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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.62.8101" ID-PMC="PMC4856904" ID-Pensoft-Pub="1314-2003-62-73" ID-Pensoft-UUID="BE59A376FFCEA6393859105CFFB1FF82" ID-PubMed="27212883" ID-Zenodo-Dep="576365" ModsDocID="1314-2003-62-73" checkinTime="1458926279483" checkinUser="pensoft" docAuthor="Khine, Phyo Kay, Lindsay, Stuart, Fraser-Jenkins, Christopher, Kluge, Juergen, Kyaw, Myint &amp; Hovenkamp, Peter" docDate="2016" docId="50CE07FAAC92F34CA38B3DE836FB6135" docLanguage="en" docName="PhytoKeys 62: 73-81" docOrigin="PhytoKeys 62" docPubDate="2016-03-25" docSource="http://dx.doi.org/10.3897/phytokeys.62.8101" docTitle="Selliguea kachinensis Hovenkamp, S. Linds., Fraser-Jenk. 2016, sp. nov." docType="treatment" docVersion="7" id="BE59A376FFCEA6393859105CFFB1FF82" lastPageNumber="78" masterDocId="BE59A376FFCEA6393859105CFFB1FF82" masterDocTitle="Selliguea kachinensis (Polypodiaceae), a new fern species of uncertain affinity from Northern Myanmar" masterLastPageNumber="81" masterPageNumber="73" pageNumber="74" updateTime="1668141681425" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Selliguea kachinensis (Polypodiaceae), a new fern species of uncertain affinity from Northern Myanmar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khine, Phyo Kay</mods:namePart>
<mods:affiliation>Faculty of Geography, Philipps University of Marburg, Deutschhausstrasse 10, D- 35032, Marburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lindsay, Stuart</mods:namePart>
<mods:affiliation>Gardens by the Bay, 18 Marina Gardens Drive, Singapore 018953</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fraser-Jenkins, Christopher</mods:namePart>
<mods:affiliation>Student Guest House, Thamel, PO Box 5555, Kathmandu, Nepal</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kluge, Juergen</mods:namePart>
<mods:affiliation>Faculty of Geography, Philipps University of Marburg, Deutschhausstrasse 10, D- 35032, Marburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kyaw, Myint</mods:namePart>
<mods:affiliation>Nature and Wildlife Conservation Division, Forest Department, Ministry of Environmental Conservation and Forestry, Office No. 39, Nay Pyi Taw, Myanmar</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hovenkamp, Peter</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, PO Box 2317 2300 RA Leiden, The Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">peter.hovenkamp@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-03-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>62</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>73</mods:start>
<mods:end>81</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.62.8101</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.62.8101</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-62-73</mods:identifier>
<mods:identifier type="Pensoft-UUID">BE59A376FFCEA6393859105CFFB1FF82</mods:identifier>
<mods:identifier type="Zenodo-Dep">576365</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127877616" LSID="urn:lsid:plazi:treatment:50CE07FAAC92F34CA38B3DE836FB6135" httpUri="http://treatment.plazi.org/id/50CE07FAAC92F34CA38B3DE836FB6135" lastPageId="5" lastPageNumber="78" pageId="1" pageNumber="74">
<subSubSection pageId="1" pageNumber="74" type="nomenclature">
<paragraph pageId="1" pageNumber="74">
<taxonomicName LSID="50CE07FA-AC92-F34C-A38B-3DE836FB6135" authority="Hovenkamp, S. Linds., Fraser-Jenk." authorityName="Hovenkamp, S. Linds., Fraser-Jenk." authorityYear="2016" class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="1" pageNumber="74" phylum="Tracheophyta" rank="species" species="kachinensis" status="sp. nov.">Selliguea kachinensis Hovenkamp, S. Linds., Fraser-Jenk.</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="74">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="74" type="type">
<paragraph pageId="1" pageNumber="74">Type.</paragraph>
<paragraph pageId="1" pageNumber="74">
Myanmar, Kachin State, Hponyin Razi, Quercus-Magnolia-
<taxonomicName genus="Araliaceae" lsidName="Araliaceae" pageId="1" pageNumber="74" rank="genus">Araliaceae</taxonomicName>
forest, epiphyte.
