271 lines
30 KiB
XML
271 lines
30 KiB
XML
<document id="218A3DA3CDF24099104A08CF7B65BEA5" ID-DOI="10.1649/0010-065X-70.4.825" ID-Zenodo-Dep="5340413" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630186311507" checkinUser="diego" docAuthor="Hisamatsu, Sadatomo, Bayless, Victoria M. & Carlton, Christopher E." docDate="2016" docId="F042A324FFFFFF8E08E0ECBFFC788F71" docLanguage="en" docName="0010-065X-70.4.825.pdf" docOrigin="The Coleopterists Bulletin 70 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-70.4.825" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Cyclocaccus intermediatus Hisamatsu 2016, new species" docType="treatment" docVersion="3" lastPageNumber="870" masterDocId="0C7BDB5CFFD3FFA30A01E84CFFFD8C1E" masterDocTitle="Revision ofCyclocaccusSharp (Coleoptera: Nitidulidae: Nitidulinae)" masterLastPageNumber="870" masterPageNumber="825" pageNumber="825" updateTime="1699179913154" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F616193B9E2CE09A59403F0B105177ED">Revision ofCyclocaccusSharp (Coleoptera: Nitidulidae: Nitidulinae)</mods:title>
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<mods:roleTerm id="5E51C84C75C26EC27F7F3EE1570C34E8">Author</mods:roleTerm>
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<mods:namePart id="4C84AA1D07ED0054248397431AA657B3">Hisamatsu, Sadatomo</mods:namePart>
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<mods:roleTerm id="0554B63EE00EFCCF44615AFB4722E1C4">Author</mods:roleTerm>
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<mods:namePart id="61F31D519DD745E1C7D13482467B637E">Bayless, Victoria M.</mods:namePart>
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<mods:name id="7AD55366FFF40090741F991AA51E07C6" type="personal">
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<mods:roleTerm id="303F287C0CCEB2B13ADE3B403434501E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DFBD89A5B1704BC309AFC112DBFA3B89">Carlton, Christopher E.</mods:namePart>
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<mods:typeOfResource id="3E8F6473341A0AC707CAFDF3F7D69946">text</mods:typeOfResource>
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<mods:title id="6D3D4DCE1C05B834A18E5F5DBC898556">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="14AD2F850D94091687902BD4A2A42733">
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<mods:date id="58720E881AF3A42E33B8DBA971739015">2016</mods:date>
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<mods:number id="287E939A06F100A74226EE3D25B12556">2016-12-31</mods:number>
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<mods:number id="0A31354A698AE29C01F854075C74FA6A">70</mods:number>
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<mods:number id="04015424A0EB164BD8D3DE533106F184">4</mods:number>
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<mods:url id="634A70A872E794A42FF0BB3539A56353">http://dx.doi.org/10.1649/0010-065x-70.4.825</mods:url>
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<mods:classification id="863A8D40EAF4FC4A9D18EE2EC4578E86">journal article</mods:classification>
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<mods:identifier id="049105E0BD4ABF89A3EA6C0F9E4127C5" type="DOI">10.1649/0010-065X-70.4.825</mods:identifier>
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<treatment id="F042A324FFFFFF8E08E0ECBFFC788F71" LSID="urn:lsid:plazi:treatment:F042A324FFFFFF8E08E0ECBFFC788F71" httpUri="http://treatment.plazi.org/id/F042A324FFFFFF8E08E0ECBFFC788F71" lastPageId="45" pageId="44">
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<subSubSection id="30F141B9FFFFFF8F08E0ECBFFC26893D" pageId="44" type="nomenclature">
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<paragraph id="78541232FFFFFF8F08E0ECBFFC26893D" blockId="44.[737,1119,1267,1341]" pageId="44">
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<heading id="231CA55EFFFFFF8F08E0ECBFFC26893D" centered="true" fontSize="10" level="2" pageId="44" reason="2">
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<emphasis id="4A9FCE20FFFFFF8F08E0ECBFFC26893D" bold="true" pageId="44">
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<taxonomicName id="BFEB69B1FFFFFF8F08E0ECBFFBA68916" authority="Hisamatsu" authorityName="Hisamatsu" authorityYear="2016" box="[737,1115,1267,1288]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" phylum="Arthropoda" rank="species" species="intermediatus" status="sp. nov.">
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<emphasis id="4A9FCE20FFFFFF8F08E0ECBFFC1A8916" bold="true" box="[737,999,1267,1288]" italics="true" pageId="44">Cyclocaccus intermediatus</emphasis>
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Hisamatsu
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</taxonomicName>
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,
