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<document id="F156AD37FBCC19A54D123F2EA07EA323" ID-DOI="10.21248/contrib.entomol.65.2.223-238" ID-ISSN="0005-805X" ID-Zenodo-Dep="5370714" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630269246923" checkinUser="felipe" docAuthor="Dathe, Holger H." docDate="2015" docId="4F5ABF2F3E59A1507DCDFB06C746F788" docLanguage="en" docName="ContribEntomol.2015.65.1.9-26.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 65 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1880" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Hylaeus (Deranchylaeus) haladanius Dathe 2015, sp. n." docType="treatment" docUuid="BEC60AB0-7090-4428-ACDA-A4C0E37767BC" docUuidSource="ZooBank" docVersion="3" lastPageNumber="20" masterDocId="B363C7573E50A15B7D70FF95C230FFAA" masterDocTitle="Studies on the systematics and taxonomy of the genus Hylaeus F. (9) Supplement to the taxonomy and distribution of Afrotropical Hylaeus F. species (Hymenoptera: Anthophila, Colletidae)" masterLastPageNumber="26" masterPageNumber="9" pageNumber="18" updateTime="1699205946262" updateUser="plazi">
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<mods:title id="B02E42BC3FD07231417215DAAD5BEE63">Studies on the systematics and taxonomy of the genus Hylaeus F. (9) Supplement to the taxonomy and distribution of Afrotropical Hylaeus F. species (Hymenoptera: Anthophila, Colletidae)</mods:title>
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<mods:namePart id="A41882CDBA4272522B0FF0D0B60FDB37">Dathe, Holger H.</mods:namePart>
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<mods:title id="8C992FB74D7B3E877ED8EE1DC3512F24">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="25AE540A73BE71071BBB35C656B818FF">2015</mods:date>
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<mods:number id="B66E347831BBF46C086C09436EE1B9EE">2015-12-21</mods:number>
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<mods:number id="A124384EFB13A89B1B94D4AC8B5CD35D">65</mods:number>
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<treatment id="4F5ABF2F3E59A1507DCDFB06C746F788" LSID="urn:lsid:zoobank.org:act:BEC60AB0-7090-4428-ACDA-A4C0E37767BC" httpUri="http://treatment.plazi.org/id/4F5ABF2F3E59A1507DCDFB06C746F788" lastPageId="11" lastPageNumber="20" pageId="9" pageNumber="18">
<subSubSection id="8FE95DB23E59A1527DCDFB06C366FB65" pageId="9" pageNumber="18" type="nomenclature">
<paragraph id="C74C0E393E59A1527DCDFB06C042FB04" blockId="9.[189,626,1171,1231]" box="[189,626,1171,1198]" pageId="9" pageNumber="18">
<heading id="9C04B9553E59A1527DCDFB06C042FB04" box="[189,626,1171,1198]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="18" reason="3">
<taxonomicName id="00F375BA3E59A1527DCDFB06C003FB04" authority="Dathe, 2015" authorityName="Dathe" authorityYear="2015" box="[189,563,1171,1198]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="18" phylum="Arthropoda" rank="species" species="haladanius" status="sp. nov." subGenus="Deranchylaeus">
<emphasis id="F587D22B3E59A1527DCDFB06C33FFB04" box="[189,271,1171,1198]" italics="true" pageId="9" pageNumber="18">Hylaeus</emphasis>
(
<emphasis id="F587D22B3E59A1527C6CFB06C384FB04" box="[284,436,1171,1198]" italics="true" pageId="9" pageNumber="18">Deranchylaeus</emphasis>
)
<emphasis id="F587D22B3E59A1527CB2FB06C003FB04" box="[450,563,1171,1198]" italics="true" pageId="9" pageNumber="18">haladanius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EEB46F503E59A1527F49FB06C042FB04" box="[569,626,1171,1198]" pageId="9" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="C74C0E393E59A1527DCDFB2CC366FB65" blockId="9.[189,626,1171,1231]" box="[189,342,1209,1231]" pageId="9" pageNumber="18">Figures 1017, 24</paragraph>
</subSubSection>
<subSubSection id="8FE95DB23E59A1527DE7FB7CC383FA2B" pageId="9" pageNumber="18" type="diagnosis">
<paragraph id="C74C0E393E59A1527DE7FB7CC383FA2B" blockId="9.[151,756,1257,1409]" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527DE7FB7CC334FAAB" bold="true" box="[151,260,1257,1281]" pageId="9" pageNumber="18">Diagnosis</emphasis>
: This species is small and dainty, and characterized by its strong sculpture. It exhibits large and shallow punctures on pro- and mesosoma; in contrast, the metasoma is only finely sculptured. The males have remarkably long antennae.
</paragraph>
</subSubSection>
<subSubSection id="8FE95DB23E59A1507DE7FA3CC09CFBCB" lastPageId="11" lastPageNumber="20" pageId="9" pageNumber="18" type="description">
<paragraph id="C74C0E393E59A1527DE7FA3CC33FFA4B" blockId="9.[151,757,1449,1985]" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527DE7FA3CC329FA6B" bold="true" box="[151,281,1449,1473]" pageId="9" pageNumber="18">Description</emphasis>
:
<emphasis id="F587D22B3E59A1527C56FA3CC36DFA6B" bold="true" box="[294,349,1449,1473]" pageId="9" pageNumber="18">Male</emphasis>
. TL 4.14.8 (4.45) mm, WL 3.03.4 (3.21) mm.
</paragraph>
<paragraph id="C74C0E393E59A1527DE7FA7CC667FBAB" blockId="9.[151,757,1449,1985]" lastBlockId="9.[794,1399,937,1985]" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527DE7FA7CC2E1F9AB" bold="true" box="[151,209,1513,1537]" pageId="9" pageNumber="18">Head</emphasis>
. Proportions HL:HW 0.971.00 (0.98), UFW:LFW 1.671.81 (1.74), outline approximately circular, with clypeus expanded downward. Scapus black, short conoid, narrower than flagellum, SL:SW 1.71.8 (1.78). Flagellum conspicuously long, yellow from segment 1, increasingly becoming darker toward apex and dorsally. Mask white to light yellow, supraclypeal area black, lateral markings tapering to orbits, terminating above bases of scapi. Foveae faciales short, forming raised bars. Clypeus CL:CW 1.341.51 (1.42), matt, moderately punctate, punctures with indistinct borders. Supraclypeal area black, basally with only a few large shallow pits; apically narrowed, delimited by high margins, here with close punctation. Frons and vertex nearly without pilosity; area of scapus bases with denser and finer punctation, otherwise with coarse, close punctation. Genae narrowed below, occiput rounded, malar area distinctly formed. Labrum and mandibles black.
</paragraph>
<paragraph id="C74C0E393E59A1527E6AFB9CC6DFFA0B" blockId="9.[794,1399,937,1985]" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527E6AFB9CC1BFFB8B" bold="true" box="[794,911,1033,1057]" pageId="9" pageNumber="18">Mesosoma</emphasis>
compact, slightly elongated, pilosity very sparse and short. Black, bright are only tegulae spots. Pronotum black, distinctly expanded; anterior margin sharp over entire width, with stepped frontbar; dorsolateral angle pointed and bent. Mesonotum and scutellum silkily shining, shagreen with large shallow punctures; mesopleurae similar. Metanotum with sharp wrinkled meshes as on basal area of propodeum. Legs black, white tibial bases, foretibiae in front and whole tarsi. Wings brownish, venation light brown. Propodeum margined, its surface usually shiny; basal and lateral areas delimited by carinae, with raised edges around wrinkled meshes; terminal area surrounded by sharp margins.
</paragraph>
<paragraph id="C74C0E393E59A1527E6AFA3CC702F90B" blockId="9.[794,1399,937,1985]" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527E6AFA3CC1BCFA6B" bold="true" box="[794,908,1449,1473]" pageId="9" pageNumber="18">Metasoma</emphasis>
narrow and slender, black. Sculpture of T1 with microscopic fine grooves, punctation minute, scattered, following terga without punctation. Terga without side fringes on depressions. Terminalia see
<figureCitation id="5FC812BC3E59A152799BF99CC705F98B" box="[1259,1333,1545,1569]" captionStart="Figs 2225" captionStartId="16.[320,360,1980,2002]" captionTargetBox="[369,1274,276,1948]" captionTargetId="figure-102@16.[320,1305,232,1971]" captionTargetPageId="16" captionText="Figs 2225: Male terminalia: metasomal sterna 7 and 8 (S7, S8), genital capsule. 22 Hylaeus (Alfkenylaeus) euphorbiae spec. nov. 23 Hylaeus (Cornylaeus) adamauanis spec. nov. 24 Hylaeus (Deranchylaeus) haladanius spec. nov. 25 Hylaeus (Pumilaeus) pumilus spec. nov., S7: left ventral view, right dorsal view. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5370726" httpUri="https://zenodo.org/record/5370726/files/figure.png" pageId="9" pageNumber="18">Fig. 24</figureCitation>
; genital capsule with prolonged gonoforcipes, these blackened in apical region; S8 compact with short lobes only as in curvicarinatus-group(2); S7 also similar to
<taxonomicName id="00F375BA3E59A1527980F9FFC164F90B" baseAuthorityName="Cameron" baseAuthorityYear="1905" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="18" phylum="Arthropoda" rank="species" species="curvicarinatus">
<emphasis id="F587D22B3E59A1527980F9FFC734F92B" box="[1264,1284,1642,1665]" italics="true" pageId="9" pageNumber="18">H</emphasis>
.
<emphasis id="F587D22B3E59A1527860F9FFC164F90B" italics="true" pageId="9" pageNumber="18">curvicarinatus</emphasis>
</taxonomicName>
, but with reduced lobes and without bristles.
</paragraph>
<paragraph id="C74C0E393E59A1527E6AF93CC746F96B" blockId="9.[794,1399,937,1985]" box="[794,1398,1705,1729]" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527E6AF93CC159F96B" bold="true" box="[794,873,1705,1729]" pageId="9" pageNumber="18">Female</emphasis>
. TL 4.75.5 (5.10) mm, WL 3.23.8 (3.49) mm.
</paragraph>
<paragraph id="C74C0E393E59A1507E6AF95CC2C0FDEB" blockId="9.[794,1399,937,1985]" lastBlockId="11.[151,757,233,1121]" lastPageId="11" lastPageNumber="20" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527E6AF95CC165F94B" bold="true" box="[794,853,1737,1761]" pageId="9" pageNumber="18">Head</emphasis>
. Proportions HL:HW 0.981.01 (0.99), UFW:LFW 1.491.60 (1.55), outline rounded trapezoid, scarcely hairy. Scapus black; flagellum of normal length, black, basoventrally partly yellow. Face totally black, in a few specimens (
<collectingCountry id="BFE44EA93E59A1527EECF8DCC689F8CB" box="[924,1209,1865,1889]" name="Central African Republic" pageId="9" pageNumber="18">Central African Republic</collectingCountry>
) with tiny white spots below on paraocular areas. Fovea facialis short, expanded and more distinct on vertex; foveal groove continues downwards as narrow wrinkled line, sepa- rated from adjacent face by a distict edge; this structure continues nearly around complex eye. Clypeus CL:CW 1.111.20 (1.16), shagreen, silkily shining, punctation only superficial and unclear. Upper supraclypeal area narrow, almost parallel-sided, above merging widely into the frons. Frons and vertex with moderate, dense punctation, scapus area densely but unclearly punctate, with white tomentum. Genae with large shallow punctation, occiput marginate, malar areas significant. Labrum with horseshoe-shaped callus, mandibles rounded bilobed.
</paragraph>
<caption id="938C5EB13E59A1527DE7F878C694F78A" pageId="9" pageNumber="18" startId="9.[151,191,2029,2051]">
<paragraph id="C74C0E393E59A1527DE7F878C694F78A" blockId="9.[151,1399,2029,2081]" pageId="9" pageNumber="18">
<emphasis id="F587D22B3E59A1527DE7F878C395F7A9" bold="true" box="[151,421,2029,2051]" pageId="9" pageNumber="18">Figs 1017 (opposite page)</emphasis>
:
<taxonomicName id="00F375BA3E59A1527CC3F87BC121F7A9" authority="Dathe, 2015" authorityName="Dathe" authorityYear="2015" box="[435,785,2030,2051]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="18" phylum="Arthropoda" rank="species" species="haladanius" status="sp. nov." subGenus="Deranchylaeus">
<emphasis id="F587D22B3E59A1527CC3F87BC3CEF7A9" box="[435,510,2030,2051]" italics="true" pageId="9" pageNumber="18">Hylaeus</emphasis>
(
<emphasis id="F587D22B3E59A1527F7FF87BC0A9F7A9" box="[527,665,2030,2051]" italics="true" pageId="9" pageNumber="18">Deranchylaeus</emphasis>
)
<emphasis id="F587D22B3E59A1527FD9F87BC121F7A9" box="[681,785,2030,2051]" italics="true" pageId="9" pageNumber="18">haladanius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EEB46F503E59A1527E69F87BC149F7A9" box="[793,889,2030,2051]" pageId="9" pageNumber="18" rank="species">spec. nov.</taxonomicNameLabel>
: head in frontal view; mesosoma lateral and dorsal; surface structure of basal terga in dorsal view.
<emphasis id="F587D22B3E59A1527F12F79EC0AEF78A" bold="true" box="[610,670,2059,2080]" pageId="9" pageNumber="18">1013</emphasis>
Typus male.
<emphasis id="F587D22B3E59A1527E5EF79EC159F78A" bold="true" box="[814,873,2059,2080]" pageId="9" pageNumber="18">1417</emphasis>
Paratypus female. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="C74C0E393E5BA1507DE7FDDCC07CFC6B" blockId="11.[151,757,233,1121]" pageId="11" pageNumber="20">
<emphasis id="F587D22B3E5BA1507DE7FDDCC33CFDCB" bold="true" box="[151,268,585,609]" pageId="11" pageNumber="20">Mesosoma</emphasis>
extended, slightly depressed, without pilosity. Coloration entirely black, only tegulae with light spots. Pronotum black, expanded, front margin raised, dorsolateral angle bent. Mesonotum and scutellum shagreen, silkily shining, with strong, dense punctation; mesopleura similarly sculptured to mesonotum, punctation larger and shallower; omaulus edged. Metanotum matt, strongly punctate. Legs entirely black, without light marks. Wing membrane brown, venation brown. Propodeum margined; basal area with marginally limited meshes similar to males, but not quite as pronounced; terminal area enclosed by sharp margins.
</paragraph>
<paragraph id="C74C0E393E5BA1507DE7FC5CC09CFBCB" blockId="11.[151,757,233,1121]" pageId="11" pageNumber="20">
<emphasis id="F587D22B3E5BA1507DE7FC5CC339FC4B" bold="true" box="[151,265,969,993]" pageId="11" pageNumber="20">Metasoma</emphasis>
slender spindle-shaped, black. T1 very finely shagreen, silkily shining, with only extremely minute, scattered punctation or without punctures, the following terga without punctation. Depressions of T1 with sparse ciliar fringes. Ventral end of fringe light yellowish.
</paragraph>
</subSubSection>
<subSubSection id="8FE95DB23E5BA1507DE7FB1CC079FB6B" pageId="11" pageNumber="20" type="distribution">
<paragraph id="C74C0E393E5BA1507DE7FB1CC079FB6B" blockId="11.[151,756,1161,1217]" pageId="11" pageNumber="20">
<emphasis id="F587D22B3E5BA1507DE7FB1CC317FB0B" bold="true" box="[151,295,1161,1185]" pageId="11" pageNumber="20">Distribution:</emphasis>
<collectingCountry id="BFE44EA93E5BA1507C4CFB1CC3A8FB0B" box="[316,408,1161,1185]" name="Burundi" pageId="11" pageNumber="20">Burundi</collectingCountry>
,
<collectingCountry id="BFE44EA93E5BA1507CC1FB1CC015FB0B" box="[433,549,1161,1185]" name="Cameroon" pageId="11" pageNumber="20">Cameroon</collectingCountry>
,
<collectingCountry id="BFE44EA93E5BA1507F4DFB1CC2C8FB6B" name="Central African Republic" pageId="11" pageNumber="20">Central African Republic</collectingCountry>
,
<collectingCountry id="BFE44EA93E5BA1507C72FB3CC363FB6B" box="[258,339,1193,1217]" name="Nigeria" pageId="11" pageNumber="20">Nigeria</collectingCountry>
,
<collectingCountry id="BFE44EA93E5BA1507C2DFB3CC3D6FB6B" box="[349,486,1193,1217]" name="South Africa" pageId="11" pageNumber="20">South Africa</collectingCountry>
,
<collectingCountry id="BFE44EA93E5BA1507C80FB3CC075FB6B" box="[496,581,1193,1217]" name="Zambia" pageId="11" pageNumber="20">Zambia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8FE95DB23E5BA1507DE7FB7CC1E6FDAB" pageId="11" pageNumber="20" type="materials_examined">
<paragraph id="C74C0E393E5BA1507DE7FB7CC382FAAB" blockId="11.[151,757,1257,2049]" box="[151,434,1257,1281]" pageId="11" pageNumber="20">
<emphasis id="F587D22B3E5BA1507DE7FB7CC31FFAAB" bold="true" box="[151,303,1257,1281]" pageId="11" pageNumber="20">Type material</emphasis>
:
<specimenCount id="D1F5C5B03E5BA1507C4CFB7CC35EFAAB" box="[316,366,1257,1281]" pageId="11" pageNumber="20" type="male">18 ♂</specimenCount>
,
<specimenCount id="D1F5C5B03E5BA1507C0AFB7CC39DFAAB" box="[378,429,1257,1281]" pageId="11" pageNumber="20" type="female">41 ♀</specimenCount>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507DE7FA9CC38DFACB" blockId="11.[151,757,1257,2049]" pageId="11" pageNumber="20">
<materialsCitation id="779B04643E5BA1507DE7FA9CC388FACB" collectingDate="2010-05-11" collectionCode="SANC" collectorName="C. D. Eardley &amp; In" country="Burundi" elevation="1401" latitude="-2.9866667" location="Ruvubu National Park" longLatPrecision="13" longitude="30.468666" municipality="Ruvubu River" pageId="11" pageNumber="20" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="F587D22B3E5BA1507DE7FA9CC33AFA8B" bold="true" box="[151,266,1289,1313]" pageId="11" pageNumber="20">
<typeStatus id="1848B09B3E5BA1507DE7FA9CC33AFA8B" box="[151,266,1289,1313]" pageId="11" pageNumber="20" type="holotype">Holotypus</typeStatus>
</emphasis>
:
<specimenCount id="D1F5C5B03E5BA1507C69FA9FC31EFA8B" box="[281,302,1290,1313]" pageId="11" pageNumber="20" type="male"></specimenCount>
<collectingCountry id="BFE44EA93E5BA1507C46FA9CC3A3FA8B" box="[310,403,1289,1313]" name="Burundi" pageId="11" pageNumber="20">Burundi</collectingCountry>
:
<location id="C22C58E23E5BA1507CEFFA9CC0A3FA8B" LSID="urn:lsid:plazi:treatment:4F5ABF2F3E59A1507DCDFB06C746F788:C22C58E23E5BA1507CEFFA9CC0A3FA8B" box="[415,659,1289,1313]" country="Burundi" latitude="-2.9866667" longLatPrecision="13" longitude="30.468666" municipality="Ruvubu River" name="Ruvubu National Park" pageId="11" pageNumber="20">Ruvubu National Park</location>
,
<collectingMunicipality id="272894433E5BA1507FD0FA9CC2E1FAEB" pageId="11" pageNumber="20">Ruvubu River</collectingMunicipality>
,
<geoCoordinate id="A2C768FE3E5BA1507D91FABCC37EFAEB" box="[225,334,1321,1345]" degrees="02" direction="south" minutes="59.20" orientation="latitude" pageId="11" pageNumber="20" precision="9" value="-2.9866667">02°59.20'S</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507C2BFABCC3FCFAEB" box="[347,460,1321,1345]" degrees="30" direction="east" minutes="28.12" orientation="longitude" pageId="11" pageNumber="20" precision="9" value="30.468666">30°28.12'E</geoCoordinate>
,
<quantity id="000BA3DC3E5BA1507CADFABCC01EFAEB" box="[477,558,1321,1345]" metricMagnitude="3" metricUnit="m" metricValue="1.401" pageId="11" pageNumber="20" unit="m" value="1401.0">
<elevation id="4CDEE90A3E5BA1507CADFABCC01EFAEB" box="[477,558,1321,1345]" metricMagnitude="3" metricUnit="m" metricValue="1.401" pageId="11" pageNumber="20" unit="m" value="1401.0">1401 m</elevation>
</quantity>
,
<date id="B34D28F93E5BA1507F4EFABCC083FAEB" box="[574,691,1321,1345]" pageId="11" pageNumber="20" value="2010-05-11">
<collectingDate id="A309D1113E5BA1507F4EFABCC083FAEB" box="[574,691,1321,1345]" pageId="11" pageNumber="20" value="2010-05-11">05.11.2010</collectingDate>
</date>
,
<collectorName id="6A066BEF3E5BA1507FB3FABCC2D8FACB" pageId="11" pageNumber="20">C.D. Eardley</collectorName>
lg.
<collectorName id="6A066BEF3E5BA1507C52FADFC309FACB" box="[290,313,1354,1377]" pageId="11" pageNumber="20">In</collectorName>
coll.
<collectionCode id="A1E296FC3E5BA1507C01FADCC388FACB" box="[369,440,1353,1377]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="11" pageNumber="20" type="Museum">SANC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507DE7FAFCC2E6FA4B" blockId="11.[151,757,1257,2049]" pageId="11" pageNumber="20">
<materialsCitation id="779B04643E5BA1507DE7FAFCC2E2FA4B" collectingDate="2010-05-11" collectionCode="SANC, SDEI" collectorName="C. D. Eardley &amp; In" country="Burundi" elevation="1401" latitude="-2.988889" location="Ruvubu National Park" longLatPrecision="21" longitude="30.470001" municipality="Ruvubu River" pageId="11" pageNumber="20" specimenCount="6" specimenCount-female="4" specimenCount-male="2" typeStatus="paratype">
<emphasis id="F587D22B3E5BA1507DE7FAFCC332FA2B" bold="true" box="[151,258,1385,1409]" pageId="11" pageNumber="20">
<typeStatus id="1848B09B3E5BA1507DE7FAFCC332FA2B" box="[151,258,1385,1409]" pageId="11" pageNumber="20" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="BFE44EA93E5BA1507C68FAFCC345FA2B" box="[280,373,1385,1409]" name="Burundi" pageId="11" pageNumber="20">Burundi</collectingCountry>
:
<location id="C22C58E23E5BA1507CF8FAFCC0BCFA2B" LSID="urn:lsid:plazi:treatment:4F5ABF2F3E59A1507DCDFB06C746F788:C22C58E23E5BA1507CF8FAFCC0BCFA2B" box="[392,652,1385,1409]" country="Burundi" latitude="-2.988889" longLatPrecision="21" longitude="30.470001" municipality="Ruvubu River" name="Ruvubu National Park" pageId="11" pageNumber="20">Ruvubu National Park</location>
,
<collectingMunicipality id="272894433E5BA1507FD0FAFCC2E1FA0B" pageId="11" pageNumber="20">Ruvubu River</collectingMunicipality>
,
<geoCoordinate id="A2C768FE3E5BA1507DAAFA1CC37AFA0B" box="[218,330,1417,1441]" degrees="02" direction="south" minutes="59" orientation="latitude" pageId="11" pageNumber="20" precision="15" seconds="20" value="-2.988889">02°59'20&quot;S</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507C3FFA1CC3F3FA0B" box="[335,451,1417,1441]" degrees="30" direction="east" minutes="28" orientation="longitude" pageId="11" pageNumber="20" precision="15" seconds="12" value="30.470001">30°28'12&quot;E</geoCoordinate>
,
<quantity id="000BA3DC3E5BA1507CBCFA1CC02EFA0B" box="[460,542,1417,1441]" metricMagnitude="3" metricUnit="m" metricValue="1.401" pageId="11" pageNumber="20" unit="m" value="1401.0">
<elevation id="4CDEE90A3E5BA1507CBCFA1CC02EFA0B" box="[460,542,1417,1441]" metricMagnitude="3" metricUnit="m" metricValue="1.401" pageId="11" pageNumber="20" unit="m" value="1401.0">1401 m</elevation>
</quantity>
,
<date id="B34D28F93E5BA1507F57FA1CC0ABFA0B" box="[551,667,1417,1441]" pageId="11" pageNumber="20" value="2010-05-11">
<collectingDate id="A309D1113E5BA1507F57FA1CC0ABFA0B" box="[551,667,1417,1441]" pageId="11" pageNumber="20" value="2010-05-11">05.11.2010</collectingDate>
</date>
,
<specimenCount id="D1F5C5B03E5BA1507FD5FA1CC0FAFA0B" box="[677,714,1417,1441]" pageId="11" pageNumber="20" type="male">2 ♂</specimenCount>
<specimenCount id="D1F5C5B03E5BA1507FBFFA1CC0C4FA0B" box="[719,756,1417,1441]" pageId="11" pageNumber="20" type="female">4 ♀</specimenCount>
(same data as holotype),
<collectorName id="6A066BEF3E5BA1507CECFA3CC013FA6B" box="[412,547,1449,1473]" pageId="11" pageNumber="20">C.D. Eardley</collectorName>
lg.
<collectorName id="6A066BEF3E5BA1507F2BFA3FC042FA6B" box="[603,626,1450,1473]" pageId="11" pageNumber="20">In</collectorName>
coll.
<collectionCode id="A1E296FC3E5BA1507FD9FA3CC0C0FA6B" box="[681,752,1449,1473]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="11" pageNumber="20" type="Museum">SANC</collectionCode>
,
<collectionCode id="A1E296FC3E5BA1507DE7FA5CC2E2FA4B" box="[151,210,1481,1505]" pageId="11" pageNumber="20">SDEI</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507DE7FA7CC32AF88B" blockId="11.[151,757,1257,2049]" pageId="11" pageNumber="20">
<materialsCitation id="779B04643E5BA1507DE7FA7CC357F98B" collectingDate="1987-07-24" collectorName="Adamaoua &amp; Beka &amp; Ngaoundere" country="Cameroon" latitude="7.2833333" location="Cameroon" longLatPrecision="1303" longitude="13.316667" pageId="11" pageNumber="20" specimenCount="1">
<collectingCountry id="BFE44EA93E5BA1507DE7FA7CC33CF9AB" box="[151,268,1513,1537]" name="Cameroon" pageId="11" pageNumber="20">Cameroon</collectingCountry>
:
<collectorName id="6A066BEF3E5BA1507C6CFA7CC3A3F9AB" box="[284,403,1513,1537]" pageId="11" pageNumber="20">Adamaoua</collectorName>
:
<collectorName id="6A066BEF3E5BA1507CD3FA7CC3E7F9AB" box="[419,471,1513,1537]" pageId="11" pageNumber="20">Beka</collectorName>
nr
<collectorName id="6A066BEF3E5BA1507F79FA7CC0BFF9AB" box="[521,655,1513,1537]" pageId="11" pageNumber="20">Ngaoundere</collectorName>
,
<geoCoordinate id="A2C768FE3E5BA1507FEFFA7CC0C4F9AB" box="[671,756,1513,1537]" degrees="07" direction="north" minutes="17" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="7.2833333">07°17'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507DE7F99CC2D9F98B" box="[151,233,1545,1569]" degrees="13" direction="east" minutes="19" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="13.316667">13°19'E</geoCoordinate>
,
<date id="B34D28F93E5BA1507D82F99CC357F98B" box="[242,359,1545,1569]" pageId="11" pageNumber="20" value="1987-07-24">
<collectingDate id="A309D1113E5BA1507D82F99CC357F98B" box="[242,359,1545,1569]" pageId="11" pageNumber="20" value="1987-07-24">24.07.1987</collectingDate>
</date>
</materialsCitation>
,
<specimenCount id="D1F5C5B03E5BA1507C00F99CC3A6F98B" box="[368,406,1545,1569]" pageId="11" pageNumber="20" type="male">3 ♂</specimenCount>
<specimenCount id="D1F5C5B03E5BA1507CEAF99CC3F0F98B" box="[410,448,1545,1569]" pageId="11" pageNumber="20" type="female">3 ♀</specimenCount>
, on
<taxonomicName id="00F375BA3E5BA1507C9CF99CC041F98B" box="[492,625,1545,1569]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="11" pageNumber="20" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
; Adamaoua: de Banyo à Tibati,
<geoCoordinate id="A2C768FE3E5BA1507C2CF9BCC381F9EB" box="[348,433,1577,1601]" degrees="06" direction="north" minutes="37" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="6.616667">06°37'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507CC7F9BCC038F9EB" box="[439,520,1577,1601]" degrees="12" direction="east" minutes="07" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="12.116667">12°07'E</geoCoordinate>
,
<date id="B34D28F93E5BA1507F63F9BCC0B8F9EB" box="[531,648,1577,1601]" pageId="11" pageNumber="20" value="1987-07-26">26.07.1987</date>
,
<specimenCount id="D1F5C5B03E5BA1507FE2F9BCC087F9EB" box="[658,695,1577,1601]" pageId="11" pageNumber="20" type="male">3 ♂</specimenCount>
<specimenCount id="D1F5C5B03E5BA1507FCCF9BCC0DEF9EB" box="[700,750,1577,1601]" pageId="11" pageNumber="20" type="female">12 ♀</specimenCount>
; Adamaoua: Louga W Ngaoundere,
<geoCoordinate id="A2C768FE3E5BA1507F4BF9DCC0A0F9CB" box="[571,656,1609,1633]" degrees="07" direction="north" minutes="05" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="7.0833335">07°05'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507FEEF9DCC0DFF9CB" box="[670,751,1609,1633]" degrees="13" direction="east" minutes="12" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="13.2">13°12'E</geoCoordinate>
,
<date id="B34D28F93E5BA1507DE7F9FCC33CF92B" box="[151,268,1641,1665]" pageId="11" pageNumber="20" value="1987-07-24">24.07.1987</date>
,
<specimenCount id="D1F5C5B03E5BA1507C66F9FCC30BF92B" box="[278,315,1641,1665]" pageId="11" pageNumber="20" type="male">1 ♂</specimenCount>
<specimenCount id="D1F5C5B03E5BA1507C31F9FCC357F92B" box="[321,359,1641,1665]" pageId="11" pageNumber="20" type="female">1 ♀</specimenCount>
; Adamaoua: Massif du Mbam, Jakiri,
<geoCoordinate id="A2C768FE3E5BA1507DE7F91CC2DBF90B" box="[151,235,1673,1697]" degrees="06" direction="north" minutes="01" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="6.016667">06°01'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507D8CF91CC37EF90B" box="[252,334,1673,1697]" degrees="10" direction="east" minutes="33" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="10.55">10°33'E</geoCoordinate>
,
<date id="B34D28F93E5BA1507C11F91CC3E6F90B" box="[353,470,1673,1697]" pageId="11" pageNumber="20" value="1987-07-22">22.07.1987</date>
,
<specimenCount id="D1F5C5B03E5BA1507C9AF91CC03FF90B" box="[490,527,1673,1697]" pageId="11" pageNumber="20" type="female">1 ♀</specimenCount>
; Adamaoua: Mayo- Darlé,
<geoCoordinate id="A2C768FE3E5BA1507D90F93CC305F96B" box="[224,309,1705,1729]" degrees="06" direction="north" minutes="28" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="6.4666667">06°28'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507C4CF93CC3BEF96B" box="[316,398,1705,1729]" degrees="11" direction="east" minutes="33" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="11.55">11°33'E</geoCoordinate>
,
<date id="B34D28F93E5BA1507CEAF93CC03FF96B" box="[410,527,1705,1729]" pageId="11" pageNumber="20" value="1987-07-24">24.07.1987</date>
,
<specimenCount id="D1F5C5B03E5BA1507F6BF93CC070F96B" box="[539,576,1705,1729]" pageId="11" pageNumber="20" type="male">1 ♂</specimenCount>
<specimenCount id="D1F5C5B03E5BA1507F37F93CC05CF96B" box="[583,620,1705,1729]" pageId="11" pageNumber="20" type="female">1 ♀</specimenCount>
; Adamaoua: Tékel,
<geoCoordinate id="A2C768FE3E5BA1507DAFF95CC303F94B" box="[223,307,1737,1761]" degrees="06" direction="north" minutes="47" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="6.7833333">06°47'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507C4DF95CC3BFF94B" box="[317,399,1737,1761]" degrees="13" direction="east" minutes="10" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="13.166667">13°10'E</geoCoordinate>
,
<date id="B34D28F93E5BA1507CEDF95CC021F94B" box="[413,529,1737,1761]" pageId="11" pageNumber="20" value="1987-07-24">24.07.1987</date>
,
<specimenCount id="D1F5C5B03E5BA1507F6FF95CC074F94B" box="[543,580,1737,1761]" pageId="11" pageNumber="20" type="male">6 ♂</specimenCount>
<specimenCount id="D1F5C5B03E5BA1507F3DF95CC04FF94B" box="[589,639,1737,1761]" pageId="11" pageNumber="20" type="female">13 ♀</specimenCount>
. W Wum,
<geoCoordinate id="A2C768FE3E5BA1507DE7F97CC2DCF8AB" box="[151,236,1769,1793]" degrees="06" direction="north" minutes="24" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="6.4">06°24'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507D85F97CC378F8AB" box="[245,328,1769,1793]" degrees="10" direction="east" minutes="03" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="10.05">10°03'E</geoCoordinate>
, 20.07.1987,
<specimenCount id="D1F5C5B03E5BA1507C95F97CC03AF8AB" box="[485,522,1769,1793]" pageId="11" pageNumber="20" type="female">2 ♀</specimenCount>
;
<materialsCitation id="779B04643E5BA1507F69F97CC326F88B" collectionCode="RBINS" collectorName="Pauly &amp; In" country="Cameroon" pageId="11" pageNumber="20" specimenCount="1">
all A.
<collectorName id="6A066BEF3E5BA1507F2EF97CC0A9F8AB" box="[606,665,1769,1793]" pageId="11" pageNumber="20">Pauly</collectorName>
lg.
<collectorName id="6A066BEF3E5BA1507FADF97FC0C4F8AB" box="[733,756,1770,1793]" pageId="11" pageNumber="20">In</collectorName>
coll.
<collectionCode id="A1E296FC3E5BA1507DB9F89CC326F88B" box="[201,278,1801,1825]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="20" type="Museum">RBINS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507DE7F8BCC312F80B" blockId="11.[151,757,1257,2049]" pageId="11" pageNumber="20">
<materialsCitation id="779B04643E5BA1507DE7F8BCC3D0F8CB" collectingDate="2010-04-04" country="Central African Republic" elevation="380" latitude="4.7666664" location="Bangui" longLatPrecision="1306" longitude="18.066668" pageId="11" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="Bangui">
<collectingCountry id="BFE44EA93E5BA1507DE7F8BCC39FF8EB" box="[151,431,1833,1857]" name="Central African Republic" pageId="11" pageNumber="20">Central African Republic</collectingCountry>
:
<quantity id="000BA3DC3E5BA1507CCDF8BCC030F8EB" box="[445,512,1833,1857]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="20" unit="km" value="85.0">85 km</quantity>
NW
<collectingRegion id="0537C0DB3E5BA1507F33F8BCC0A1F8EB" box="[579,657,1833,1857]" country="Central African Republic" name="Bangui" pageId="11" pageNumber="20">Bangui</collectingRegion>
,
<geoCoordinate id="A2C768FE3E5BA1507FEFF8BCC0C4F8EB" box="[671,756,1833,1857]" degrees="04" direction="north" minutes="46" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="4.7666664">04°46'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507DE7F8DCC2D9F8CB" box="[151,233,1865,1889]" degrees="18" direction="east" minutes="04" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="18.066668">18°04'E</geoCoordinate>
,
<quantity id="000BA3DC3E5BA1507D82F8DCC306F8CB" box="[242,310,1865,1889]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="20" unit="m" value="380.0">
<elevation id="4CDEE90A3E5BA1507D82F8DCC306F8CB" box="[242,310,1865,1889]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="20" unit="m" value="380.0">380 m</elevation>
</quantity>
,
<date id="B34D28F93E5BA1507C4EF8DCC383F8CB" box="[318,435,1865,1889]" pageId="11" pageNumber="20" value="2010-04-04">
<collectingDate id="A309D1113E5BA1507C4EF8DCC383F8CB" box="[318,435,1865,1889]" pageId="11" pageNumber="20" value="2010-04-04">04.04.2010</collectingDate>
</date>
,
<specimenCount id="D1F5C5B03E5BA1507CCBF8DCC3D0F8CB" box="[443,480,1865,1889]" pageId="11" pageNumber="20" type="female">2 ♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="779B04643E5BA1507C9AF8DCC32EF80B" collectingDate="2009-03-06" collectionCode="OLBL, SDEI" collectorName="J. Halada &amp; In" country="Central African Republic" elevation="820" latitude="5.6" location="Carnot" longLatPrecision="1305" longitude="15.9" pageId="11" pageNumber="20" specimenCount="1" specimenCount-female="1">
<quantity id="000BA3DC3E5BA1507C9AF8DCC01DF8CB" box="[490,557,1865,1889]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="20" unit="km" value="90.0">90 km</quantity>
N
<location id="C22C58E23E5BA1507F39F8DCC0A6F8CB" LSID="urn:lsid:plazi:treatment:4F5ABF2F3E59A1507DCDFB06C746F788:C22C58E23E5BA1507F39F8DCC0A6F8CB" box="[585,662,1865,1889]" country="Central African Republic" latitude="5.6" longLatPrecision="1305" longitude="15.9" name="Carnot" pageId="11" pageNumber="20">Carnot</location>
,
<geoCoordinate id="A2C768FE3E5BA1507FEFF8DCC0C4F8CB" box="[671,756,1865,1889]" degrees="05" direction="north" minutes="36" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="5.6">05°36'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507DE7F8FCC2D9F82B" box="[151,233,1897,1921]" degrees="15" direction="east" minutes="54" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="15.9">15°54'E</geoCoordinate>
,
<quantity id="000BA3DC3E5BA1507D83F8FCC308F82B" box="[243,312,1897,1921]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="20" unit="m" value="820.0">
<elevation id="4CDEE90A3E5BA1507D83F8FCC308F82B" box="[243,312,1897,1921]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="20" unit="m" value="820.0">820 m</elevation>
</quantity>
,
<date id="B34D28F93E5BA1507C32F8FCC386F82B" box="[322,438,1897,1921]" pageId="11" pageNumber="20" value="2009-03-06">
<collectingDate id="A309D1113E5BA1507C32F8FCC386F82B" box="[322,438,1897,1921]" pageId="11" pageNumber="20" value="2009-03-06">03.06.2009</collectingDate>
</date>
,
<specimenCount id="D1F5C5B03E5BA1507CB0F8FCC3D6F82B" box="[448,486,1897,1921]" pageId="11" pageNumber="20" type="female">1 ♀</specimenCount>
; all
<collectorName id="6A066BEF3E5BA1507F61F8FFC041F82B" box="[529,625,1897,1921]" pageId="11" pageNumber="20">J. Halada</collectorName>
lg.
<collectorName id="6A066BEF3E5BA1507FDAF8FFC0F1F82B" box="[682,705,1898,1921]" pageId="11" pageNumber="20">In</collectorName>
coll.
<collectionCode id="A1E296FC3E5BA1507DE7F81CC2E9F80B" box="[151,217,1929,1953]" pageId="11" pageNumber="20">OLBL</collectionCode>
,
<collectionCode id="A1E296FC3E5BA1507D93F81CC32EF80B" box="[227,286,1929,1953]" pageId="11" pageNumber="20">SDEI</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507DE7F83CC31EF7AB" blockId="11.[151,757,1257,2049]" pageId="11" pageNumber="20">
<materialsCitation id="779B04643E5BA1507DE7F83CC31AF7AB" collectingDate="2011-08-05" collectionCode="OLBL" collectorName="Gashaka Gumfi NP &amp; J. Halada &amp; In" country="Nigeria" elevation="450" latitude="7.35" location="Serti" longLatPrecision="1303" longitude="11.533334" pageId="11" pageNumber="20" specimenCount="1" specimenCount-female="1">
<collectingCountry id="BFE44EA93E5BA1507DE7F83CC2D9F86B" box="[151,233,1961,1985]" name="Nigeria" pageId="11" pageNumber="20">Nigeria</collectingCountry>
:
<collectorName id="6A066BEF3E5BA1507D82F83CC38FF86B" box="[242,447,1961,1985]" pageId="11" pageNumber="20">Gashaka Gumfi NP</collectorName>
,
<quantity id="000BA3DC3E5BA1507CB9F83CC03CF86B" box="[457,524,1961,1985]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="20" unit="km" value="30.0">30 km</quantity>
SE
<location id="C22C58E23E5BA1507F42F83CC054F86B" LSID="urn:lsid:plazi:treatment:4F5ABF2F3E59A1507DCDFB06C746F788:C22C58E23E5BA1507F42F83CC054F86B" box="[562,612,1961,1985]" country="Nigeria" latitude="7.35" longLatPrecision="1303" longitude="11.533334" name="Serti" pageId="11" pageNumber="20">Serti</location>
env.,
<geoCoordinate id="A2C768FE3E5BA1507FEFF83CC0C4F86B" box="[671,756,1961,1985]" degrees="07" direction="north" minutes="21" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="7.35">07°21'N</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507DE7F85CC2D9F84B" box="[151,233,1993,2017]" degrees="11" direction="east" minutes="32" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="11.533334">11°32'E</geoCoordinate>
, ca.
<quantity id="000BA3DC3E5BA1507C66F85CC36BF84B" box="[278,347,1993,2017]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="20" unit="m" value="450.0">
<elevation id="4CDEE90A3E5BA1507C66F85CC36BF84B" box="[278,347,1993,2017]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="20" unit="m" value="450.0">450 m</elevation>
</quantity>
, 24.04.
<date id="B34D28F93E5BA1507CC0F85CC016F84B" box="[432,550,1993,2017]" pageId="11" pageNumber="20" value="2011-08-05">
<collectingDate id="A309D1113E5BA1507CC0F85CC016F84B" box="[432,550,1993,2017]" pageId="11" pageNumber="20" value="2011-08-05">08.05.2011</collectingDate>
</date>
,
<specimenCount id="D1F5C5B03E5BA1507F40F85CC065F84B" box="[560,597,1993,2017]" pageId="11" pageNumber="20" type="female">1 ♀</specimenCount>
,
<collectorName id="6A066BEF3E5BA1507F11F85FC0F1F84B" box="[609,705,1993,2017]" pageId="11" pageNumber="20">J. Halada</collectorName>
lg.
<collectorName id="6A066BEF3E5BA1507DE7F87FC29EF7AB" box="[151,174,2026,2049]" pageId="11" pageNumber="20">In</collectorName>
coll.
<collectionCode id="A1E296FC3E5BA1507D97F87CC31AF7AB" box="[231,298,2025,2049]" pageId="11" pageNumber="20">OLBL</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507E6AFF7CC155FEEB" blockId="11.[794,1399,233,385]" pageId="11" pageNumber="20">
<materialsCitation id="779B04643E5BA1507E6AFF7CC151FEEB" collectingDate="2011-01-31" collectionCode="SANC" collectorName="C. D. Eardley &amp; In" country="South Africa" latitude="-30.93" location="Vosburg" longLatPrecision="20" longitude="24.934721" pageId="11" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Eastern Cape">
<collectingCountry id="BFE44EA93E5BA1507E6AFF7CC194FEAB" box="[794,932,233,257]" name="South Africa" pageId="11" pageNumber="20">South Africa</collectingCountry>
:
<collectingRegion id="0537C0DB3E5BA1507EDDFF7FC60EFEAB" box="[941,1086,233,257]" country="South Africa" name="Eastern Cape" pageId="11" pageNumber="20">Eastern Cape</collectingRegion>
:
<quantity id="000BA3DC3E5BA1507936FF7CC6B9FEAB" box="[1094,1161,233,257]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="20" unit="km" value="18.0">18 km</quantity>
S
<location id="C22C58E23E5BA15079D2FF7CC6CDFEAB" LSID="urn:lsid:plazi:treatment:4F5ABF2F3E59A1507DCDFB06C746F788:C22C58E23E5BA15079D2FF7CC6CDFEAB" box="[1186,1277,233,257]" country="South Africa" latitude="-30.93" longLatPrecision="20" longitude="24.934721" name="Vosburg" pageId="11" pageNumber="20" stateProvince="Eastern Cape">Vosburg</location>
,
<geoCoordinate id="A2C768FE3E5BA1507877FF7CC746FEAB" box="[1287,1398,233,257]" degrees="30" direction="south" minutes="55" orientation="latitude" pageId="11" pageNumber="20" precision="15" seconds="48" value="-30.93">30°55'48&quot;S</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA1507E6AFE9CC1BEFE8B" box="[794,910,265,289]" degrees="24" direction="east" minutes="56" orientation="longitude" pageId="11" pageNumber="20" precision="15" seconds="05" value="24.934721">24°56'05&quot;E</geoCoordinate>
,
<date id="B34D28F93E5BA1507EECFE9CC621FE8B" box="[924,1041,265,289]" pageId="11" pageNumber="20" value="2011-01-31">
<collectingDate id="A309D1113E5BA1507EECFE9CC621FE8B" box="[924,1041,265,289]" pageId="11" pageNumber="20" value="2011-01-31">31.01.2011</collectingDate>
</date>
,
<specimenCount id="D1F5C5B03E5BA1507950FE9CC675FE8B" box="[1056,1093,265,289]" pageId="11" pageNumber="20" type="male">1 ♂</specimenCount>
,
<collectorName id="6A066BEF3E5BA1507925FE9CC6D1FE8B" box="[1109,1249,265,289]" pageId="11" pageNumber="20">C.D. Eardley</collectorName>
lg.
<collectorName id="6A066BEF3E5BA1507858FE9FC70FFE8B" box="[1320,1343,266,289]" pageId="11" pageNumber="20">In</collectorName>
coll.
<collectionCode id="A1E296FC3E5BA1507E6AFEBCC151FEEB" box="[794,865,297,321]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="11" pageNumber="20" type="Museum">SANC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507E6AFEDCC738FE2B" blockId="11.[794,1399,233,385]" pageId="11" pageNumber="20">
<materialsCitation id="779B04643E5BA1507E6AFEDCC734FE2B" collectingDate="2009-12-12" collectionCode="OLBL" collectorName="F. Kantner &amp; In" country="Zambia" elevation="1250" latitude="-10.5" location="Kasama" longLatPrecision="1298" longitude="31.2" pageId="11" pageNumber="20" specimenCount="1" specimenCount-female="1">
<collectingCountry id="BFE44EA93E5BA1507E6AFEDCC15FFECB" box="[794,879,329,353]" name="Zambia" pageId="11" pageNumber="20">Zambia</collectingCountry>
:
<quantity id="000BA3DC3E5BA1507EF2FEDCC1F5FECB" box="[898,965,329,353]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="20" unit="km" value="35.0">35 km</quantity>
S
<location id="C22C58E23E5BA1507E82FEDFC678FECB" LSID="urn:lsid:plazi:treatment:4F5ABF2F3E59A1507DCDFB06C746F788:C22C58E23E5BA1507E82FEDFC678FECB" box="[1010,1096,330,353]" country="Zambia" latitude="-10.5" longLatPrecision="1298" longitude="31.2" name="Kasama" pageId="11" pageNumber="20">Kasama</location>
,
<geoCoordinate id="A2C768FE3E5BA150792DFEDCC69AFECB" box="[1117,1194,329,353]" degrees="10" direction="south" minutes="30" orientation="latitude" pageId="11" pageNumber="20" precision="925" value="-10.5">10°30'S</geoCoordinate>
<geoCoordinate id="A2C768FE3E5BA15079CBFEDCC73DFECB" box="[1211,1293,329,353]" degrees="31" direction="east" minutes="12" orientation="longitude" pageId="11" pageNumber="20" precision="925" value="31.2">31°12'E</geoCoordinate>
,
<quantity id="000BA3DC3E5BA1507851FEDCC742FECB" box="[1313,1394,329,353]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="20" unit="m" value="1250.0">
<elevation id="4CDEE90A3E5BA1507851FEDCC742FECB" box="[1313,1394,329,353]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="20" unit="m" value="1250.0">1250 m</elevation>
</quantity>
,
<date id="B34D28F93E5BA1507E6AFEFCC1BEFE2B" box="[794,910,361,385]" pageId="11" pageNumber="20" value="2009-12-12">
<collectingDate id="A309D1113E5BA1507E6AFEFCC1BEFE2B" box="[794,910,361,385]" pageId="11" pageNumber="20" value="2009-12-12">12.12.2009</collectingDate>
</date>
,
<specimenCount id="D1F5C5B03E5BA1507EE8FEFCC18DFE2B" box="[920,957,361,385]" pageId="11" pageNumber="20" type="female">1 ♀</specimenCount>
,
<collectorName id="6A066BEF3E5BA1507EB9FEFFC608FE2B" box="[969,1080,362,385]" pageId="11" pageNumber="20">F. Kantner</collectorName>
lg.
<collectorName id="6A066BEF3E5BA1507901FEFFC6B8FE2B" box="[1137,1160,362,385]" pageId="11" pageNumber="20">In</collectorName>
coll.
<collectionCode id="A1E296FC3E5BA15079B1FEFCC734FE2B" box="[1217,1284,361,385]" pageId="11" pageNumber="20">OLBL</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507E6AFE3CC1E6FDAB" blockId="11.[794,1399,425,513]" pageId="11" pageNumber="20">
<emphasis id="F587D22B3E5BA1507E6AFE3CC1AAFE6B" bold="true" box="[794,922,425,449]" pageId="11" pageNumber="20">Flower visit</emphasis>
: On flowers of
<taxonomicName id="00F375BA3E5BA1507930FE3FC18BFE4B" class="Magnoliopsida" family="Hypericaceae" genus="Harungana" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="20" phylum="Tracheophyta" rank="species" species="madagascariensis">
<emphasis id="F587D22B3E5BA1507930FE3FC747FE6B" box="[1088,1399,425,449]" italics="true" pageId="11" pageNumber="20">Harungana madagascariensis</emphasis>
(Hypericaceae)
</taxonomicName>
and
<taxonomicName id="00F375BA3E5BA1507E80FE5CC645FE4B" box="[1008,1141,457,481]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="11" pageNumber="20" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
; all records from
<collectingCountry id="BFE44EA93E5BA150785EFE5CC17DFDAB" name="Cameroon" pageId="11" pageNumber="20">Cameroon</collectingCountry>
by A. PAULY.
</paragraph>
</subSubSection>
<subSubSection id="8FE95DB23E5BA1507E6AFDBCC6F7FCCB" pageId="11" pageNumber="20" type="etymology">
<paragraph id="C74C0E393E5BA1507E6AFDBCC6F7FCCB" blockId="11.[794,1399,553,865]" pageId="11" pageNumber="20">
<emphasis id="F587D22B3E5BA1507E6AFDBCC1DBFDEB" bold="true" box="[794,1003,553,577]" pageId="11" pageNumber="20">Derivatio nominis</emphasis>
: The new species is dedicated to members of the Halada family (Hamr near Chlum u Třeboně,
<collectingCountry id="BFE44EA93E5BA1507E0DFDFCC618FD2B" box="[893,1064,617,641]" name="Czech Republic" pageId="11" pageNumber="20">Czech Republic</collectingCountry>
). Jiři Halada, the senior, and his sons Marek and Michal collected sometimes together with other Czech partners extensive material of
<taxonomicName id="00F375BA3E5BA1507852FD3CC746FD6B" box="[1314,1398,681,705]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="F587D22B3E5BA1507852FD3CC746FD6B" box="[1314,1398,681,705]" italics="true" pageId="11" pageNumber="20">Hylaeus</emphasis>
</taxonomicName>
from numerous African countries and provided much of their collection to the Biology Centre of the Oberösterreichisches Landesmuseum, Linz (
<collectingCountry id="BFE44EA93E5BA15079F8FC9CC6EDFC8B" box="[1160,1245,777,801]" name="Austria" pageId="11" pageNumber="20">Austria</collectingCountry>
). The material was collected with expertise, the specimens are excellently prepared it is a joy to work with.
</paragraph>
</subSubSection>
<subSubSection id="8FE95DB23E5BA1507E6AFC1CC746F788" pageId="11" pageNumber="20" type="discussion">
<paragraph id="C74C0E393E5BA1507E6AFC1CC15EFB0B" blockId="11.[794,1399,905,2082]" pageId="11" pageNumber="20">
<emphasis id="F587D22B3E5BA1507E6AFC1CC1A2FC0B" bold="true" box="[794,914,905,929]" pageId="11" pageNumber="20">Discussion</emphasis>
: This species was discovered only in the wake of a comprehensive revision of the subgenus
<taxonomicName id="00F375BA3E5BA1507852FC3CC641FC4B" authority="(DATHE 2014)" baseAuthorityName="DATHE" baseAuthorityYear="2014" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Deranchylaeus">
<emphasis id="F587D22B3E5BA1507852FC3CC746FC6B" box="[1314,1398,937,961]" italics="true" pageId="11" pageNumber="20">Hylaeus</emphasis>
(
<emphasis id="F587D22B3E5BA1507E53FC5CC18DFC4B" box="[803,957,969,993]" italics="true" pageId="11" pageNumber="20">Deranchylaeus</emphasis>
) (
<bibRefCitation id="A36273C83E5BA1507EABFC5CC65AFC4B" author="DATHE, H. H." box="[987,1130,969,993]" pageId="11" pageNumber="20" pagination="1 - 84" refId="ref13161" refString="DATHE, H. H. 2014: Studies on the systematics and taxonomy of the genus Hylaeus F. (8). Revision of the Afrotropic subgenus Hylaeus (Deranchylaeus) BRIDWELL (Hymenoptera: Anthophila, Colletidae). - Zootaxa 3874 (1): 1 - 84." type="journal article" year="2014">DATHE 2014</bibRefCitation>
)
</taxonomicName>
. Although it had been represented in the checked material, it was not recognized because of its strikingly coarse sculpture on head and mesosoma and was initially assigned to other subgenera for which such features are characteristic. These include in particular the
<taxonomicName id="00F375BA3E5BA1507EBCFBFCC610FB2B" box="[972,1056,1129,1153]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="F587D22B3E5BA1507EBCFBFCC610FB2B" box="[972,1056,1129,1153]" italics="true" pageId="11" pageNumber="20">Hylaeus</emphasis>
</taxonomicName>
subgenera
<taxonomicName id="00F375BA3E5BA15079E8FBFCC732FB2B" box="[1176,1282,1129,1153]" class="Insecta" family="Colletidae" genus="Metylaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="F587D22B3E5BA15079E8FBFCC732FB2B" box="[1176,1282,1129,1153]" italics="true" pageId="11" pageNumber="20">Metylaeus</emphasis>
</taxonomicName>
and
<taxonomicName id="00F375BA3E5BA1507847FBFCC159FB0B" authorityName="SNELLING" authorityYear="1985" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="15" phylum="Arthropoda" rank="subGenus" subGenus="Alfkenylaeus">
<emphasis id="F587D22B3E5BA1507847FBFCC159FB0B" italics="true" pageId="11" pageNumber="20">Alfkenylaeus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C74C0E393E5BA1507E6AFB3CC1EDF92B" blockId="11.[794,1399,905,2082]" pageId="11" pageNumber="20">
The new species is consistent with
<taxonomicName id="00F375BA3E5BA15079EAFB3CC768FB6B" box="[1178,1368,1193,1217]" class="Insecta" family="Colletidae" genus="Metylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="F587D22B3E5BA15079EAFB3CC734FB6B" box="[1178,1284,1193,1217]" italics="true" pageId="11" pageNumber="20">Metylaeus</emphasis>
species
</taxonomicName>
in the sculpture of the pro- and mesosoma, the anteriorly sharp and laterally tapered pronotum, a margined omaulus and sharp margins around the terminal area of the propodeum; in contrast,
<taxonomicName id="00F375BA3E5BA150794BFABFC6FFFAEB" authority="Dathe, 2015" authorityName="Dathe" authorityYear="2015" box="[1083,1231,1321,1345]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="species" species="haladanius" status="sp. nov.">
<emphasis id="F587D22B3E5BA150794BFABFC67FFAEB" box="[1083,1103,1322,1345]" italics="true" pageId="11" pageNumber="20">H</emphasis>
.
<emphasis id="F587D22B3E5BA150792BFABCC6FFFAEB" box="[1115,1231,1321,1345]" italics="true" pageId="11" pageNumber="20">haladanius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EEB46F503E5BA15079ADFABFC775FAEB" box="[1245,1349,1322,1345]" pageId="11" pageNumber="20" rank="species">spec. nov.</taxonomicNameLabel>
has no occipital carina, the metanotum is evenly rounded and punctate and without any remarkable callosities or projections. The male terminalia (
<figureCitation id="5FC812BC3E5BA15079D1FA1CC6DBFA0B" box="[1185,1259,1417,1441]" captionStart="Figs 2225" captionStartId="16.[320,360,1980,2002]" captionTargetBox="[369,1274,276,1948]" captionTargetId="figure-102@16.[320,1305,232,1971]" captionTargetPageId="16" captionText="Figs 2225: Male terminalia: metasomal sterna 7 and 8 (S7, S8), genital capsule. 22 Hylaeus (Alfkenylaeus) euphorbiae spec. nov. 23 Hylaeus (Cornylaeus) adamauanis spec. nov. 24 Hylaeus (Deranchylaeus) haladanius spec. nov. 25 Hylaeus (Pumilaeus) pumilus spec. nov., S7: left ventral view, right dorsal view. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5370726" httpUri="https://zenodo.org/record/5370726/files/figure.png" pageId="11" pageNumber="20">Fig. 24</figureCitation>
) are fundamentally different from
<taxonomicName id="00F375BA3E5BA1507957FA3CC6A1FA6B" box="[1063,1169,1449,1473]" class="Insecta" family="Colletidae" genus="Metylaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="F587D22B3E5BA1507957FA3CC6A1FA6B" box="[1063,1169,1449,1473]" italics="true" pageId="11" pageNumber="20">Metylaeus</emphasis>
</taxonomicName>
. They appear rather similar to
<taxonomicName id="00F375BA3E5BA1507EFCFA5FC604FA4B" box="[908,1076,1481,1505]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="species" species="xanthostoma">
<emphasis id="F587D22B3E5BA1507EFCFA5FC604FA4B" box="[908,1076,1481,1505]" italics="true" pageId="11" pageNumber="20">H. xanthostoma</emphasis>
</taxonomicName>
as a reduced variant. Within the subgenus
<taxonomicName id="00F375BA3E5BA1507ED9FA7CC673F9AB" baseAuthorityName="DATHE" baseAuthorityYear="2014" box="[937,1091,1513,1537]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Deranchylaeus">
<emphasis id="F587D22B3E5BA1507ED9FA7CC673F9AB" box="[937,1091,1513,1537]" italics="true" pageId="11" pageNumber="20">Deranchylaeus</emphasis>
</taxonomicName>
the new species more closely resembles
<taxonomicName id="00F375BA3E5BA1507EF9F99FC601F98B" box="[905,1073,1545,1569]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="species" species="xanthostoma">
<emphasis id="F587D22B3E5BA1507EF9F99FC1ADF98B" box="[905,925,1546,1569]" italics="true" pageId="11" pageNumber="20">H</emphasis>
.
<emphasis id="F587D22B3E5BA1507ED9F99CC601F98B" box="[937,1073,1545,1569]" italics="true" pageId="11" pageNumber="20">xanthostoma</emphasis>
</taxonomicName>
, because of its similarly coarse thoracic sculpture, at least in the Western populations, for which a larger geographical variation was observed (
<taxonomicName id="00F375BA3E5BA1507E53F9FCC1E0F92B" authority="Dathe, 2015" authorityName="Dathe" authorityYear="2015" box="[803,976,1641,1665]" pageId="11" pageNumber="10" rank="subGenus" status="subgen. nov." subGenus="Dathe">
<bibRefCitation id="A36273C83E5BA1507E53F9FCC1E0F92B" author="DATHE, H. H." box="[803,976,1641,1665]" pageId="11" pageNumber="20" pagination="1 - 84" refId="ref13161" refString="DATHE, H. H. 2014: Studies on the systematics and taxonomy of the genus Hylaeus F. (8). Revision of the Afrotropic subgenus Hylaeus (Deranchylaeus) BRIDWELL (Hymenoptera: Anthophila, Colletidae). - Zootaxa 3874 (1): 1 - 84." type="journal article" year="2014">DATHE 2014: 76</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="C74C0E393E5BA1507E6AF91CC746F788" blockId="11.[794,1399,905,2082]" pageId="11" pageNumber="20">
The present example shows again quite well the mosaic-like distribution of character complexes, which is apparently characteristic of Afrotropical
<taxonomicName id="00F375BA3E5BA1507851F95CC15DF8AB" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="F587D22B3E5BA1507851F95CC746F94B" box="[1313,1398,1737,1761]" italics="true" pageId="11" pageNumber="20">Hylaeus</emphasis>
species.
</taxonomicName>
There are few characters that separate individual groups. The sickle-shaped mandibles of
<emphasis id="F587D22B3E5BA1507870F89CC747F88B" box="[1280,1399,1801,1825]" italics="true" pageId="11" pageNumber="20">Nothylaeus</emphasis>
are a rare example But they are probably the exception, because even the stylus formation of the gonoforcipes of the male genital capsule can be found in similar forms: even in
<taxonomicName id="00F375BA3E5BA1507E1EF81CC1C6F80B" authorityName="SNELLING" authorityYear="1985" box="[878,1014,1929,1953]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="15" phylum="Arthropoda" rank="subGenus" subGenus="Alfkenylaeus">
<emphasis id="F587D22B3E5BA1507E1EF81CC1C6F80B" box="[878,1014,1929,1953]" italics="true" pageId="11" pageNumber="20">Alfkenylaeus</emphasis>
</taxonomicName>
and
<taxonomicName id="00F375BA3E5BA150795BF81CC6F8F80B" baseAuthorityName="DATHE" baseAuthorityYear="2014" box="[1067,1224,1929,1953]" class="Insecta" family="Colletidae" genus="Hylaeus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Deranchylaeus">
<emphasis id="F587D22B3E5BA150795BF81CC6F8F80B" box="[1067,1224,1929,1953]" italics="true" pageId="11" pageNumber="20">Deranchylaeus</emphasis>
</taxonomicName>
species. Thornlike structures are common in
<emphasis id="F587D22B3E5BA1507903F83CC6DBF86B" box="[1139,1259,1961,1985]" italics="true" pageId="11" pageNumber="20">Cornylaeus</emphasis>
(metasoma) and
<taxonomicName id="00F375BA3E5BA1507E3DF85CC189F84B" box="[845,953,1993,2017]" class="Insecta" family="Colletidae" genus="Metylaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="F587D22B3E5BA1507E3DF85CC189F84B" box="[845,953,1993,2017]" italics="true" pageId="11" pageNumber="20">Metylaeus</emphasis>
</taxonomicName>
(mesosoma), but they may be missing even in individuals of the same species. Correspondingly, it is difficult to write straightforward and yet
</paragraph>
</subSubSection>
</treatment>
</document>