<geoCoordinate degrees="27.601421" direction="north" orientation="latitude" precision="1" value="27.601421">27.601421°N</geoCoordinate>
,
<geoCoordinate degrees="96.988873" direction="east" orientation="longitude" precision="1" value="96.98888">96.988873°E</geoCoordinate>
, 1715 m, G. Miehe, P.K. Khine [
<normalizedToken originalValue="“Kine”">&quot;Kine&quot;</normalizedToken>
], L. Shein, M. Kyaw, P. Ma, S. Lan Wan 13-094-159, 19 Nov. 2013 (holotype: L; isotype: SING).
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74" type="epiphytic">
<paragraph pageId="1" pageNumber="74">Epiphytic.</paragraph>
<paragraph lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74">
Rhizome long-creeping, branched, 3.2-4.5 mm diam. when dry (c. 6 mm diam. after soaking in boiling water), black and shiny when dry with a glaucous waxy layer, the younger parts densely covered with scales, irregularly rooting from the ventral side, in cross-section with scattered sclerification in the peripheral, epidermal and subepidermal region; phyllopodia c. 2 cm distant, c. 1-3 mm high. Scales deciduous, mostly absent from older parts of the rhizome, basifixed and slightly to strongly auriculate, or pseudopeltate, or sometimes fully peltate, c. 0.5
<normalizedToken originalValue="×">x</normalizedToken>
2.0-3.5(-4.0) mm, gradually narrowed from the base to a long narrow acumen, brown or blackish near the attachment, central region brown and thick, the margin and acumen thinner and lighter, margin irregularly dentate, more strongly so towards the base. Fronds pendent, simple, monomorphic, stipitate, all parts densely hairy with multicellular, uniseriate, soft hairs to 1.5(-1.7) mm long, the longest hairs inserted on midrib and veins; sparse,
<pageBreakToken pageId="2" pageNumber="75" start="start">long</pageBreakToken>
narrow pale strongly toothed scales present among the hairs on the abaxial midrib (particularly towards the base of the lamina), stipe 0.8-5.0 cm long, c. 1 mm thick; lamina 18-58
<normalizedToken originalValue="×">x</normalizedToken>
3.3-7.2 cm, oblong - narrowly elliptic, the basal 1-6 cm often narrowed, base truncate to cordate, apex acuminate, texture thin-herbaceous, glaucous when fresh, translucent when dry, margin very narrowly hyaline, without notches. Venation anastomosing, primary veins straight or slightly curved, at 60-90 degrees to the midrib, secondary veins hardly distinct, delimiting c. 5-6 rows of rectangular areoles with anastomosing tertiary veins and free veins in all directions, ending in hydathodes. Sori in a single row between each pair of primary veins, usually one per areole, but sometimes absent from the first one or two areolae closest to the midrib and occasionally two in areolae closest to the margin, c. 2 mm in diameter when ripe. Sporangia long-stalked, capsules c. 0.2 mm long, bearing 2-6 uniseriate, c. 0.4-0.8 mm long hairs, annulus with 14-16 indurated cells. Spores 29-46
<normalizedToken originalValue="×">x</normalizedToken>
25-34
<normalizedToken originalValue="µm">µm</normalizedToken>
in lateral view, perispore with a 0.1-0.3
<normalizedToken originalValue="µm">µm</normalizedToken>
thick, finely colliculate basal layer, rather densely set with narrow, fragile spines, spines c. 2
<normalizedToken originalValue="µm">µm</normalizedToken>
long by 0.5
<normalizedToken originalValue="µm">µm</normalizedToken>
thick at the base, somewhat narrowed to a blunt apex, apparently easily breaking off at the base leaving a low round scar.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="75" type="additional specimens seen">
<paragraph pageId="2" pageNumber="75">Additional specimens seen.</paragraph>
<paragraph pageId="2" pageNumber="75">
Myanmar, Kachin State. Hponyin Razi: G. Miehe, P.K. Khine [
<normalizedToken originalValue="“Kine”">&quot;Kine&quot;</normalizedToken>
], L. Shein, M. Kyaw, P. Ma, S. Lan Wan 13-096-034, 23 Nov. 2013, above 1300 m, road site (L); G. Miehe, P.K. Khine [
<normalizedToken originalValue="“Kine”">&quot;Kine&quot;</normalizedToken>
], L. Shein, M. Kyaw, P. Ma, S. Lan Wan 13-131-013, 11 Nov. 2013, 1600 m, road site (SING).
</paragraph>
<paragraph pageId="2" pageNumber="75">
Hponkan Razi: P.K. Khine [
<normalizedToken originalValue="“Kine”">&quot;Kine&quot;</normalizedToken>
], J. Kluge, A.S. Lanwan, D.R. Lanwan, P. Lanwan 14-031-020, 14 Oct. 2014,
<geoCoordinate degrees="27.548702" direction="north" orientation="latitude" precision="1" value="27.548702">27.548702 N</geoCoordinate>
,
<geoCoordinate degrees="97.032742" direction="east" orientation="longitude" precision="1" value="97.032745">97.032742 E</geoCoordinate>
, 1565 m, evergreen broadleaved (L). Above Ziadam: P.K. Khine [
<normalizedToken originalValue="“Kine”">&quot;Kine&quot;</normalizedToken>
], J. Kluge, A.S. Lanwan, D.R. Lanwan, P. Lanwan 14-047-022, 21 Oct. 2014,
<geoCoordinate degrees="27.585061" direction="north" orientation="latitude" precision="1" value="27.58506">27.585061°N</geoCoordinate>
,
<geoCoordinate degrees="97.104085" direction="east" orientation="longitude" precision="1" value="97.10409">97.104085°E</geoCoordinate>
, 1448 m, evergreen broadleaved (L, SING). Pisa District: Wang Jun &amp; Zhou Lian Xuan 5431, 18 Apr. 2009, Pangjia (
<geoCoordinate degrees="26" direction="north" minutes="31.589" orientation="latitude" precision="1" value="26.526484">26°31.589'N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="18.473" orientation="longitude" precision="1" value="98.307884">98°18.473'E</geoCoordinate>
, 717 m) to Wuru (
<geoCoordinate degrees="26" direction="north" minutes="31.589" orientation="latitude" precision="1" value="26.526484">26°31.589'N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="18.473" orientation="longitude" precision="1" value="98.307884">98°18.473'E</geoCoordinate>
), in forest on a tree (CDBI). Ridan: Xia Nianhe, Deng Yunfei, Zhou Wei &amp; Wu Linfang 1519, 20 Mar. 2009, around the Waqkure village, c. 2 miles from Ridan, E. side of Namai Kha river,
<geoCoordinate degrees="27" direction="north" minutes="11.876" orientation="latitude" precision="1" value="27.197933">27°11.876'N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="15.165" orientation="longitude" precision="1" value="98.25275">98°15.165'E</geoCoordinate>
, 1400 m., in forest (CDBI).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="75" type="etymology">
<paragraph pageId="2" pageNumber="75">Etymology.</paragraph>
<paragraph pageId="2" pageNumber="75">The name derives from Kachin State, where the species is found.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="75" type="ecology">
<paragraph pageId="2" pageNumber="75">Ecology.</paragraph>
<paragraph pageId="2" pageNumber="75">
Based on the specimens collected in the &quot;northern forest complex&quot;,
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="2" pageNumber="75" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="2" pageNumber="75">Selliguea kachinensis</emphasis>
</taxonomicName>
grows on heavily moss-covered trees in primary evergreen broadleaved forest (dominated by
<taxonomicName authorityName="Dumortier" authorityYear="1829" genus="Fagaceae" lsidName="Fagaceae" pageId="2" pageNumber="75" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="75">Fagaceae</emphasis>
</taxonomicName>
,
<taxonomicName genus="Lauraceae" lsidName="Lauraceae" pageId="2" pageNumber="75" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="75">Lauraceae</emphasis>
</taxonomicName>
,
<taxonomicName genus="Araliaceae" lsidName="Araliaceae" pageId="2" pageNumber="75" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="75">Araliaceae</emphasis>
</taxonomicName>
, and
<taxonomicName genus="Magnoliaceae" lsidName="Magnoliaceae" pageId="2" pageNumber="75" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="75">Magnoliaceae</emphasis>
</taxonomicName>
) between 1300 m and 1715 m. It was found occasionally on trunks at 4 m from the ground (Figure
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitat of Selliguea kachinensis: a thick bryophyte covered trunk in Hponyin Razi at about 1,700 m (photograph by P. K. Khine)." figureDoi="10.3897/phytokeys.62.8101.figure3" httpUri="https://binary.pensoft.net/fig/82803" pageId="2" pageNumber="75">3</figureCitation>
), more frequently in the moss cover of trunks and thicker branches above 8 m and in the tree crowns, but is absent in the outer canopy. It is locally abundant together with
<taxonomicName class="Magnoliopsida" family="Orchidaceae" genus="Drynaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Drynaria propinqua" order="Asparagales" pageId="2" pageNumber="75" phylum="Angiospermae" rank="species" species="propinqua">
<emphasis italics="true" pageId="2" pageNumber="75">Drynaria propinqua</emphasis>
</taxonomicName>
(Wall. ex Mett.) J.Sm. (Figure
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Selliguea kachinensis: a thick branch covered with other Polypodiaceae such as Drynaria propinqua, and Orchidaceae in Hponkan Razi at 1,600 m (photograph by P. K. Khine)." figureDoi="10.3897/phytokeys.62.8101.figure4" httpUri="https://binary.pensoft.net/fig/82804" pageId="2" pageNumber="75">4</figureCitation>
). It was not found growing on steep rock cliffs or open banks along trails. During our visits in November 2013, and October 2014 we did not observe any wilting of the fronds (in contrast to
<taxonomicName class="Filicopsida" family="Oleandraceae" genus="Oleandra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Oleandra neriiformis" order="Polypodiales" pageId="2" pageNumber="75" phylum="Pteridophyta" rank="species" species="neriiformis">
<emphasis italics="true" pageId="2" pageNumber="75">Oleandra neriiformis</emphasis>
</taxonomicName>
Cav. and
<taxonomicName authorityName="C. Presl" authorityYear="1836" baseAuthorityName="Hook." class="Filicopsida" family="Oleandraceae" genus="Oleandra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Oleandra wallichii" order="Polypodiales" pageId="2" pageNumber="75" phylum="Pteridophyta" rank="species" species="wallichii">
<emphasis italics="true" pageId="2" pageNumber="75">Oleandra wallichii</emphasis>
</taxonomicName>
C.Presl which are lithophytes/epiphytes with a somewhat similar habit) and so could not assess whether it is deciduous or evergreen, but the herbaceous texture suggests that it is deciduous.
</paragraph>
<caption ID-Zenodo-Dep="979161" doi="10.3897/phytokeys.62.8101.figure1" httpUri="https://binary.pensoft.net/fig/82801" pageId="2" pageNumber="75" start="Figure 1" startId="F1">
<paragraph pageId="2" pageNumber="75">
<emphasis bold="true" pageId="2" pageNumber="75">Figure 1.</emphasis>
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="2" pageNumber="75" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="2" pageNumber="75">Selliguea kachinensis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="2" pageNumber="75">A</emphasis>
habit
<emphasis bold="true" pageId="2" pageNumber="75">B</emphasis>
detail showing venation pattern. After Kine
<emphasis italics="true" pageId="2" pageNumber="75">et al</emphasis>
. 14-047-022 (L). Drawing by
<normalizedToken originalValue="Esmée">Esmee</normalizedToken>
Winkel.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="979163" doi="10.3897/phytokeys.62.8101.figure2" httpUri="https://binary.pensoft.net/fig/82802" pageId="2" pageNumber="75" start="Figure 2" startId="F2">
<paragraph pageId="2" pageNumber="75">
<emphasis bold="true" pageId="2" pageNumber="75">Figure 2.</emphasis>
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="2" pageNumber="75" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="2" pageNumber="75">Selliguea kachinensis</emphasis>
</taxonomicName>
- Distribution.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="979165" doi="10.3897/phytokeys.62.8101.figure3" httpUri="https://binary.pensoft.net/fig/82803" pageId="2" pageNumber="75" start="Figure 3" startId="F3">
<paragraph pageId="2" pageNumber="75">
<emphasis bold="true" pageId="2" pageNumber="75">Figure 3.</emphasis>
Habitat of
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="2" pageNumber="75" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="2" pageNumber="75">Selliguea kachinensis</emphasis>
</taxonomicName>
: a thick bryophyte covered trunk in Hponyin Razi at about 1,700 m (photograph by P. K. Khine).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="979167" doi="10.3897/phytokeys.62.8101.figure4" httpUri="https://binary.pensoft.net/fig/82804" pageId="2" pageNumber="75" start="Figure 4" startId="F4">
<paragraph pageId="2" pageNumber="75">
<emphasis bold="true" pageId="2" pageNumber="75">Figure 4.</emphasis>
Habitat of
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="2" pageNumber="75" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="2" pageNumber="75">Selliguea kachinensis</emphasis>
</taxonomicName>
: a thick branch covered with other
<taxonomicName family="Polypodiaceae" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="2" pageNumber="75" rank="family">Polypodiaceae</taxonomicName>
such as
<taxonomicName class="Magnoliopsida" family="Orchidaceae" genus="Drynaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Drynaria propinqua" order="Asparagales" pageId="2" pageNumber="75" phylum="Angiospermae" rank="species" species="propinqua">
<emphasis italics="true" pageId="2" pageNumber="75">Drynaria propinqua</emphasis>
</taxonomicName>
, and
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asparagales" pageId="2" pageNumber="75" phylum="Angiospermae" rank="family">Orchidaceae</taxonomicName>
in Hponkan Razi at 1,600 m (photograph by P. K. Khine).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="76" type="climate">
<paragraph pageId="3" pageNumber="76">
<pageBreakToken pageId="3" pageNumber="76" start="start">Climate</pageBreakToken>
.
</paragraph>
<paragraph pageId="3" pageNumber="76">The climate station nearest to the collection sites is Putao (450 m a.s.l) in an intramontane basin 50 km to the southeast. It records approximately 4000 mm rainfall between May and October with a pronounced dry season from November to January. We expect that the annual rainfall at altitudes between 1400 and 1800 m a.s.l. exceeds 5000 mm plus an unknown amount of fog precipitation from clouds shrouding the mountains between April and November.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="76" type="distribution, conservation and threats">
<paragraph pageId="3" pageNumber="76">Distribution, conservation and threats.</paragraph>
<paragraph pageId="3" pageNumber="76">
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="3" pageNumber="76" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="3" pageNumber="76">Selliguea kachinensis</emphasis>
</taxonomicName>
is currently known from five locations, all in the north of Kachin State, Myanmar. On the basis of these occurrences, the Extent of Occurrence is 4738 km2, while the known Area of Occupancy is 20 km2. However, as exploration of the area has been very fragmentary, we have little information on the actual occurrence of this species, which could well be more widely distributed along the rim of the Mali Kha / Irawaddy River basin. To date, forests where
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="3" pageNumber="76" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="3" pageNumber="76">Selliguea kachinensis</emphasis>
</taxonomicName>
is found are among the least disturbed submontane evergreen broadleaved forests of the Southeastern Himalaya. Drastically increased population could extend swidden farming and might lead to a reduction of the population, but we have no information on any concrete and current threats to the habitat of the species. Accordingly, we propose a status of Data Deficient (
<bibRefCitation author="IUCN" journalOrPublisher="Blumea" pageId="6" pageNumber="79" refId="B5" refString="IUCN, 2014. Guidelines for Using the IUCN Red List Categories and Criteria." title="Guidelines for Using the IUCN Red List Categories and Criteria" year="2014">IUCN 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="78" pageId="4" pageNumber="77" type="discussion: generic placement">
<paragraph pageId="4" pageNumber="77">
<pageBreakToken pageId="4" pageNumber="77" start="start">Discussion</pageBreakToken>
: generic placement.
</paragraph>
<paragraph lastPageId="5" lastPageNumber="78" pageId="4" pageNumber="77">
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea kachinensis</emphasis>
</taxonomicName>
does not fit easily into the genus
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea</emphasis>
</taxonomicName>
, which contains mostly species with a more coriaceous texture and a distinctly cartilaginous, often notched, margin (although a thin-herbaceous texture is notably present in
<taxonomicName authorityName="Hovenk" authorityYear="2011" class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea pui" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="pui">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea pui</emphasis>
</taxonomicName>
Hovenkamp). An alternative position would be in the related genus
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris</emphasis>
</taxonomicName>
. This would agree with the rather distinctive, glabrescent rhizome, which is similar to that of
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris lehmanii" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="species" species="lehmanii">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris lehmanii</emphasis>
</taxonomicName>
(Mett.) Ching or
<taxonomicName authorityName="W. M. Chu" authorityYear="1992" class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris tomentosa" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="species" species="tomentosa">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris tomentosa</emphasis>
</taxonomicName>
W.M.Chu, and with a number of other characters (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of morphological characteristics of Selliguea kachinensis with Selliguea and Arthromeris." httpUri="http://table.plazi.org/id/21E3FD286DB0C464DD51734F13475A97" pageId="4" pageNumber="77" tableUuid="21E3FD286DB0C464DD51734F13475A97">1</tableCitation>
) but it would seriously weaken the diagnostic value of that genus, as all species so far placed in
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris</emphasis>
</taxonomicName>
have imparipinnate fronds with articulate pinnae (
<bibRefCitation author="Lu, SG" editor="Wu, Z" journalOrPublisher="Science Press, Beijing" pageId="6" pageNumber="79" pagination="768 - 772" refId="B6" refString="Lu, SG, Hovenkamp, PH, 2013. Arthromeris. In: Wu, Z, Raven, PH, Deyuan, H, Eds., Flora of China 2-3. Science Press, Beijing: 768 - 772" title="Arthromeris." volumeTitle="Flora of China 2 - 3." year="2013">Lu and Hovenkamp 2013</bibRefCitation>
;
<bibRefCitation author="Tagawa, M" journalOrPublisher="Flora of Thailand" pageId="6" pageNumber="79" pagination="486 - 580" refId="B8" refString="Tagawa, M, Iwatsuki, K, 1989. Polypodiaceae. Flora of Thailand 3: 486 - 580" title="Polypodiaceae." volume="3" year="1989">Tagawa and Iwatsuki 1989</bibRefCitation>
). There are several distinctive characters in
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea kachinensis</emphasis>
</taxonomicName>
that argue against placement in either of these genera, and for the erection of a new genus. The often somewhat lyrate base of the lamina of
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea kachinensis</emphasis>
</taxonomicName>
is distinctive and not encountered in any other species of either
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea</emphasis>
</taxonomicName>
or
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris</emphasis>
</taxonomicName>
. The rhizome of
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea kachinensis</emphasis>
</taxonomicName>
is also distinct in that the cross-section shows sclerification only in the peripheral, subepidermal region (Figure
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selliguea kachinensis. A cross section of rhizome B rhizome scales C lamina margin D sporangium E spore F detail of spore, scale bar. All from the holotype, Miehe et al. 13 - 094 - 159 (L). Scale bar: 10 µm (A-E); 1 µm (F)." figureDoi="10.3897/phytokeys.62.8101.figure5" httpUri="https://binary.pensoft.net/fig/82805" pageId="4" pageNumber="77">5a</figureCitation>
). Sclerification occurs frequently in
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea</emphasis>
</taxonomicName>
, rarely in
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris</emphasis>
</taxonomicName>
, but in both cases takes the form of sclerified strands in the central part of the rhizome, or a continuous, sclerified band well below the epidermis. The rhizome scales (Figure
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selliguea kachinensis. A cross section of rhizome B rhizome scales C lamina margin D sporangium E spore F detail of spore, scale bar. All from the holotype, Miehe et al. 13 - 094 - 159 (L). Scale bar: 10 µm (A-E); 1 µm (F)." figureDoi="10.3897/phytokeys.62.8101.figure5" httpUri="https://binary.pensoft.net/fig/82805" pageId="4" pageNumber="77">5b</figureCitation>
) do not show any distinctive characters. A dense indument of long hairs similar to the indument of
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea kachinensis</emphasis>
</taxonomicName>
(Figure
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selliguea kachinensis. A cross section of rhizome B rhizome scales C lamina margin D sporangium E spore F detail of spore, scale bar. All from the holotype, Miehe et al. 13 - 094 - 159 (L). Scale bar: 10 µm (A-E); 1 µm (F)." figureDoi="10.3897/phytokeys.62.8101.figure5" httpUri="https://binary.pensoft.net/fig/82805" pageId="4" pageNumber="77">5c, d</figureCitation>
) occurs in some species of
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris</emphasis>
</taxonomicName>
, but in
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea</emphasis>
</taxonomicName>
,
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea trisecta" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="trisecta">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea trisecta</emphasis>
</taxonomicName>
(Baker) Fraser-Jenk. and
<taxonomicName authorityName="X. C. Zhang &amp; L. J. He" authorityYear="2015" baseAuthorityName="Mett. ex Kuhn" class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea erythrocarpa" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="erythrocarpa">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea erythrocarpa</emphasis>
</taxonomicName>
(Mett. ex Kuhn) Hovenkamp, S. Linds., Fraser-Jenk.
<emphasis italics="true" pageId="4" pageNumber="77">comb. nov.</emphasis>
(basionym:
<taxonomicName authorityName="Mett. ex Kuhn" authorityYear="1869" class="Filicopsida" family="Polypodiaceae" genus="Polypodium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Polypodium erythrocarpum" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="species" species="erythrocarpum">
<emphasis italics="true" pageId="4" pageNumber="77">Polypodium erythrocarpum</emphasis>
</taxonomicName>
Mett. ex Kuhn, Linnaea 36: 135. 1869) are also hairy, while
<taxonomicName authorityName="Fraser-Jenk" authorityYear="2008" baseAuthorityName="C. Chr." class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea chrysotricha" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="chrysotricha">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea chrysotricha</emphasis>
</taxonomicName>
(C.Chr.) Fraser-Jenk. also has hairs (albeit short and stiff ones) on the capsules of the sporangia. The spore ornamentation (Figure
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selliguea kachinensis. A cross section of rhizome B rhizome scales C lamina margin D sporangium E spore F detail of spore, scale bar. All from the holotype, Miehe et al. 13 - 094 - 159 (L). Scale bar: 10 µm (A-E); 1 µm (F)." figureDoi="10.3897/phytokeys.62.8101.figure5" httpUri="https://binary.pensoft.net/fig/82805" pageId="4" pageNumber="77">5e, f</figureCitation>
) is matched in
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea</emphasis>
</taxonomicName>
by e.g.
<taxonomicName authorityName="H. Ohashi &amp; K. Ohashi" authorityYear="2009" baseAuthorityName="Hayata" class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea quasidivaricata" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="quasidivaricata">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea quasidivaricata</emphasis>
</taxonomicName>
(Hayata) H.Ohashi &amp; K.Ohashi and
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea yakushimensis" order="Polypodiales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="yakushimensis">
<emphasis italics="true" pageId="4" pageNumber="77">Selliguea yakushimensis</emphasis>
</taxonomicName>
(Makino) Fraser-Jenk. and in
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris</emphasis>
</taxonomicName>
by e.g.
<taxonomicName authorityName="Ching" authorityYear="1933" baseAuthorityName="Hook." class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris tenuicauda" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="species" species="tenuicauda">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris tenuicauda</emphasis>
</taxonomicName>
(Hook.) Ching and
<taxonomicName authorityName="Ching" authorityYear="1933" baseAuthorityName="Mett." class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris lehmannii" order="Polypodiales" pageId="4" pageNumber="77" phylum="Pteridophyta" rank="species" species="lehmannii">
<emphasis italics="true" pageId="4" pageNumber="77">Arthromeris lehmannii</emphasis>
</taxonomicName>
(Mett.) Ching (
<bibRefCitation author="Tryon, AF" journalOrPublisher="Springer, New York" pageId="6" pageNumber="79" publicationUrl="10.1007/978-1-4613-8991-0" refId="B10" refString="Tryon, AF, Lugardon, B, 1991. Spores of the Pteridophyta. Springer, New York, DOI: 10.1007/978-1-4613-8991-0" title="Spores of the Pteridophyta" url="10.1007/978-1-4613-8991-0" year="1991">Tryon and Lugardon 1991</bibRefCitation>
). Thus, there are
<pageBreakToken pageId="5" pageNumber="78" start="start">arguments</pageBreakToken>
both for and against placement in
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea</emphasis>
</taxonomicName>
or in
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="5" pageNumber="78" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Arthromeris</emphasis>
</taxonomicName>
and there are arguments in favour of erecting a new genus. We have decided not to do the latter, as the generic taxonomy of the Selligueoid ferns is at the moment unsettled, has been burdened already by the erection of numerous small genera (
<bibRefCitation author="Hovenkamp, P" journalOrPublisher="Blumea" pageId="6" pageNumber="79" pagination="1 - 108" refId="B4" refString="Hovenkamp, P, 1998. An account of the Malay-Pacific species of Selliguea (Polypodiaceae). Blumea 43: 1 - 108" title="An account of the Malay-Pacific species of Selliguea (Polypodiaceae)." volume="43" year="1998">Hovenkamp 1998</bibRefCitation>
), and it is beginning to become clear that the best option to avoid paraphyletic groups may be to accept a large genus
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea</emphasis>
</taxonomicName>
(
<bibRefCitation author="He, L" journalOrPublisher="Bishen Singh Mahendra Pal Singh, Dehra Dun" pageId="6" pageNumber="79" refId="B3" refString="He, L, Schneider, H, Hovenkamp, P, Marquardt, J, Wei, R, Zhang, X, Xiang, Q, in prep. . Taxonomy in the face of confusion: towards a natural classification of selligueoid ferns despite homoplasy and rapid radiation." title="Taxonomy in the face of confusion: towards a natural classification of selligueoid ferns despite homoplasy and rapid radiation" year="in prep.">He et al. in prep.</bibRefCitation>
). As alternative to a monotypic genus, we prefer a placement in the genus
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea</emphasis>
</taxonomicName>
over one in
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="5" pageNumber="78" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Arthromeris</emphasis>
</taxonomicName>
in anticipation of a generic reorganization along these lines.
</paragraph>
<caption ID-Zenodo-Dep="979169" doi="10.3897/phytokeys.62.8101.figure5" httpUri="https://binary.pensoft.net/fig/82805" pageId="5" pageNumber="78" start="Figure 5" startId="F5">
<paragraph pageId="5" pageNumber="78">
<emphasis bold="true" pageId="5" pageNumber="78">Figure 5.</emphasis>
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea kachinensis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="5" pageNumber="78">A</emphasis>
cross section of rhizome
<emphasis bold="true" pageId="5" pageNumber="78">B</emphasis>
rhizome scales
<emphasis bold="true" pageId="5" pageNumber="78">C</emphasis>
lamina margin
<emphasis bold="true" pageId="5" pageNumber="78">D</emphasis>
sporangium
<emphasis bold="true" pageId="5" pageNumber="78">E</emphasis>
spore
<emphasis bold="true" pageId="5" pageNumber="78">F</emphasis>
detail of spore, scale bar. All from the holotype, Miehe
<emphasis italics="true" pageId="5" pageNumber="78">et al</emphasis>
. 13-094-159 (L). Scale bar: 10
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="5" pageNumber="78">A-E</emphasis>
); 1
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="5" pageNumber="78">F</emphasis>
).
</paragraph>
</caption>
<caption ID-Table-UUID="21E3FD286DB0C464DD51734F13475A97" httpUri="http://table.plazi.org/id/21E3FD286DB0C464DD51734F13475A97" pageId="5" pageNumber="78" start="Table 1" startId="T1">
<paragraph pageId="5" pageNumber="78">
<emphasis bold="true" pageId="5" pageNumber="78">Table 1.</emphasis>
Comparison of morphological characteristics of
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea kachinensis</emphasis>
</taxonomicName>
with
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea</emphasis>
</taxonomicName>
and
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="5" pageNumber="78" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Arthromeris</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="78">
<table pageId="5" pageNumber="78">
<tr pageId="5" pageNumber="78">
<th colspan="1" isEmpty="true" pageId="5" pageNumber="78" rowspan="1">-</th>
<th colspan="1" pageId="5" pageNumber="78" rowspan="1">
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="78" rowspan="1">
<taxonomicName class="Polypodiopsida" family="Polypodiaceae" genus="Selliguea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selliguea kachinensis" order="Polypodiales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis italics="true" pageId="5" pageNumber="78">Selliguea kachinensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="78" rowspan="1">
<taxonomicName class="Filicopsida" family="Polypodiaceae" genus="Arthromeris" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Arthromeris" order="Polypodiales" pageId="5" pageNumber="78" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="78">Arthromeris</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="5" pageNumber="78">
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Rhizome with scattered sclerification</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Absent or central</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Peripheral</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Absent or central</td>
</tr>
<tr pageId="5" pageNumber="78">
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Rhizome with continuous sclerified band</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Often</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">No</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">No</td>
</tr>
<tr pageId="5" pageNumber="78">
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Lamina shape</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Simple to pinnate</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Simple</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Pinnate, pinnae articulate</td>
</tr>
<tr pageId="5" pageNumber="78">
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Lamina texture</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Mostly coriaceous</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Thin-herbaceous</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Thin-herbaceous to herbaceous</td>
</tr>
<tr pageId="5" pageNumber="78">
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Lamina indument</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Glabrous to short-hairy</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Soft hairy</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Glabrous to densely soft-hairy</td>
</tr>
<tr pageId="5" pageNumber="78">
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Lamina margin</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Mostly cartilaginous, often notched</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Not differentiated</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Often distinctly flat-cartilaginous, not notched</td>
</tr>
<tr pageId="5" pageNumber="78">
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Sporangial indument</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Rarely present, short stiff hairs</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Soft long hairs</td>
<td colspan="1" pageId="5" pageNumber="78" rowspan="1">Absent</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>