus="Leewenhoekia" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Leewenhoekia stipitata" order="Asterales" pageId="0" pageNumber="1" rank="species" species="stipitata">Leewenhoekia stipitata</taxonomicName>
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, orth. var.: F. von Mueller, Syst. Census Austral. Pl.: 86. 1882.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Type.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Australia. Western Australia</emphasis>
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: Swan River, [1833]
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<emphasis italics="true" pageId="0" pageNumber="1">
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K. von
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<normalizedToken originalValue="Hügel">Huegel</normalizedToken>
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s.n.
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</emphasis>
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(lectotype, here designated: W 0047173, all
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<taxonomicName class="Magnoliopsida" family="Stylidiaceae" genus="Levenhookia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Levenhookia" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Levenhookia</emphasis>
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</taxonomicName>
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material [i.e. excluding the individual of
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<taxonomicName class="Liliopsida" family="Restionaceae" genus="Centrolepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Centrolepis" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Centrolepis</emphasis>
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</taxonomicName>
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]).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Annual herb 2-18 cm high. Glandular hairs 0.1-0.5 mm long. Stem dark red to reddish brown, sometimes paler or greenish distally, simple or branched to varying degrees with porrect or ascending lateral branches, glandular-hairy. Leaves cauline, scattered, green (occasionally tinged red) or red-purple, lamina narrowly oblanceolate to oblanceolate, elliptic or ovate, 2.5-12 mm long including the petiole, 0.5-3(-4.5) mm wide, subacute to acute or obtuse, glandular-hairy on the abaxial surface and margins. Flowers in umbels, corymbs or short racemes, 1-ca. 150 per plant; bracts narrowly oblanceolate to oblanceolate, elliptic or linear, 2-10 mm long, glandular-hairy like the leaves; pedicels 3-18 mm long, glandular-hairy. Hypanthium depressed globose, globose or ellipsoid, 0.7-1.5 mm long, 0.7-1.8 mm wide, glandular-hairy. Calyx lobes
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<normalizedToken originalValue="±">+/-</normalizedToken>
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equal or with the anterior pair scarcely longer than the rest, 1.3-2.3 mm long, acute, moderately to sparsely glandular-hairy. Corolla bright to pale pink (rarely white) with a white throat, sometimes with two, elongated red-pink markings towards the base of each lobe near the margins; lobes evenly arranged or with the upper (posterior) ones
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<normalizedToken originalValue="±">+/-</normalizedToken>
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paired vertically, sometimes weakly recurved, obovate or elliptic with a slender claw, rounded, scarcely apiculate or bluntly pointed, glabrous or sometimes with sparse glandular hairs abaxially; anterior lobes equal to or slightly shorter and narrower than the posterior pair, 2.5-4.5 mm long, 1.6-2.8 mm wide; posterior lobes 2.7-4.5 mm long, 1.7-3.3 mm wide; tube creamy white, 0.2-0.5 mm long, obscured by the calyx lobes, glabrous. Labellum ventral, 2-3.5 mm long including a 0.9-1.5 mm long claw; hood yellow or whitish with dark red-maroon markings, sparsely glandular-hairy abaxially, scarcely papillate adaxially along the margins, papillate abaxially with a short, blunt, dark pink-red or sometimes yellow appendage at the cleft apex; basal appendages yellowish or white, rounded, 0.4-0.6 mm long. Column sheath white or yellowish, glabrous, 1.3-2.5 mm high on the posterior side with a thickened rim and broad anterior cleft, 3 pendulous appendages present on the inner surface towards the throat. Column creamy white, free, erect, slightly thickened distally, 2-4 mm long; stigmatic lobes to ca. 1 mm long, incurved, the lower-most developing while the column is hooded, the uppermost developing later. Capsule globose or ovoid, 1-2.3 mm long excluding calyx lobes. Seeds 0.4-0.5 mm long, 0.25-0.3 mm wide.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnostic features">
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<paragraph pageId="0" pageNumber="1">Diagnostic features.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Stylidiaceae" genus="Levenhookia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Levenhookia stipitata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stipitata">
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<emphasis italics="true" pageId="0" pageNumber="1">Levenhookia stipitata</emphasis>
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</taxonomicName>
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has glandular-hairy stems, bracts and calyx lobes, long pedicels (3-18 mm), a short corolla tube (obscured by the calyx lobes), a small, blunt, papillate appendage at the tip of the labellum, and a prominent column sheath (more than half the length of column). The corolla lobes may have a pair of elongated markings near the base (see notes below).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="phenology">
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<paragraph pageId="0" pageNumber="1">Phenology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Flowering from September to December; fruits have been collected from late October to December.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is widespread in Western Australia (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Comparative distributions and floral morphologies A, B L. stipitata, with a disjunct distribution in Western Australia and South Australia and flowers on long pedicels and with a prominent column sheath (J. A. Wege 1874). Inset showing coloured markings at the base of the corolla lobes that have been recorded for many populations (J. A. Wege 1873) C, D L. octomaculata, a Western Australian endemic with prominent corolla markings and a densely papillate labellum (J. A. Wege 2074) E, F L. chippendalei, widespread in arid areas of Western Australia and the Northern Territory, with the incised apical labellum appendage visible in the lower right flower (N. Gibson 6559, S. van Leeuwen, M. A. Langley & K. Brown). Photos by J. A. Wege (B, D) and K. Brown (F). Scale bar on maps 1000 km." figureDoi="10.3897/phytokeys.151.51909.figure7" httpUri="https://binary.pensoft.net/fig/420779" pageId="0" pageNumber="1">7A</figureCitation>
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), occurring in all bioregions within the South-West Province as well as the adjacent Yalgoo and Coolgardie bioregions. It has a more restricted distribution in South Australia where it is mostly confined to the Eyre Peninsula between Pinkawillinie Conservation Park and Point Boston, with an outlying record from the central Yorke Peninsula.
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</paragraph>
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<caption doi="10.3897/phytokeys.151.51909.figure7" httpUri="https://binary.pensoft.net/fig/420779" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
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Comparative distributions and floral morphologies
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<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
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<taxonomicName lsidName="L. stipitata" pageId="0" pageNumber="1" rank="species" species="stipitata">
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<emphasis italics="true" pageId="0" pageNumber="1">L. stipitata</emphasis>
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</taxonomicName>
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, with a disjunct distribution in Western Australia and South Australia and flowers on long pedicels and with a prominent column sheath (
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<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1874</emphasis>
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). Inset showing coloured markings at the base of the corolla lobes that have been recorded for many populations (
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<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1873</emphasis>
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)
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<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
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<taxonomicName lsidName="L. octomaculata" pageId="0" pageNumber="1" rank="species" species="octomaculata">
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<emphasis italics="true" pageId="0" pageNumber="1">L. octomaculata</emphasis>
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</taxonomicName>
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, a Western Australian endemic with prominent corolla markings and a densely papillate labellum (
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<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 2074</emphasis>
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)
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<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
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<taxonomicName lsidName="L. chippendalei" pageId="0" pageNumber="1" rank="species" species="chippendalei">
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<emphasis italics="true" pageId="0" pageNumber="1">L. chippendalei</emphasis>
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</taxonomicName>
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, widespread in arid areas of Western Australia and the Northern Territory, with the incised apical labellum appendage visible in the lower right flower (
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<emphasis italics="true" pageId="0" pageNumber="1">N. Gibson 6559</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="1">S. van Leeuwen</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="1">M.A. Langley & K. Brown</emphasis>
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). Photos by J.A. Wege (
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<emphasis bold="true" pageId="0" pageNumber="1">B, D</emphasis>
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) and K. Brown (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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). Scale bar on maps 1000 km.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="habitat">
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<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Stylidiaceae" genus="Levenhookia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Levenhookia stipitata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stipitata">
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<emphasis italics="true" pageId="0" pageNumber="1">Levenhookia stipitata</emphasis>
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</taxonomicName>
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grows in sand, sandy loam or clayey sand over limestone or laterite, more rarely in association with granite outcropping, usually on plains and hill-slopes but sometimes in depressions or seasonally damp habitats. Associated vegetation is varied and includes heathland, woodland,
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<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus marginata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="marginata">
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<emphasis italics="true" pageId="0" pageNumber="1">Eucalyptus marginata</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="K.D.Hill & L.A.S.Johnson" authorityYear="1995" class="Magnoliopsida" family="Myrtaceae" genus="Corymbia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corymbia calophylla" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="calophylla">
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<emphasis italics="true" pageId="0" pageNumber="1">Corymbia calophylla</emphasis>
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</taxonomicName>
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forest,
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<taxonomicName class="Magnoliopsida" family="Proteaceae" genus="Banksia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Banksia" order="Proteales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Banksia</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. wandoo" pageId="0" pageNumber="1" rank="species" species="wandoo">
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<emphasis italics="true" pageId="0" pageNumber="1">E. wandoo</emphasis>
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</taxonomicName>
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or
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<taxonomicName lsidName="E. cladocalyx" pageId="0" pageNumber="1" rank="species" species="cladocalyx">
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<emphasis italics="true" pageId="0" pageNumber="1">E. cladocalyx</emphasis>
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</taxonomicName>
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woodland, open mallee woodland or shrubland, and
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Acacia</emphasis>
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</taxonomicName>
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or
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<taxonomicName class="Agaricomycetes" family="Tricholomataceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melaleuca" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Melaleuca</emphasis>
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</taxonomicName>
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tall shrubland.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="conservation status">
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<paragraph pageId="0" pageNumber="1">Conservation status.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This common species is not considered to be under threat (
|
||
<bibRefCitation author="IUCN" journalOrPublisher="IUCN Gland, Switzerland and Cambridge, UK" pageId="0" pageNumber="1" refId="B17" refString="IUCN, 2012. IUCN Red List Categories and Criteria: version 3.1. IUCN Species Survival Commission. IUCN Gland, Switzerland and Cambridge, UK" title="IUCN Red List Categories and Criteria: version 3.1. IUCN Species Survival Commission." year="2012">IUCN (2012)</bibRefCitation>
|
||
: Least Concern); however, in South Australia, it is listed as Rare (Schedule 9) under the
|
||
<emphasis italics="true" pageId="0" pageNumber="1">National Parks and Wildlife Act 1972</emphasis>
|
||
(
|
||
<bibRefCitation author="Government of South Australia" journalOrPublisher="Four Gables Press, Kardinya, Western Australia" pageId="0" pageNumber="1" publicationUrl="https://www.legislation.sa.gov.au/LZ/C/A/NATIONAL%20PARKS%20AND%20WILDLIFE%20ACT%201972.aspx" refId="B15" refString="Government of South Australia, 2018. National Parks and Wildlife Act 1972. Version: 4.10.2018. https://www.legislation.sa.gov.au/LZ/C/A/NATIONAL%20PARKS%20AND%20WILDLIFE%20ACT%201972.aspx" title="National Parks and Wildlife Act 1972. Version: 4.10.2018." url="https://www.legislation.sa.gov.au/LZ/C/A/NATIONAL%20PARKS%20AND%20WILDLIFE%20ACT%201972.aspx" year="2018">Government of South Australia 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
From the Latin
|
||
<emphasis italics="true" pageId="0" pageNumber="1">stipitatus</emphasis>
|
||
(stipitate, provided with a stipe or little stalk), presumably with reference to the corolla segments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="vernacular name">
|
||
<paragraph pageId="0" pageNumber="1">Vernacular name.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Common Stylewort (
|
||
<bibRefCitation author="Erickson, R" journalOrPublisher="Paterson Brokensha, Perth" pageId="0" pageNumber="1" refId="B10" refString="Erickson, R, 1958. Triggerplants. Paterson Brokensha, Perth" title="Triggerplants." year="1958">Erickson 1958</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="typification">
|
||
<paragraph pageId="0" pageNumber="1">Typification.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The only
|
||
<normalizedToken originalValue="Hügel">Huegel</normalizedToken>
|
||
specimen that is known is held at Naturhistorisches Museum Wien (W 0047173) and bears annotations by Bentham (as '
|
||
<taxonomicName authorityName="Benth" authorityYear="1837" class="Magnoliopsida" family="Stylidiaceae" genus="Stylidium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stylidium stipitatum" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stipitatum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stylidium stipitatum</emphasis>
|
||
</taxonomicName>
|
||
'), Sonder (as '
|
||
<taxonomicName authorityName="Sond" authorityYear="1845" class="Magnoliopsida" family="Stylidiaceae" genus="Coleostylis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleostylis umbellulata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umbellulata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Coleostylis umbellulata</emphasis>
|
||
</taxonomicName>
|
||
') and Mildbraed (as '
|
||
<taxonomicName class="Magnoliopsida" family="Stylidiaceae" genus="Levenhookia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Levenhookia stipitata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stipitata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Levenhookia stipitata</emphasis>
|
||
</taxonomicName>
|
||
'). The collection is mixed, comprising four individuals of
|
||
<taxonomicName lsidName="L. stipitata" pageId="0" pageNumber="1" rank="species" species="stipitata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">L. stipitata</emphasis>
|
||
</taxonomicName>
|
||
(and associated fragments in the attached packets) and a single individual of
|
||
<taxonomicName authorityName="Roem & Schult" authorityYear="1817" baseAuthorityName="R. Br." class="Liliopsida" family="Restionaceae" genus="Centrolepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Centrolepis aristata" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="aristata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Centrolepis aristata</emphasis>
|
||
</taxonomicName>
|
||
(R.Br.) Poir. (
|
||
<taxonomicName family="Centrolepidaceae" lsidName="" pageId="0" pageNumber="1" rank="family">Centrolepidaceae</taxonomicName>
|
||
) mounted with the
|
||
<taxonomicName class="Magnoliopsida" family="Stylidiaceae" genus="Levenhookia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Levenhookia" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Levenhookia</emphasis>
|
||
</taxonomicName>
|
||
in the centre of the sheet. All
|
||
<taxonomicName class="Magnoliopsida" family="Stylidiaceae" genus="Levenhookia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Levenhookia" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Levenhookia</emphasis>
|
||
</taxonomicName>
|
||
material on this sheet is designated herein as the lectotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="notes">
|
||
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Corolla markings are often absent in
|
||
<taxonomicName lsidName="L. stipitata" pageId="0" pageNumber="1" rank="species" species="stipitata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">L. stipitata</emphasis>
|
||
</taxonomicName>
|
||
; however, two discrete markings at the base of each lobe can be consistently present in populations in Western Australia and South Australia (e.g.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">D.E. Murfet 2270 & R.L. Taplin</emphasis>
|
||
(AD),
|
||
<emphasis italics="true" pageId="0" pageNumber="1">D.E. Murfet 4494 & A. Lowrie</emphasis>
|
||
(AD),
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1562 & B.P. Miller</emphasis>
|
||
(PERTH),
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1873</emphasis>
|
||
(PERTH: Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Comparative distributions and floral morphologies A, B L. stipitata, with a disjunct distribution in Western Australia and South Australia and flowers on long pedicels and with a prominent column sheath (J. A. Wege 1874). Inset showing coloured markings at the base of the corolla lobes that have been recorded for many populations (J. A. Wege 1873) C, D L. octomaculata, a Western Australian endemic with prominent corolla markings and a densely papillate labellum (J. A. Wege 2074) E, F L. chippendalei, widespread in arid areas of Western Australia and the Northern Territory, with the incised apical labellum appendage visible in the lower right flower (N. Gibson 6559, S. van Leeuwen, M. A. Langley & K. Brown). Photos by J. A. Wege (B, D) and K. Brown (F). Scale bar on maps 1000 km." figureDoi="10.3897/phytokeys.151.51909.figure7" httpUri="https://binary.pensoft.net/fig/420779" pageId="0" pageNumber="1">7B</figureCitation>
|
||
, inset),
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege</emphasis>
|
||
2068 (PERTH)) or variably present within a population (e.g.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 777</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 2077</emphasis>
|
||
). A comparison with
|
||
<taxonomicName lsidName="L. octomaculata" pageId="0" pageNumber="1" rank="species" species="octomaculata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">L. octomaculata</emphasis>
|
||
</taxonomicName>
|
||
, an allied species named for its corolla markings, is provided under the notes for that species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="illustrations">
|
||
<paragraph pageId="0" pageNumber="1">Illustrations.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">L. Diels & E. Pritzel, Bot. Jahrb. Syst. 35: 598, fig. 67 A-C. 1905 [reproduced by J. Mildbraed in H.G.A. Engler, Pflanzenr. 35: 29, fig. 10A-C. 1908]; R. Erickson, Triggerplants 201, Pl. 57, No. 2 and 208, Pl. 58, Nos. 8-14. 1958; B.J. Grieve & W.E. Blackall, How to know W. Austral. wildfl. 4: 766, no. 6. 1982; J. Wheeler, N. Marchant & M. Lewington, Fl. South West 2: 903. 2002.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="1">Selected specimens examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Australia. Western Australia</emphasis>
|
||
: Mt Merivale, 20 km E of Esperance, 25 Oct 1995,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. Archer 168</emphasis>
|
||
(MEL); Brixton Street Wetlands, Kenwick, 10 Nov 2011,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">K.L. Brown 888 & G. Paczkowska</emphasis>
|
||
(PERTH); Darling Range escarpment, W of Walyunga Reserve, 17 Dec 1971,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N.T. Burbidge 7882</emphasis>
|
||
(CANB); Dryandra State Forest, NE of Congelin, at the sigma bend of Patonga Rd, 24 Oct 1991,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">W. Greuter 23183</emphasis>
|
||
(PERTH); Sheepwash State Forest, E boundary at 'The
|
||
<normalizedToken originalValue="Pass’">Pass'</normalizedToken>
|
||
, 13 Dec 1998,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B.G. Hammersley 2137</emphasis>
|
||
(PERTH); Reserve 23229/1255, Brookton Hwy, Armadale, 5 km SE of Kinsella Rd, 11 Dec 2004,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">F. Hort</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J. Hort & B. Hort 2451</emphasis>
|
||
(PERTH); Wongamine Nature Reserve, ca. 13 km NE of Toodyay, 7 Oct 1995,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T.R. Lally & B.J. Lepschi 775</emphasis>
|
||
(PERTH); Sukey Hill, 3.5 km E of Cranbrook, 13 Nov 1995,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T.R. Lally & B.J. Lepschi 880</emphasis>
|
||
(PERTH); 1.85 km along Park Rd from Great Eastern Hwy, John Forrest National Park, 14 Nov 1995,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 120</emphasis>
|
||
(PERTH); 20 m NE along gravel track off Del Park Rd, 6.7 km from South Western Hwy, 29 Nov 1995,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 156 & P. French</emphasis>
|
||
(PERTH); ca. 900 m from Eagle Bay settlement on Meelup Beach Rd, 8 Nov 2002,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 777</emphasis>
|
||
(PERTH); 1.6 km E on Cadda Rd from Brand Hwy, Badgingarra National Park,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1688 & W.S. Armbruster</emphasis>
|
||
, 20 Oct 2009 (PERTH); ca. 1.7 km on Canning Road from Pickering Brook Road, Korung National Park, 12 Nov 2003,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1100</emphasis>
|
||
(PERTH); Qualen Rd, just W of Kent Rd, Wandoo National Park, 29 Nov 2008,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege & B.P. Miller JAW 1562</emphasis>
|
||
(PERTH); 6.4 km W of Brand Hwy on Bibby Rd, SW of Badgingarra, 20 Oct 2011,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1873</emphasis>
|
||
(PERTH); 4.7 km E of Brand Hwy on Wanamal West Rd, Boonanarring Nature Reserve, 20 Oct 2011,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 1874</emphasis>
|
||
(CANB, MEL, PERTH); 6.4 km E of Sundalara Rd on Tomkins Rd, NE of Eneabba, 30 Oct 2018,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">J.A. Wege 2077</emphasis>
|
||
(PERTH);
|
||
<emphasis bold="true" pageId="0" pageNumber="1">South Australia</emphasis>
|
||
: Wanilla, some 25 km NW of Port Lincoln, 8 Nov 1968,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C.R. Alcock 2542</emphasis>
|
||
(AD, MEL); Granite outcrop NW of Lienerts/Woolford Track in Pinkawillinie C[onservation] P[ark], 3 Nov 2009,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T.S. Te 859 & T.S. Croft</emphasis>
|
||
(AD); Site BS162-MIN01101, Patchid 20131 [9.1 km direct ENE of Roger Corner, Yorke Peninsula], 13 Oct 2004,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">L.M.B. Heard & N.R. Neagle BS162-1754</emphasis>
|
||
(AD); Verran Tanks Conservation Park, 10 Oct 1991,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">D.E. Murfet 1299b</emphasis>
|
||
(AD); Murrunatta Conservation Park, 15 Oct 1995,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">D.E. Murfet & R.L. Taplin 2270</emphasis>
|
||
(AD).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |