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<document id="05B13DD23622DE96D22877BDC046656A" ID-CLB-Dataset="20340" ID-DOI="10.11646/zootaxa.5105.3.2" ID-GBIF-Dataset="4f25a761-8f7b-47d3-84c9-ad2917b90f74" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332731" ID-ZooBank="83013C70-69E4-42BD-8CDD-0FA320927290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646597716981" checkinUser="plazi" docAuthor="Bhansali, Sneha &amp; Wesener, Thomas" docDate="2022" docId="03FBA627FFA2AD0AE389FA06FA15FD12" docLanguage="en" docName="zootaxa.5105.3.2.pdf" docOrigin="Zootaxa 5105 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zephronia golovatchi Srisonchai, Sutcharit &amp; Likhitrakarn 2021" docType="key" docVersion="6" lastPageNumber="377" masterDocId="FFC2DE5FFFB1AD1EE31EFF84FF89FFAF" masterDocTitle="New Thai giant pill-millipede species, with new genetic barcoding data (Diplopoda Sphaerotheriida, Zephroniidae)" masterLastPageNumber="380" masterPageNumber="357" pageNumber="376" updateTime="1699317184639" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="573AB8EED8E78356C2A7303D0DBB0746" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9FC09D6E2CEA639E6BF6EEA012AB7D75">
<mods:title id="0BECE84E33B081CC992E5A8DCBAD64E6">New Thai giant pill-millipede species, with new genetic barcoding data (Diplopoda Sphaerotheriida, Zephroniidae)</mods:title>
</mods:titleInfo>
<mods:name id="82DE618CCADCA3B3CA622438A80992B5" type="personal">
<mods:role id="E705BC27CEF884DC5A170A39F0F45DF2">
<mods:roleTerm id="6798657D7F6C6B5B8F20529ACAC649C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FCB5E30AE25A0AAD12F5699A6A2D0812">Bhansali, Sneha</mods:namePart>
<mods:nameIdentifier id="A66FDE782F87E34666EE6DCFA37803C4" type="ORCID">0000-0002-5440-9958</mods:nameIdentifier>
<mods:affiliation id="0D3E297C5992B8AF1D06EEBCCE82816D">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="B0A31405A5164B6FCAF507F615E1EBF0" type="email">sneharbhansali@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E6F368A2DD823F17E8E6506FAC4A1241" type="personal">
<mods:role id="8F9D44361FBFF82B0F03CA850F6CF288">
<mods:roleTerm id="7F57C74D62F448C74F9B1EEB73A12E37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C0750BCDFF1F62DBDAEF18FB02F69CD">Wesener, Thomas</mods:namePart>
<mods:nameIdentifier id="D4608A5338297CAE8A0C0D068811AA64" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
<mods:affiliation id="DD7B11CBC530FA838361762CF76E3A42">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany. &amp; T. Wesener @ leibniz-zfmk. de; https: // orcid. org / 0000 - 0002 - 2028 - 3541</mods:affiliation>
<mods:nameIdentifier id="E6E2F13FCA28D59E2FDB3700423ABD04" type="email">esener@leibniz-zfmk.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="C3E36BD158A283802940D4EADCAAE323">text</mods:typeOfResource>
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<mods:titleInfo id="2B768502DCEA4F9D60FA9D5BFE27BC05">
<mods:title id="AC5738D7209B59626A89FE83F59FCE89">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5C55B7BD3C3A8F53627E37B04E062CB4">
<mods:date id="B0DE70D8896A1203B96602E50C9A88F6">2022</mods:date>
<mods:detail id="F6D91EBEEDB576DE9C8E8D6C66D2D042" type="pubDate">
<mods:number id="4AD35B1398ADE819A909DD80A837A444">2022-03-04</mods:number>
</mods:detail>
<mods:detail id="FD77D8252B0A1501F93D822970A1B846" type="volume">
<mods:number id="4B5968501AA5CF19A21818D949D709D3">5105</mods:number>
</mods:detail>
<mods:detail id="6D4F5C590F5E93AF774D6973BFAD4555" type="issue">
<mods:number id="ACD90B08A540CCCEB83F76C89A31A644">3</mods:number>
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<mods:classification id="12191177CA4F83BBC874E725393A95F9">journal article</mods:classification>
<mods:identifier id="093746E0BBB7E7EF255E8E3F7E16F8D2" type="CLB-Dataset">20340</mods:identifier>
<mods:identifier id="4922BC0ABDE1EA85721FCB6EBAF62CF0" type="DOI">10.11646/zootaxa.5105.3.2</mods:identifier>
<mods:identifier id="BA0871FB5C9BD62498197F2F57B4AB33" type="GBIF-Dataset">4f25a761-8f7b-47d3-84c9-ad2917b90f74</mods:identifier>
<mods:identifier id="FE2E41691E7C116F46382998E927EDF9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F0159DEE31D395BD089D89C9C98700C4" type="Zenodo-Dep">6332731</mods:identifier>
<mods:identifier id="6F94D6C577CC5508ED0B065EE6F5DD8E" type="ZooBank">83013C70-69E4-42BD-8CDD-0FA320927290</mods:identifier>
</mods:mods>
<treatment id="03FBA627FFA2AD0AE389FA06FA15FD12" ID-DOI="http://doi.org/10.5281/zenodo.6333455" ID-GBIF-Taxon="193890964" ID-Zenodo-Dep="6333455" LSID="urn:lsid:plazi:treatment:03FBA627FFA2AD0AE389FA06FA15FD12" httpUri="http://treatment.plazi.org/id/03FBA627FFA2AD0AE389FA06FA15FD12" lastPageId="20" lastPageNumber="377" pageId="19" pageNumber="376">
<subSubSection id="C34844BAFFA2AD0DE389FA06FC4BFA33" box="[151,962,1410,1437]" pageId="19" pageNumber="376" type="nomenclature">
<paragraph id="8BED1731FFA2AD0DE389FA06FC4BFA33" blockId="19.[151,962,1410,1437]" box="[151,962,1410,1437]" pageId="19" pageNumber="376">
<heading id="D0A5A05DFFA2AD0DE389FA06FC4BFA33" bold="true" box="[151,962,1410,1437]" fontSize="11" level="1" pageId="19" pageNumber="376" reason="1">
<taxonomicName id="4C526CB2FFA2AD0DE389FA06FC4BFA33" authority="Srisonchai, Sutcharit &amp; Likhitrakarn, 2021" authorityName="Srisonchai, Sutcharit &amp; Likhitrakarn" authorityYear="2021" box="[151,962,1410,1437]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="376" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="B926CB23FFA2AD0DE389FA06FC4BFA33" bold="true" box="[151,962,1410,1437]" pageId="19" pageNumber="376">
<emphasis id="B926CB23FFA2AD0DE389FA06FE12FA32" bold="true" box="[151,411,1410,1437]" italics="true" pageId="19" pageNumber="376">Zephronia golovatchi</emphasis>
<bibRefCitation id="EFC36AC0FFA2AD0DE2BCFA06FC4BFA33" author="Srisonchai, R. &amp; Sutcharit, C. &amp; Likhitrakarn, N." box="[418,962,1410,1436]" pageId="19" pageNumber="376" pagination="19 - 56" refId="ref14171" refString="Srisonchai, R., Sutcharit, C. &amp; Likhitrakarn, N. (2021) The giant pill-millipede genus Zephronia Gray, 1832 from Thailand, with a redescription of Z. siamensis Hirst, 1907 and descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae). ZooKeys, 1067, 19 - 56. https: // doi. org / 10.3897 / zookeys. 1067.72369" type="journal article" year="2021">Srisonchai, Sutcharit &amp; Likhitrakarn, 2021</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFA2AD0DE389FA4FFC93F9FE" pageId="19" pageNumber="376" type="materials_examined">
<paragraph id="8BED1731FFA2AD0DE389FA4FFC93F9FE" blockId="19.[151,1436,1482,1689]" pageId="19" pageNumber="376">
<emphasis id="B926CB23FFA2AD0DE389FA4FFEEDFA4A" bold="true" box="[151,356,1482,1509]" pageId="19" pageNumber="376">New locality data</emphasis>
:
<materialsCitation id="3B3A1D6CFFA2AD0DE26FFA4FFDFFF9A6" ID-GBIF-Occurrence="3698753332" collectingDate="2017-06-10" collectionCode="ZFMK" latitude="14.516444" location="Highway" longLatPrecision="1" longitude="101.398026" municipality="Pak Ching District" pageId="19" pageNumber="376" specimenCode="MYR6262" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima">
<specimenCount id="9D54DCB8FFA2AD0DE26FFA4FFE15FA4A" box="[369,412,1483,1509]" pageId="19" pageNumber="376" type="male">1 ♂</specimenCount>
,
<emphasis id="B926CB23FFA2AD0DE2B9FA4FFDF5FA4B" bold="true" box="[423,636,1482,1509]" pageId="19" pageNumber="376">
<collectionCode id="ED438FF4FFA2AD0DE2B9FA4FFE72FA4A" box="[423,507,1483,1509]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="376" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBF4BF4AFFA2AD0DE11FFA4EFDF5FA4B" box="[513,636,1482,1508]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="376" type="Museum">MYR6262</specimenCode>
</emphasis>
,
<collectingRegion id="4996D9D3FFA2AD0DE196FA4FFC5CFA4A" box="[648,981,1483,1509]" country="Thailand" name="Nakhon Ratchasima" pageId="19" pageNumber="376">Nakhon Ratchasima Province</collectingRegion>
,
<collectingMunicipality id="6B898D4BFFA2AD0DE0FEFA4FFB38FA4A" box="[992,1201,1483,1509]" pageId="19" pageNumber="376">Pak Ching District</collectingMunicipality>
, near
<location id="8E8D41EAFFA2AD0DE7EFFA4FFAD1FA4A" LSID="urn:lsid:plazi:treatment:03FBA627FFA2AD0AE389FA06FA15FD12:8E8D41EAFFA2AD0DE7EFFA4FFAD1FA4A" box="[1265,1368,1483,1509]" latitude="14.516444" longLatPrecision="1" longitude="101.398026" municipality="Pak Ching District" name="Highway" pageId="19" pageNumber="376" stateProvince="Nakhon Ratchasima">Highway</location>
3052,
<geoCoordinate id="EE6671F6FFA2AD0DE389FA6AFEB8F9A6" box="[151,305,1518,1545]" degrees="14" direction="north" minutes="30" orientation="latitude" pageId="19" pageNumber="376" precision="1" seconds="59.2" value="14.516444">14°30'59.2''N</geoCoordinate>
,
<geoCoordinate id="EE6671F6FFA2AD0DE221FA6AFE6DF9A6" box="[319,484,1518,1545]" degrees="101" direction="east" minutes="23" orientation="longitude" pageId="19" pageNumber="376" precision="1" seconds="52.9" value="101.398026">101°23'52.9''E</geoCoordinate>
,
<date id="FFEC31F1FFA2AD0DE2EDFA6BFDFFF9A6" box="[499,630,1519,1545]" pageId="19" pageNumber="376" value="2017-06-10">
<collectingDate id="EFA8C819FFA2AD0DE2EDFA6BFDFFF9A6" box="[499,630,1519,1545]" pageId="19" pageNumber="376" value="2017-06-10">10.VI.2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3A1D6CFFA2AD0DE19AFA6BFB74F9A6" ID-GBIF-Occurrence="3698753305" box="[644,1277,1518,1545]" collectingDate="2017-06-10" collectionCode="ZFMK" latitude="14.516444" location="Highway" longLatPrecision="1" longitude="101.398026" municipality="Pak Ching District" pageId="19" pageNumber="376" specimenCode="MYR6779" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Nakhon Ratchasima">
<specimenCount id="9D54DCB8FFA2AD0DE19AFA6BFD3BF9A6" box="[644,690,1519,1545]" pageId="19" pageNumber="376" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D54DCB8FFA2AD0DE1DFFA6BFC8DF9A6" box="[705,772,1519,1545]" pageId="19" pageNumber="376" type="female">2 ♀♀</specimenCount>
,
<emphasis id="B926CB23FFA2AD0DE00DFA6BFC65F9A7" bold="true" box="[787,1004,1518,1545]" pageId="19" pageNumber="376">
<collectionCode id="ED438FF4FFA2AD0DE00DFA6BFCEEF9A6" box="[787,871,1519,1545]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="376" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBF4BF4AFFA2AD0DE06FFA6AFC65F9A7" box="[881,1004,1518,1544]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="376" type="Museum">MYR6779</specimenCode>
</emphasis>
, same data as previous
</materialsCitation>
;
<materialsCitation id="3B3A1D6CFFA2AD0DE615FA6BFD90F982" ID-GBIF-Occurrence="3698753324" collectingDate="2017-06-10" collectionCode="ZFMK" latitude="14.516444" location="Highway" longLatPrecision="1" longitude="101.398026" municipality="Pak Ching District" pageId="19" pageNumber="376" specimenCode="MYR7236" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima">
<specimenCount id="9D54DCB8FFA2AD0DE615FA6BFAB3F9A6" box="[1291,1338,1519,1545]" pageId="19" pageNumber="376" type="male">1 ♂</specimenCount>
,
<emphasis id="B926CB23FFA2AD0DE656FA6BFE9BF983" bold="true" pageId="19" pageNumber="376">
<collectionCode id="ED438FF4FFA2AD0DE656FA6BFA15F9A6" box="[1352,1436,1519,1545]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="376" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBF4BF4AFFA2AD0DE389F996FE9BF983" box="[151,274,1554,1580]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="376" type="Museum">MYR7236</specimenCode>
</emphasis>
, same data as previous
</materialsCitation>
;
<materialsCitation id="3B3A1D6CFFA2AD0DE13BF997FC93F9FE" ID-GBIF-Occurrence="3698753321" collectingDate="1984-09-29" collectingDateMax="1984-10-06" collectingDateMin="1984-09-29" collectorName="Karsholt &amp; Lomholdt &amp; Nielsen." elevation="700" location="Khao Yai National Park" pageId="19" pageNumber="376" specimenCount="2" specimenCount-female="2" stateProvince="Nakhon Nayok">
<specimenCount id="9D54DCB8FFA2AD0DE13BF997FDECF982" box="[549,613,1555,1581]" pageId="19" pageNumber="376" type="female">2 ♀♀</specimenCount>
,
<emphasis id="B926CB23FFA2AD0DE16CF997FC8DF983" bold="true" box="[626,772,1554,1581]" pageId="19" pageNumber="376">NHMD K53</emphasis>
,
<collectingRegion id="4996D9D3FFA2AD0DE00CF997FBA3F982" box="[786,1066,1555,1581]" country="Thailand" name="Nakhon Nayok" pageId="19" pageNumber="376">Nakhon Nayok Province</collectingRegion>
,
<location id="8E8D41EAFFA2AD0DE72BF997FACAF982" LSID="urn:lsid:plazi:treatment:03FBA627FFA2AD0AE389FA06FA15FD12:8E8D41EAFFA2AD0DE72BF997FACAF982" box="[1077,1347,1555,1581]" name="Khao Yai National Park" pageId="19" pageNumber="376" stateProvince="Nakhon Nayok">Khao Yai National Park</location>
,
<quantity id="4CAABAD4FFA2AD0DE650F997FA1EF982" box="[1358,1431,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="376" unit="m" value="700.0">
<elevation id="007FF002FFA2AD0DE650F997FA1EF982" box="[1358,1431,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="376" unit="m" value="700.0">700 m</elevation>
</quantity>
, leg.
<date id="FFEC31F1FFA2AD0DE3D9F9B3FE08F9FE" box="[199,385,1591,1617]" pageId="19" pageNumber="376" value="1984-09-29" valueMax="1984-10-06" valueMin="1984-09-29">
<collectingDate id="EFA8C819FFA2AD0DE3D9F9B3FE08F9FE" box="[199,385,1591,1617]" pageId="19" pageNumber="376" value="1984-09-29" valueMax="1984-10-06" valueMin="1984-09-29">29.IX–6.X.1984</collectingDate>
</date>
, leg.
<collectorName id="26A772E7FFA2AD0DE2A2F9B3FD96F9FE" box="[444,543,1591,1617]" pageId="19" pageNumber="376">Karsholt</collectorName>
,
<collectorName id="26A772E7FFA2AD0DE135F9B3FD10F9FE" box="[555,665,1591,1617]" pageId="19" pageNumber="376">Lomholdt</collectorName>
&amp;
<collectorName id="26A772E7FFA2AD0DE1A3F9B3FC93F9FE" box="[701,794,1591,1617]" pageId="19" pageNumber="376">Nielsen.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFA2AD0DE3D9F9DFFCF3F8AB" pageId="19" pageNumber="376" type="discussion">
<paragraph id="8BED1731FFA2AD0DE3D9F9DFFC0BF936" blockId="19.[151,1436,1482,1689]" pageId="19" pageNumber="376">
<emphasis id="B926CB23FFA2AD0DE3D9F9DFFEBDF9DA" bold="true" box="[199,308,1627,1653]" pageId="19" pageNumber="376">Remarks</emphasis>
. Both localities lie close to the
<typeStatus id="54E9A993FFA2AD0DE190F9DFFD37F9DA" box="[654,702,1627,1653]" pageId="19" pageNumber="376">type</typeStatus>
locality (
<figureCitation id="13690BB4FFA2AD0DE038F9DFFC09F9DA" box="[806,896,1627,1653]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="19" pageNumber="376">Fig. 2A</figureCitation>
). However, specimens from both localities differ significantly genetically and might represent two different species.
</paragraph>
<subSection id="E2DD0EDAFFA2AD0DE389F96FFCF3F8AB" box="[151,890,1770,1797]" pageId="19" pageNumber="376" type="multiple">
<paragraph id="8BED1731FFA2AD0DE389F96FFCF3F8AB" blockId="19.[151,890,1770,1797]" box="[151,890,1770,1797]" pageId="19" pageNumber="376">
<heading id="D0A5A05DFFA2AD0DE389F96FFCF3F8AB" bold="true" box="[151,890,1770,1797]" fontSize="11" level="1" pageId="19" pageNumber="376" reason="1">
<emphasis id="B926CB23FFA2AD0DE389F96FFCF3F8AB" bold="true" box="[151,890,1770,1797]" pageId="19" pageNumber="376">
Updated key to confirmed species of
<taxonomicName id="4C526CB2FFA2AD0DE17DF96EFD6BF8AA" authorityName="Gray" authorityYear="1832" box="[611,738,1770,1797]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B926CB23FFA2AD0DE17DF96EFD6BF8AA" bold="true" box="[611,738,1770,1797]" italics="true" pageId="19" pageNumber="376">Zephronia</emphasis>
</taxonomicName>
in
<collectingCountry id="F34557A1FFA2AD0DE016F96EFCF3F8AB" box="[776,890,1770,1796]" name="Thailand" pageId="19" pageNumber="376">Thailand</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSection>
</subSubSection>
<subSubSection id="C34844BAFFA2AD0AE389F8ADFA15FD12" lastPageId="20" lastPageNumber="377" pageId="19" pageNumber="376" type="key">
<key id="FFC34000FFA2AD0AE389F8ADFA15FD12" lastPageId="20" lastPageNumber="377" pageId="19" pageNumber="376">
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<paragraph id="8BED1731FFA2AD0DE389F8ADFA15F8F4" blockId="19.[151,1437,1833,2031]" pageId="19" pageNumber="376">
<keyLead id="30A3B404FFA2AD0DE389F8ADFA15F8F4" pageId="19" pageNumber="376">
<emphasis id="B926CB23FFA2AD0DE389F8ADFF2AF891" bold="true" box="[151,163,1833,1854]" pageId="19" pageNumber="376">1</emphasis>
: Tarsi of leg pair 4–21 with single apical spine, third leg pair without apical spine. Male antennae, antennomere 6 swollen, not axe-shaped...................................... ..
<taxonomicName id="4C526CB2FFA2AD0DE028F8C3FB16F8F3" ID-CoL="9BDFN" authority="Rosenmejer &amp; Wesener" authorityName="Rosenmejer &amp; Wesener" authorityYear="2021" box="[822,1183,1862,1884]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="376" phylum="Arthropoda" rank="species" species="viridisoma">
<emphasis id="B926CB23FFA2AD0DE028F8C3FC3FF8F4" box="[822,950,1862,1884]" italics="true" pageId="19" pageNumber="376">Z. viridisoma</emphasis>
Rosenmejer &amp; Wesener
</taxonomicName>
in
<bibRefCitation id="EFC36AC0FFA2AD0DE7A0F8C2FA15F8F4" author="Rosenmejer, T. &amp; Enghoff, H. &amp; Moritz, L. &amp; Wesener, T." box="[1214,1436,1862,1884]" pageId="19" pageNumber="376" pagination="208 - 232" refId="ref13687" refString="Rosenmejer, T., Enghoff, H., Moritz, L. &amp; Wesener, T. (2021) Integrative description of new giant pill-millipedes from southern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae). European Journal of Taxonomy, 762, 208 - 232. https: // doi. org / 10.5852 / ejt. 2021.762.1457" type="journal article" year="2021">
Rosenmejer
<emphasis id="B926CB23FFA2AD0DE628F8C3FAEEF8F4" box="[1334,1383,1862,1884]" italics="true" pageId="19" pageNumber="376">et al.</emphasis>
2021
</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA2AD0DE389F8E0FA15F8D7" blockId="19.[151,1437,1833,2031]" box="[151,1436,1891,1914]" pageId="19" pageNumber="376">
<keyLead id="30A3B404FFA2AD0DE389F8E0FA15F8D7" box="[151,1436,1891,1914]" pageId="19" pageNumber="376">
- Tarsi of leg pair 4–21 with 3 apical spines, third leg pair with apical spine. Male antennae, antennomere 6 axe-shaped......
<emphasis id="B926CB23FFA2AD0DE68EF8E7FA15F8D7" bold="true" box="[1424,1436,1891,1912]" pageId="19" pageNumber="376">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A60F94FFA2AD0DE389F805FA15F841" pageId="19" pageNumber="376">
<paragraph id="8BED1731FFA2AD0DE389F805FA15F81C" blockId="19.[151,1437,1833,2031]" pageId="19" pageNumber="376">
<keyLead id="30A3B404FFA2AD0DE389F805FA15F81C" pageId="19" pageNumber="376">
<emphasis id="B926CB23FFA2AD0DE389F805FF2AF839" bold="true" box="[151,163,1921,1942]" pageId="19" pageNumber="376">2</emphasis>
: Conspicuous colour pattern with a vivid green, glabrous or dull, not golden setae. Palpi of gnathochilarium, sensilla in clusters.............................................................................................
<emphasis id="B926CB23FFA2AD0DE68EF81AFA15F81C" bold="true" box="[1424,1436,1950,1971]" pageId="19" pageNumber="376">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA2AD0DE389F838FA15F841" blockId="19.[151,1437,1833,2031]" pageId="19" pageNumber="376">
<keyLead id="30A3B404FFA2AD0DE389F838FA15F841" pageId="19" pageNumber="376">
- Brown, grey, or brown-greenish color pattern with dense coat of 'golden' setae. Palpi of gnathochilarium, sensilla in single field................................................................................................
<emphasis id="B926CB23FFA2AD0DE68EF85DFA15F841" bold="true" box="[1424,1436,2009,2030]" pageId="19" pageNumber="376">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A60F94FFA5AD0AE389FF13FA15FE8D" pageId="20" pageNumber="377">
<paragraph id="8BED1731FFA5AD0AE389FF13FA15FF65" blockId="20.[151,1437,151,701]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FF13FA15FF65" pageId="20" pageNumber="377">
<emphasis id="B926CB23FFA5AD0AE389FF13FF2AFF03" bold="true" box="[151,163,151,172]" pageId="20" pageNumber="377">3</emphasis>
: Small bodied, around
<quantity id="4CAABAD4FFA5AD0AE2A1FF1CFD8FFF02" box="[447,518,152,173]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="377" unit="mm" value="20.0">20 mm</quantity>
. Unique color pattern, green-whitish with red on the anal shield. Tergites setose, dull. Anterior telopod, podomere 3 with up to 7 crenulated teeth. Antennae, antennomere 6 with 50–60 cones....
<taxonomicName id="4C526CB2FFA5AD0AE7AAFF31FA15FF65" ID-CoL="5D2SZ" authority="Hirst, 1907" authorityName="Hirst" authorityYear="1907" box="[1204,1436,181,203]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B926CB23FFA5AD0AE7AAFF31FAA3FF65" box="[1204,1322,181,202]" italics="true" pageId="20" pageNumber="377">Z. siamensis</emphasis>
<bibRefCitation id="EFC36AC0FFA5AD0AE62EFF31FA15FF65" author="Hirst, A. S." box="[1328,1436,181,203]" pageId="20" pageNumber="377" pagination="215 - 219" refId="ref13153" refString="Hirst, A. S. (1907) On four new pill-millipedes from the Malay Peninsula and Siam. The Annals and Magazine of Natural History, Series 7, 20 (117), 215 - 219. https: // doi. org / 10.1080 / 00222930709487327" type="journal article" year="1907">Hirst, 1907</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA5AD0AE389FF57FA15FE8D" blockId="20.[151,1437,151,701]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FF57FA15FE8D" pageId="20" pageNumber="377">
- Large, around
<quantity id="4CAABAD4FFA5AD0AE270FF56FE3AFF47" box="[366,435,210,232]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="377" unit="mm" value="40.0">40 mm</quantity>
. Unique color pattern, with a vivid green-lightbrownish stripes. Tergite glabrous, shiny. Anterior telopod variable, without or only very few sclerotized teeth. Antennae, antennomere 6 with 70–90 cones........................................................................
<taxonomicName id="4C526CB2FFA5AD0AE05EFE89FB13FE8C" authority="Likhitrakarn &amp; Golovatch" authorityName="Likhitrakarn &amp; Golovatch" authorityYear="2021" box="[832,1178,269,291]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="phrain">
<emphasis id="B926CB23FFA5AD0AE05EFE89FC10FE8D" box="[832,921,269,290]" italics="true" pageId="20" pageNumber="377">Z. phrain</emphasis>
Likhitrakarn &amp; Golovatch
</taxonomicName>
in
<bibRefCitation id="EFC36AC0FFA5AD0AE7A6FE89FA15FE8D" author="Likhitrakarn, N. &amp; Golovatch, S. I. &amp; Srisonchai, R. &amp; Sutcharit, C." box="[1208,1436,269,291]" pageId="20" pageNumber="377" pagination="12 - 26" refId="ref13325" refString="Likhitrakarn, N., Golovatch, S. I., Srisonchai, R. &amp; Sutcharit, C. (2021) Two New Species of the Giant Pill-Millipede Genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae). Tropical Natural History, 21, 12 - 26." type="journal article" year="2021">
Likhitrakarn
<emphasis id="B926CB23FFA5AD0AE628FE89FAEEFE8D" box="[1334,1383,269,290]" italics="true" pageId="20" pageNumber="377">et al.</emphasis>
2021
</bibRefCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A60F94FFA5AD0AE389FEAEFA15FEF2" pageId="20" pageNumber="377">
<paragraph id="8BED1731FFA5AD0AE389FEAEFA15FE90" blockId="20.[151,1437,151,701]" box="[151,1436,298,320]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FEAEFA15FE90" box="[151,1436,298,320]" pageId="20" pageNumber="377">
<emphasis id="B926CB23FFA5AD0AE389FEAEFF2AFE90" bold="true" box="[151,163,298,319]" pageId="20" pageNumber="377">4</emphasis>
: Male second leg, mesal margin extended in short membranous process...........................................
<emphasis id="B926CB23FFA5AD0AE68EFEAEFA15FE90" bold="true" box="[1424,1436,298,319]" pageId="20" pageNumber="377">5</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA5AD0AE389FECCFA15FEF2" blockId="20.[151,1437,151,701]" box="[151,1436,327,349]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FECCFA15FEF2" box="[151,1436,327,349]" pageId="20" pageNumber="377">
- Male second leg, mesal margin not extended................................................................
<emphasis id="B926CB23FFA5AD0AE68EFECCFA15FEF2" bold="true" box="[1424,1436,328,349]" pageId="20" pageNumber="377">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A60F94FFA5AD0AE389FEE1FA15FE38" pageId="20" pageNumber="377">
<paragraph id="8BED1731FFA5AD0AE389FEE1FA15FED5" blockId="20.[151,1437,151,701]" box="[151,1436,357,379]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FEE1FA15FED5" box="[151,1436,357,379]" pageId="20" pageNumber="377">
<emphasis id="B926CB23FFA5AD0AE389FEE1FF2AFED5" bold="true" box="[151,163,357,378]" pageId="20" pageNumber="377">5</emphasis>
: Endotergum, two rows of marginal bristles which project beyond tergite margin.................................. ...
<emphasis id="B926CB23FFA5AD0AE68EFEE1FA15FED5" bold="true" box="[1424,1436,357,378]" pageId="20" pageNumber="377">6</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA5AD0AE389FE07FA15FE38" blockId="20.[151,1437,151,701]" box="[151,1436,386,408]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FE07FA15FE38" box="[151,1436,386,408]" pageId="20" pageNumber="377">
- Endotergum, two rows of marginal bristles which end clearly before the tergite margin................
<taxonomicName id="4C526CB2FFA5AD0AE7FFFE07FAC3FE37" ID-CoL="9BDJT" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[1249,1354,387,408]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFA5AD0AE7FFFE07FAC3FE37" box="[1249,1354,387,408]" italics="true" pageId="20" pageNumber="377">Z. erawani</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFA5AD0AE64EFE06FA15FE38" bold="true" box="[1360,1436,386,407]" pageId="20" pageNumber="377">
<taxonomicNameLabel id="A2157658FFA5AD0AE64EFE06FA15FE38" box="[1360,1436,386,407]" pageId="20" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A60F94FFA5AD0AE389FE1BFA15FDA2" pageId="20" pageNumber="377">
<paragraph id="8BED1731FFA5AD0AE389FE1BFA15FE7D" blockId="20.[151,1437,151,701]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FE1BFA15FE7D" pageId="20" pageNumber="377">
<emphasis id="B926CB23FFA5AD0AE389FE1BFF2AFE1B" bold="true" box="[151,163,415,436]" pageId="20" pageNumber="377">6</emphasis>
: Female vulval operculum margin straight and wide.Antennae, antennomere 6 with 90–100 cones. Male anal shield bell-shaped....................................................................... ...
<taxonomicName id="4C526CB2FFA5AD0AE75EFE39FA15FE7D" authority="Srisonchai et al., 2021" authorityName="Srisonchai" authorityYear="2021" box="[1088,1436,445,467]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="B926CB23FFA5AD0AE75EFE39FB36FE7D" box="[1088,1215,445,466]" italics="true" pageId="20" pageNumber="377">Z. golovatchi</emphasis>
<bibRefCitation id="EFC36AC0FFA5AD0AE7DBFE39FA15FE7D" author="Srisonchai, R. &amp; Sutcharit, C. &amp; Likhitrakarn, N." box="[1221,1436,445,467]" pageId="20" pageNumber="377" pagination="19 - 56" refId="ref14171" refString="Srisonchai, R., Sutcharit, C. &amp; Likhitrakarn, N. (2021) The giant pill-millipede genus Zephronia Gray, 1832 from Thailand, with a redescription of Z. siamensis Hirst, 1907 and descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae). ZooKeys, 1067, 19 - 56. https: // doi. org / 10.3897 / zookeys. 1067.72369" type="journal article" year="2021">
Srisonchai
<emphasis id="B926CB23FFA5AD0AE62EFE39FAE8FE7D" box="[1328,1377,445,466]" italics="true" pageId="20" pageNumber="377">et al.</emphasis>
, 2021
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA5AD0AE389FE5FFA15FDA2" blockId="20.[151,1437,151,701]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FE5FFA15FDA2" pageId="20" pageNumber="377">
- Female vulval operculum narrow in posterior view. Antennae, antennomere 6 with around 75 cones. Male anal shield rectangular............................................................... ..
<taxonomicName id="4C526CB2FFA5AD0AE743FE7CFA15FDA2" authority="Srisonchai et al. 2021" authorityName="Srisonchai" authorityYear="2021" box="[1117,1436,503,525]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="enghoffi">
<emphasis id="B926CB23FFA5AD0AE743FE7CFB4CFDA3" box="[1117,1221,503,525]" italics="true" pageId="20" pageNumber="377">Z. enghoffi</emphasis>
<bibRefCitation id="EFC36AC0FFA5AD0AE7D5FE73FA15FDA2" author="Srisonchai, R. &amp; Sutcharit, C. &amp; Likhitrakarn, N." box="[1227,1436,503,525]" pageId="20" pageNumber="377" pagination="19 - 56" refId="ref14171" refString="Srisonchai, R., Sutcharit, C. &amp; Likhitrakarn, N. (2021) The giant pill-millipede genus Zephronia Gray, 1832 from Thailand, with a redescription of Z. siamensis Hirst, 1907 and descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae). ZooKeys, 1067, 19 - 56. https: // doi. org / 10.3897 / zookeys. 1067.72369" type="journal article" year="2021">
Srisonchai
<emphasis id="B926CB23FFA5AD0AE628FE7CFAEEFDA3" box="[1334,1383,503,525]" italics="true" pageId="20" pageNumber="377">et al.</emphasis>
2021
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A60F94FFA5AD0AE389FD91FA15FDE8" pageId="20" pageNumber="377">
<paragraph id="8BED1731FFA5AD0AE389FD91FA15FD85" blockId="20.[151,1437,151,701]" box="[151,1436,533,555]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FD91FA15FD85" box="[151,1436,533,555]" pageId="20" pageNumber="377">
<emphasis id="B926CB23FFA5AD0AE389FD91FF2AFD85" bold="true" box="[151,163,533,554]" pageId="20" pageNumber="377">7</emphasis>
: Brown. Inner area of endotergum without setae.........
<taxonomicName id="4C526CB2FFA5AD0AE007FD91FB13FD84" authority="Likhitrakarn &amp; Golovatch" authorityName="Likhitrakarn &amp; Golovatch" authorityYear="2021" box="[793,1178,533,555]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="lannaensis">
<emphasis id="B926CB23FFA5AD0AE007FD91FC10FD85" box="[793,921,533,554]" italics="true" pageId="20" pageNumber="377">Z. lannaensis</emphasis>
Likhitrakarn &amp; Golovatch
</taxonomicName>
in
<bibRefCitation id="EFC36AC0FFA5AD0AE7A6FD91FA15FD85" author="Likhitrakarn, N. &amp; Golovatch, S. I. &amp; Srisonchai, R. &amp; Sutcharit, C." box="[1208,1436,533,555]" pageId="20" pageNumber="377" pagination="12 - 26" refId="ref13325" refString="Likhitrakarn, N., Golovatch, S. I., Srisonchai, R. &amp; Sutcharit, C. (2021) Two New Species of the Giant Pill-Millipede Genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae). Tropical Natural History, 21, 12 - 26." type="journal article" year="2021">
Likhitrakarn
<emphasis id="B926CB23FFA5AD0AE628FD91FAEEFD85" box="[1334,1383,533,554]" italics="true" pageId="20" pageNumber="377">et al.</emphasis>
2021
</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA5AD0AE389FDB7FA15FDE8" blockId="20.[151,1437,151,701]" box="[151,1436,562,584]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FDB7FA15FDE8" box="[151,1436,562,584]" pageId="20" pageNumber="377">
- Greyish. Inner area of endotergum with setae...............................................................
<emphasis id="B926CB23FFA5AD0AE68EFDB6FA15FDE8" bold="true" box="[1424,1436,562,583]" pageId="20" pageNumber="377">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A60F94FFA5AD0AE389FDCBFA15FD12" pageId="20" pageNumber="377">
<paragraph id="8BED1731FFA5AD0AE389FDCBFA15FD2D" blockId="20.[151,1437,151,701]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FDCBFA15FD2D" pageId="20" pageNumber="377">
<emphasis id="B926CB23FFA5AD0AE389FDCBFF2AFDCB" bold="true" box="[151,163,591,612]" pageId="20" pageNumber="377">8</emphasis>
: Small,
<quantity id="4CAABAD4FFA5AD0AE23BFDD4FE02FDCA" box="[293,395,592,613]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="20" pageNumber="377" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">20–22 mm</quantity>
long. Legs with 7–9 ventral spines. Male antennae, antennomere 6 with ca. 50 cones. Pleurite 1 extended into long, sharp tip. Endotergum with two rows of marginal bristles........................
<taxonomicName id="4C526CB2FFA5AD0AE777FDE9FA15FD2D" authority="Srisonchai et al. 2021" authorityName="Srisonchai" authorityYear="2021" box="[1129,1436,621,643]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFA5AD0AE777FDE9FB4CFD2D" box="[1129,1221,621,642]" italics="true" pageId="20" pageNumber="377">Z. panhai</emphasis>
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Srisonchai
<emphasis id="B926CB23FFA5AD0AE628FDE9FAEEFD2D" box="[1334,1383,621,642]" italics="true" pageId="20" pageNumber="377">et al.</emphasis>
2021
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</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BED1731FFA5AD0AE389FD0FFA15FD12" blockId="20.[151,1437,151,701]" pageId="20" pageNumber="377">
<keyLead id="30A3B404FFA5AD0AE389FD0FFA15FD12" pageId="20" pageNumber="377">
- Medium,
<quantity id="4CAABAD4FFA5AD0AE25FFD0EFE0EFD0F" box="[321,391,650,672]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="377" unit="mm" value="40.0">40 mm</quantity>
long. Legs with 10–12 ventral spines. Male antennae, antennomere 6 with 78–88 apical cones. Pleurite 1 weakly extended, well-rounded. Endotergum with three rows of marginal bristles...............
<taxonomicName id="4C526CB2FFA5AD0AE7AEFD2CFAC3FD13" ID-CoL="9BDJV" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[1200,1354,679,701]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="377" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFA5AD0AE7AEFD2CFAC3FD13" box="[1200,1354,679,701]" italics="true" pageId="20" pageNumber="377">Z. chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFA5AD0AE64EFD2CFA15FD12" bold="true" box="[1360,1436,680,701]" pageId="20" pageNumber="377">
<taxonomicNameLabel id="A2157658FFA5AD0AE64EFD2CFA15FD12" box="[1360,1436,680,701]" pageId="20" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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