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<taxonomicNameLabel id="51AC735BFFFFFF8F0967ED42FC26893D" box="[870,987,1294,1315]" pageId="44" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="30F141B9FFFFFF8F0917ED64FBD68923" box="[790,1067,1320,1341]" pageId="44" type="description">
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<paragraph id="78541232FFFFFF8F0917ED64FBD68923" blockId="44.[737,1119,1267,1341]" box="[790,1067,1320,1341]" pageId="44">
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(
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<figureCitation id="E0D00EB7FFFFFF8F091DED64FC838923" box="[796,894,1320,1341]" captionStart="Fig" captionStartId="34.[175,206,993,1012]" captionTargetBox="[175,1154,190,946]" captionTargetId="figure-280@34.[151,1171,183,970]" captionTargetPageId="34" captionText="Fig. 15. Dorsal habitus of Cyclocaccus species in the C. morulus species-group. A) C. lescheni, holotype, B) C. stonyx, holotype, Costa Rica, C) C. stonyx, Panama, D) C. stonyx, Ecuador, E) C. morulus, holotype, F) C. smileyeyes, holotype, Bolivia, G) C. intermediatus, holotype. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340444" httpUri="https://zenodo.org/record/5340444/files/figure.png" pageId="44">Figs. 15G</figureCitation>
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,
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<figureCitation id="E0D00EB7FFFFFF8F0988ED64FC498923" box="[905,948,1320,1341]" captionStart="Fig" captionStartId="35.[197,228,1565,1584]" captionTargetBox="[251,1117,192,1529]" captionTargetId="figure-0@35.[233,1133,183,1542]" captionTargetPageId="35" captionText="Fig. 16. Diagnostic characters of the C. morulus species-group. Cyclocaccus morulus: A) Ventral habitus, B) Head, dorsal view, C) Head, ventral view, D) Labrum, E) Mentum and labium, ventral view, F) Right maxilla, dorsal view, G) Right mandible, dorsal view, H) Scutellum, I) Male abdominal tergite VII, J) Male abdominal sternite VII. Cyclocaccus intermediatus: K) Male abdominal tergite VII. Cyclocaccus morulus: L) Metendosternite, dorsal view, M) Male protibia and protarsus, dorsal view, N) Male mesotibia and mesotarsus, dorsal view, O) Male metatibia and metatarsus, dorsal view, P) Male abdominal tergite VIII and spiculum gastrale, ventral view, Q) Left metanepisternum. Scale bars: A = 1 mm; B–C, I–K, P–Q = 0.5 mm; D–H, L–O = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340446" httpUri="https://zenodo.org/record/5340446/files/figure.png" pageId="44">16K</figureCitation>
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,
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<figureCitation id="E0D00EB7FFFFFF8F09C1ED64FC278923" box="[960,986,1320,1341]" captionStart="Fig" captionStartId="40.[175,206,1686,1705]" captionTargetBox="[203,1125,226,1630]" captionTargetId="figure-1@40.[172,1152,183,1662]" captionTargetPageId="40" captionText="Fig. 21. Cyclocaccus intermediatus, Peru. A) Tegmen, holotype, ventral view, B) Median lobe, holotype, ventral view, C) Right antenna, holotype, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340458" httpUri="https://zenodo.org/record/5340458/files/figure.png" pageId="44">21</figureCitation>
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, Map 5)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="30F141B9FFFFFF8F08C4ED1FFC328A49" pageId="44" type="materials_examined">
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<paragraph id="78541232FFFFFF8F08C4ED1FFC328A49" blockId="44.[685,1172,1362,1730]" pageId="44">
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<emphasis id="4A9FCE20FFFFFF8F08C4ED1FFCB58976" bold="true" box="[709,840,1363,1384]" pageId="44">Type Series.</emphasis>
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<materialsCitation id="C883186FFFFFFF8F0955ED1FFC2A89F3" collectingDate="1989-07-02" collectionCode="SEMC" collectorName="Plot & J. S. Ashe & R. A. Leschen" county="Tambopata Prov." elevation="200" latitude="-12.55" location="Reserva" longLatPrecision="1293" longitude="-69.05" municipality="Maldonado" pageId="44" specimenCount="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="holotype">
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||
<typeStatus id="A750AC90FFFFFF8F0955ED1FFC518976" box="[852,940,1363,1384]" pageId="44" type="holotype">Holotype</typeStatus>
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||
(
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<collectionCode id="1EFA8AF7FFFFFF8F09C3ED1FFBF58976" box="[962,1032,1363,1384]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="44" type="Museum">SEMC</collectionCode>
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):
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||
<specimenCount id="6EEDD9BBFFFFFF8F0E20ED1EFBCE8976" box="[1057,1075,1362,1384]" pageId="44" type="male">♂</specimenCount>
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, ‘PERU:
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<collectingCounty id="91356ABEFFFFFF8F08ACED22FCA8899D" box="[685,853,1390,1411]" pageId="44">Tambopata Prov.</collectingCounty>
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/
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<collectingRegion id="BA2FDCD0FFFFFF8F096AED22FC03899D" box="[875,1022,1390,1411]" country="Peru" name="Madre de Dios" pageId="44">Madre de Dios</collectingRegion>
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Dpto. /
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<locationDeviation id="69E60B10FFFFFF8F0E55ED22FCE28983" location="Puerto" pageId="44">
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<quantity id="BF13BFD7FFFFFF8F0E55ED22FB6E899D" box="[1108,1171,1390,1411]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="44" unit="km" value="15.0">15 km</quantity>
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||
NE Puerto
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||
</locationDeviation>
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||
//
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||
<collectingMunicipality id="98308848FFFFFF8F0954EDC4FC398983" box="[853,964,1416,1437]" pageId="44">Maldonado</collectingMunicipality>
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,
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<location id="7D3444E9FFFFFF8F09DAEDC4FBDA8983" LSID="urn:lsid:plazi:treatment:F042A324FFFFFF8E08E0ECBFFC788F71:7D3444E9FFFFFF8F09DAEDC4FBDA8983" box="[987,1063,1416,1437]" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Maldonado" name="Reserva" pageId="44" stateProvince="Madre de Dios">Reserva</location>
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||
/
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||
<collectingRegion id="BA2FDCD0FFFFFF8F0E56EDC4FB698983" box="[1111,1172,1416,1437]" country="Peru" name="Cusco" pageId="44">Cuzco</collectingRegion>
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<location id="7D3444E9FFFFFF8F08ACEDEFFCE589A6" LSID="urn:lsid:plazi:treatment:F042A324FFFFFF8E08E0ECBFFC788F71:7D3444E9FFFFFF8F08ACEDEFFCE589A6" box="[685,792,1443,1464]" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Maldonado" name="Amazonico" pageId="44" stateProvince="Madre de Dios">Amazónico</location>
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/
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||
<geoCoordinate id="1DDF74F5FFFFFF8F093DEDEFFC7789A6" box="[828,906,1442,1464]" degrees="12" direction="south" minutes="33" orientation="latitude" pageId="44" precision="925" value="-12.55">12°33′S</geoCoordinate>
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,
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||
<geoCoordinate id="1DDF74F5FFFFFF8F099DEDEFFC0C89A6" box="[924,1009,1442,1464]" degrees="69" direction="west" minutes="03" orientation="longitude" pageId="44" precision="925" value="-69.05">69°03′W</geoCoordinate>
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/
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<quantity id="BF13BFD7FFFFFF8F0E14EDEFFBA189A6" box="[1045,1116,1443,1464]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="44" unit="m" value="200.0">
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<elevation id="F3C6F501FFFFFF8F0E14EDEFFBA189A6" box="[1045,1116,1443,1464]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="44" unit="m" value="200.0">200 m</elevation>
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</quantity>
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||
,
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||
<collectorName id="D51E77E4FFFFFF8F0E6FEDEFFB6989A6" box="[1134,1172,1443,1464]" pageId="44">Plot</collectorName>
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#Z2U19 //
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||
<date id="0C5534F2FFFFFF8F0917EDF1FC7A89CC" box="[790,903,1469,1490]" pageId="44" value="1989-07-02">
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||
<collectingDate id="1C11CD1AFFFFFF8F0917EDF1FC7A89CC" box="[790,903,1469,1490]" pageId="44" value="1989-07-02">2 July 1989</collectingDate>
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</date>
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||
,
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||
<collectorName id="D51E77E4FFFFFF8F0993EDF1FC0989CC" box="[914,1012,1469,1490]" pageId="44">J. S. Ashe</collectorName>
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||
, /
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||
<collectorName id="D51E77E4FFFFFF8F0E0DEDF1FB6989CC" box="[1036,1172,1469,1490]" pageId="44">R. A. Leschen</collectorName>
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||
#366 / ex., flight
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||
<collectingMethod id="A1AA6A25FFFFFF8F0954ED94FC5B89F3" box="[853,934,1496,1517]" pageId="44">intercept</collectingMethod>
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||
trap’
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||
</materialsCitation>
|
||
.
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||
<materialsCitation id="C883186FFFFFFF8F09E7ED94FC318A49" collectingDate="2000-10-20" collectingDateMax="2000-10-21" collectingDateMin="2000-10-20" collectionCode="SEMC" collectorName="R. Brooks" country="Peru" county="Cocha Salvador" elevation="310" latitude="-12.003612" location="Manu National Park" longLatPrecision="21" longitude="-71.52667" municipality="Reserved Zone" pageId="44" specimenCount="1" specimenCount-female="1" stateProvince="Marde de Dios" typeStatus="paratype">
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||
<typeStatus id="A750AC90FFFFFF8F09E7ED94FBC789F3" box="[998,1082,1496,1517]" pageId="44" type="paratype">Paratype</typeStatus>
|
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: [
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||
<emphasis id="4A9FCE20FFFFFF8F0E4DED94FB7189F3" bold="true" box="[1100,1164,1496,1517]" pageId="44">
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||
<collectingCountry id="00FC52A2FFFFFF8F0E4DED94FB7189F3" box="[1100,1164,1496,1517]" name="Peru" pageId="44">PERU</collectingCountry>
|
||
</emphasis>
|
||
]
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||
<specimenCount id="6EEDD9BBFFFFFF8F08ACEDBEFD368A16" box="[685,715,1522,1544]" pageId="44" type="female">1♀</specimenCount>
|
||
,
|
||
<collectingCounty id="91356ABEFFFFFF8F08E4EDBEFC708A19" box="[741,909,1522,1543]" pageId="44">Cocha Salvador</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="98308848FFFFFF8F09A5EDBEFBB98A19" box="[932,1092,1522,1543]" pageId="44">Reserved Zone</collectingMunicipality>
|
||
,
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||
<location id="7D3444E9FFFFFF8F0E5DEDBEFCCA8A3C" LSID="urn:lsid:plazi:treatment:F042A324FFFFFF8E08E0ECBFFC788F71:7D3444E9FFFFFF8F0E5DEDBEFCCA8A3C" country="Peru" county="Cocha Salvador" latitude="-12.003612" longLatPrecision="21" longitude="-71.52667" municipality="Reserved Zone" name="Manu National Park" pageId="44" stateProvince="Marde de Dios">Manu National Park</location>
|
||
,
|
||
<collectingRegion id="BA2FDCD0FFFFFF8F0945EE41FC268A3C" box="[836,987,1549,1570]" inferredBySuffix="true" pageId="44">Marde de Dios</collectingRegion>
|
||
,
|
||
<quantity id="BF13BFD7FFFFFF8F09E8EE41FBD18A3C" box="[1001,1068,1549,1570]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="44" unit="m" value="310.0">
|
||
<elevation id="F3C6F501FFFFFF8F09E8EE41FBD18A3C" box="[1001,1068,1549,1570]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="44" unit="m" value="310.0">310 m</elevation>
|
||
</quantity>
|
||
, ex:
|
||
<collectingMethod id="A1AA6A25FFFFFF8F0E61EE41FCD68A23" pageId="44">flight intercept trap</collectingMethod>
|
||
,
|
||
<geoCoordinate id="1DDF74F5FFFFFF8F0937EE64FC5B8A23" box="[822,934,1575,1597]" degrees="12" direction="south" minutes="00" orientation="latitude" pageId="44" precision="15" seconds="13" value="-12.003612">12°00′13”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1DDF74F5FFFFFF8F09B0EE64FBD78A23" box="[945,1066,1575,1597]" degrees="71" direction="west" minutes="31" orientation="longitude" pageId="44" precision="15" seconds="36" value="-71.52667">71°31′36″W</geoCoordinate>
|
||
,
|
||
<date id="0C5534F2FFFFFF8F0E35EE64FD238A49" pageId="44" value="2000-10-20" valueMax="2000-10-21" valueMin="2000-10-20">
|
||
<collectingDate id="1C11CD1AFFFFFF8F0E35EE64FD238A49" pageId="44" value="2000-10-20" valueMax="2000-10-21" valueMin="2000-10-20">20–21. X. 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="D51E77E4FFFFFF8F08E8EE0EFCB78A49" box="[745,842,1602,1623]" pageId="44">R. Brooks</collectorName>
|
||
leg. (
|
||
<collectionCode id="1EFA8AF7FFFFFF8F0981EE0EFC388A49" box="[896,965,1602,1623]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="44" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30F141B9FFFFFF8F08C4EE11FBF68ADF" pageId="44" type="etymology">
|
||
<paragraph id="78541232FFFFFF8F08C4EE11FBF68ADF" blockId="44.[685,1172,1362,1730]" pageId="44">
|
||
<emphasis id="4A9FCE20FFFFFF8F08C4EE11FCC58A6C" bold="true" box="[709,824,1629,1650]" pageId="44">Etymology.</emphasis>
|
||
The epithet of this new species is derived from the somewhat intermediate character between congeners within the
|
||
<taxonomicName id="BFEB69B1FFFFFF8F09CDEEDDFBBF8AB9" authorityName="Sharp" authorityYear="1891" box="[972,1090,1681,1703]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" phylum="Arthropoda" rank="species" species="laeticulus">
|
||
<emphasis id="4A9FCE20FFFFFF8F09CDEEDDFBBF8AB9" box="[972,1090,1681,1703]" italics="true" pageId="44">C. laeticulus</emphasis>
|
||
</taxonomicName>
|
||
speciesgroup and
|
||
<taxonomicName id="BFEB69B1FFFFFF8F0915EEE0FC818ADC" authorityName="Hisamatsu" authorityYear="2016" box="[788,892,1708,1730]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" phylum="Arthropoda" rank="species" species="morulus">
|
||
<emphasis id="4A9FCE20FFFFFF8F0915EEE0FC818ADC" box="[788,892,1708,1730]" italics="true" pageId="44">C. morulus</emphasis>
|
||
</taxonomicName>
|
||
species-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30F141B9FFFEFF8E0AC4E88EFE8D8F1B" pageId="45" type="diagnosis">
|
||
<paragraph id="78541232FFFEFF8E0AC4E88EFD7A8E08" blockId="45.[173,660,194,1729]" pageId="45">
|
||
<emphasis id="4A9FCE20FFFEFF8E0AC4E88EFED18CC9" bold="true" box="[197,300,194,215]" pageId="45">Diagnosis.</emphasis>
|
||
This species differs from other congeners by the following combination of characters: body dark reddish brown; apical margin of abdominal tergite VII widely rounded; metathoracic discrimen absent; subcoxal lines on abdominal sternite III absent; pro- and mesotarsal segments II and III of males moderately dilated; tegmen long, subparallel-sided at basal half, then strongly convergent apically; TL/TW = 2.32 (
|
||
<emphasis id="4A9FCE20FFFEFF8E0804E9DBFDEC8DB2" box="[517,529,407,428]" italics="true" pageId="45">n</emphasis>
|
||
= 1); apical margin of tegmen narrowly rounded in ventral view; median lobe subparallel-sided; apical margin of median lobe truncate; apices of gonocoxites square, with apicolateral corners widely rounded.
|
||
</paragraph>
|
||
<paragraph id="78541232FFFEFF8E0AC4EA57FE8D8F1B" blockId="45.[173,660,194,1729]" pageId="45">
|
||
<emphasis id="4A9FCE20FFFEFF8E0AC4EA57FE9B8E2E" bold="true" box="[197,358,539,560]" pageId="45">Similar Species.</emphasis>
|
||
This new species is similar to
|
||
<taxonomicName id="BFEB69B1FFFEFF8E0AACEA79FEE78E55" authorityName="Hisamatsu" authorityYear="2016" box="[173,282,565,587]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" phylum="Arthropoda" rank="species" species="morulus">
|
||
<emphasis id="4A9FCE20FFFEFF8E0AACEA79FEE78E55" box="[173,282,565,587]" italics="true" pageId="45">C. morulus</emphasis>
|
||
</taxonomicName>
|
||
in having a dark reddish brown body coloration, but can be distinguished by the following combination of characters: apical margin of abdominal tergite VII widely rounded; metathoracic discrimen absent; metanepisternum densely punctate; abdominal sternite III without subcoxal lines; tegmen strongly convergent apically from basal half in ventral view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30F141B9FFFEFF8E0AC4EB46FBC48DFF" pageId="45" type="description">
|
||
<paragraph id="78541232FFFEFF8E0AC4EB46FB578D42" blockId="45.[173,660,194,1729]" lastBlockId="45.[706,1194,194,879]" pageId="45">
|
||
<emphasis id="4A9FCE20FFFEFF8E0AC4EB46FEC28F01" bold="true" box="[197,319,778,799]" pageId="45">Description.</emphasis>
|
||
Length 1.88–2.03 (1.95) mm, width 1.53–1.70 (1.62) mm, depth 0.92–1.02 (0.97) mm (
|
||
<emphasis id="4A9FCE20FFFEFF8E0B24EB0CFECC8F4B" box="[293,305,832,853]" italics="true" pageId="45">n</emphasis>
|
||
= 2).
|
||
<emphasis id="4A9FCE20FFFEFF8E0B72EB0CFE508F4B" bold="true" box="[371,429,832,853]" pageId="45">Male.</emphasis>
|
||
Body (
|
||
<figureCitation id="E0D00EB7FFFEFF8E0BFBEB0CFDAE8F4B" box="[506,595,832,853]" captionStart="Fig" captionStartId="34.[175,206,993,1012]" captionTargetBox="[175,1154,190,946]" captionTargetId="figure-280@34.[151,1171,183,970]" captionTargetPageId="34" captionText="Fig. 15. Dorsal habitus of Cyclocaccus species in the C. morulus species-group. A) C. lescheni, holotype, B) C. stonyx, holotype, Costa Rica, C) C. stonyx, Panama, D) C. stonyx, Ecuador, E) C. morulus, holotype, F) C. smileyeyes, holotype, Bolivia, G) C. intermediatus, holotype. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340444" httpUri="https://zenodo.org/record/5340444/files/figure.png" pageId="45">Fig. 15G</figureCitation>
|
||
) oval; dorsal disc dark reddish brown; apical half of head, mouthparts, antennal segments I–VIII, lateral margins of pronotum, venter of head, hypomera, elytral epipleura, and legs paler.
|
||
<emphasis id="4A9FCE20FFFEFF8E0BE5EBE6FDDD8FA1" bold="true" box="[484,544,938,959]" pageId="45">Head:</emphasis>
|
||
Dorsal disc densely punctate; interspaces reticulate. Labrum feebly notched at middle. Antenna (
|
||
<figureCitation id="E0D00EB7FFFEFF8E0804EB93FDA78FEA" box="[517,602,991,1012]" captionStart="Fig" captionStartId="40.[175,206,1686,1705]" captionTargetBox="[203,1125,226,1630]" captionTargetId="figure-1@40.[172,1152,183,1662]" captionTargetPageId="40" captionText="Fig. 21. Cyclocaccus intermediatus, Peru. A) Tegmen, holotype, ventral view, B) Median lobe, holotype, ventral view, C) Right antenna, holotype, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340458" httpUri="https://zenodo.org/record/5340458/files/figure.png" pageId="45">Fig. 21C</figureCitation>
|
||
) with club long and large, ACL/ASI-VIII = 0.77 (
|
||
<emphasis id="4A9FCE20FFFEFF8E0850EBB6FDA08811" box="[593,605,1018,1039]" italics="true" pageId="45">n</emphasis>
|
||
= 1); antennal segment XI with 2 oval antennal sensillae; approximate ratio of each segment (
|
||
<emphasis id="4A9FCE20FFFEFF8E0814EC63FDDC885A" box="[533,545,1071,1092]" italics="true" pageId="45">n</emphasis>
|
||
= 1) 4.23: 2.70: 2.30: 1.37: 1.73: 1.20: 1.00: 1.55: 3.00: 3.31: 6.27.
|
||
<emphasis id="4A9FCE20FFFEFF8E0AE6EC28FEAE8867" bold="true" box="[231,339,1124,1145]" pageId="45">Pronotum:</emphasis>
|
||
3.24–3.28 (3.26) times as wide as long (
|
||
<emphasis id="4A9FCE20FFFEFF8E0B07EC33FEEF888A" box="[262,274,1151,1172]" italics="true" pageId="45">n</emphasis>
|
||
= 2); disc densely punctate; disc with 2 kinds of punctures, large punctures, slightly smaller than eye-facet, and minute punctures; interspaces reticulate. Scutellum with disc densely punctate; punctures on disc similar to those of pronotum; interspaces reticulate.
|
||
<emphasis id="4A9FCE20FFFEFF8E0BE6ED4FFDD08906" bold="true" box="[487,557,1283,1304]" pageId="45">Elytra:</emphasis>
|
||
0.93–0.96 (0.94) times as long as wide (
|
||
<emphasis id="4A9FCE20FFFEFF8E0BE1ED52FE11892D" box="[480,492,1310,1331]" italics="true" pageId="45">n</emphasis>
|
||
= 2), 3.54–3.61 (3.58) times as long as pronotum (
|
||
<emphasis id="4A9FCE20FFFEFF8E083AED75FDBA8950" box="[571,583,1337,1358]" italics="true" pageId="45">n</emphasis>
|
||
= 2), subparallel-sided from base to basal 1/3; each elytron with 9 regular rows of punctures about as large as eye facet; punctures on interstices similar to those on pronotum; abdominal tergite VII with apical margin widely rounded.
|
||
<emphasis id="4A9FCE20FFFEFF8E0BD7EDF1FDDD89CC" bold="true" box="[470,544,1469,1490]" pageId="45">Venter:</emphasis>
|
||
Prosternum strongly convex along midline, with a tuft of setae at middle. Metaventrite strongly convex, with axillary space; metathoracic discrimen absent. Metanepisternum densely punctate. Abdominal sternite III without subcoxal lines; lateral margins of abdominal sternite VII serrate.
|
||
<emphasis id="4A9FCE20FFFEFF8E0802EE11FDC48A6C" bold="true" box="[515,569,1629,1650]" pageId="45">Legs:</emphasis>
|
||
Slender; pro- and mesotarsal segments I–III moderately dilated; tarsal claws simple.
|
||
<emphasis id="4A9FCE20FFFEFF8E0BD5EEDEFDC58AB9" bold="true" box="[468,568,1682,1703]" pageId="45">Genitalia:</emphasis>
|
||
Tegmen (
|
||
<figureCitation id="E0D00EB7FFFEFF8E0AB5EEE0FEEE8ADF" box="[180,275,1708,1729]" captionStart="Fig" captionStartId="40.[175,206,1686,1705]" captionTargetBox="[203,1125,226,1630]" captionTargetId="figure-1@40.[172,1152,183,1662]" captionTargetPageId="40" captionText="Fig. 21. Cyclocaccus intermediatus, Peru. A) Tegmen, holotype, ventral view, B) Median lobe, holotype, ventral view, C) Right antenna, holotype, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340458" httpUri="https://zenodo.org/record/5340458/files/figure.png" pageId="45">Fig. 21A</figureCitation>
|
||
) subparallel-sided at basal 1/2, then strongly convergent apically; TL/TW = 2.32 (
|
||
<emphasis id="4A9FCE20FFFEFF8E0E6BE88EFB8B8CC9" box="[1130,1142,194,215]" italics="true" pageId="45">n</emphasis>
|
||
= 1); apical margin of tegmen narrowly rounded in ventral view; median lobe (
|
||
<figureCitation id="E0D00EB7FFFEFF8E09C7E8BBFBDE8D12" box="[966,1059,247,268]" captionStart="Fig" captionStartId="40.[175,206,1686,1705]" captionTargetBox="[203,1125,226,1630]" captionTargetId="figure-1@40.[172,1152,183,1662]" captionTargetPageId="40" captionText="Fig. 21. Cyclocaccus intermediatus, Peru. A) Tegmen, holotype, ventral view, B) Median lobe, holotype, ventral view, C) Right antenna, holotype, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340458" httpUri="https://zenodo.org/record/5340458/files/figure.png" pageId="45">Fig. 21B</figureCitation>
|
||
) subparallelsided in ventral view; apical margin of median lobe truncate, shallowly emarginate at middle in ventral view; sclerites of internal sac as in
|
||
<figureCitation id="E0D00EB7FFFEFF8E0E51E90BFB5B8D42" box="[1104,1190,327,348]" captionStart="Fig" captionStartId="40.[175,206,1686,1705]" captionTargetBox="[203,1125,226,1630]" captionTargetId="figure-1@40.[172,1152,183,1662]" captionTargetPageId="40" captionText="Fig. 21. Cyclocaccus intermediatus, Peru. A) Tegmen, holotype, ventral view, B) Median lobe, holotype, ventral view, C) Right antenna, holotype, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340458" httpUri="https://zenodo.org/record/5340458/files/figure.png" pageId="45">Fig. 21B</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="78541232FFFEFF8E08C2E92DFBC48DFF" blockId="45.[706,1194,194,879]" pageId="45">
|
||
<emphasis id="4A9FCE20FFFEFF8E08C2E92DFCED8D68" bold="true" box="[707,784,353,374]" pageId="45">Female.</emphasis>
|
||
Prosternum without tuft of setae at middle. Pro- and mesotarsal segments I–III more weakly dilated than those of males. Ovipositor (
|
||
<figureCitation id="E0D00EB7FFFEFF8E0E4BE9DBFB5F8DB2" box="[1098,1186,407,428]" captionStart="Fig" captionStartId="40.[175,206,1686,1705]" captionTargetBox="[203,1125,226,1630]" captionTargetId="figure-1@40.[172,1152,183,1662]" captionTargetPageId="40" captionText="Fig. 21. Cyclocaccus intermediatus, Peru. A) Tegmen, holotype, ventral view, B) Median lobe, holotype, ventral view, C) Right antenna, holotype, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340458" httpUri="https://zenodo.org/record/5340458/files/figure.png" pageId="45">Fig. 21D</figureCitation>
|
||
) with squared apices of gonocoxites; apicolateral corners of gonocoxites widely rounded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30F141B9FFFEFF8E08DAE9AAFC128DE5" box="[731,1007,486,507]" pageId="45" type="distribution">
|
||
<paragraph id="78541232FFFEFF8E08DAE9AAFC128DE5" blockId="45.[706,1194,194,879]" box="[731,1007,486,507]" pageId="45">
|
||
<emphasis id="4A9FCE20FFFEFF8E08DAE9AAFCA18DE5" bold="true" box="[731,860,486,507]" pageId="45">Distribution.</emphasis>
|
||
<collectingCountry id="00FC52A2FFFEFF8E0969E9AAFC6E8DE5" box="[872,915,486,507]" name="Peru" pageId="45">Peru</collectingCountry>
|
||
(Map 5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30F141B9FFFEFF8E08DAEA4DFC788F71" pageId="45" type="discussion">
|
||
<paragraph id="78541232FFFEFF8E08DAEA4DFC788F71" blockId="45.[706,1194,194,879]" pageId="45">
|
||
<emphasis id="4A9FCE20FFFEFF8E08DAEA4DFCB08E08" bold="true" box="[731,845,513,534]" pageId="45">Comments.</emphasis>
|
||
<taxonomicName id="BFEB69B1FFFEFF8E095BEA4CFBB28E08" authorityName="Hisamatsu" authorityYear="2016" box="[858,1103,512,534]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" phylum="Arthropoda" rank="species" species="intermediatus">
|
||
<emphasis id="4A9FCE20FFFEFF8E095BEA4CFBB28E08" box="[858,1103,512,534]" italics="true" pageId="45">Cyclocaccus intermediatus</emphasis>
|
||
</taxonomicName>
|
||
has some characters of the
|
||
<taxonomicName id="BFEB69B1FFFEFF8E0977EA57FC098E2F" authorityName="Sharp" authorityYear="1891" box="[886,1012,539,561]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" phylum="Arthropoda" rank="species" species="laeticulus">
|
||
<emphasis id="4A9FCE20FFFEFF8E0977EA57FC098E2F" box="[886,1012,539,561]" italics="true" pageId="45">C. laeticulus</emphasis>
|
||
</taxonomicName>
|
||
species-group, as follows: apical margin of abdominal tergite VII widely rounded; subcoxal lines on abdominal sternite III absent; pro- and mesotarsal segments II and III of males moderately dilated; and tegmen narrowing to apically in ventral view. However, it differs in the following combination of characters that constitute the basis for including it in the
|
||
<taxonomicName id="BFEB69B1FFFEFF8E08C3EAA3FCCB8F1B" authorityName="Hisamatsu" authorityYear="2016" box="[706,822,751,773]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" phylum="Arthropoda" rank="species" species="morulus">
|
||
<emphasis id="4A9FCE20FFFEFF8E08C3EAA3FCCB8F1B" box="[706,822,751,773]" italics="true" pageId="45">C. morulus</emphasis>
|
||
</taxonomicName>
|
||
species-group: prosternum strongly convex along midline, with tuft of setae at middle; metanepisternum densely punctate; lateral margins of abdominal sternite VII serrate; apices of gonocoxites square.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |