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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="3D04C9134C3357AF827C326761DF8161" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Pax ellipita Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="D52287DC-6ED8-4E56-B435-F5CB56650469" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamani, Alireza</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marusik, Yuri M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1035</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>145</mods:start>
<mods:end>193</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
<mods:identifier type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
<mods:identifier type="Pensoft-UUID">55CAE096B4A451D88EA433175454DA3F</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181753571" LSID="urn:lsid:zoobank.org:act:D52287DC-6ED8-4E56-B435-F5CB56650469" httpUri="http://treatment.plazi.org/id/3D04C9134C3357AF827C326761DF8161" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/D52287DC-6ED8-4E56-B435-F5CB56650469" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita" status="sp. nov.">Pax ellipita</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">Figs 5A, B</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">, 7</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">, 10A-E</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Kermanshah Province</emphasis>
: north of Kermanshah,
<geoCoordinate degrees="34" direction="north" minutes="28" orientation="latitude" precision="925" value="34.466667">34°28'N</geoCoordinate>
,
<geoCoordinate degrees="47" direction="east" minutes="00" orientation="longitude" precision="925" value="47.0">47°00'E</geoCoordinate>
, 18.06.1975 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation ID-GBIF-Occurrence="3093634308" collectingDate="1977-04-11" collectorName="B. Prazan" country="Iran" latitude="32.85" location="river Saimareh" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" specimenCount="1" specimenCount-female="1" stateProvince="Lorestan Province">
<collectingCountry name="Iran">Iran</collectingCountry>
:
<emphasis italics="true" pageId="0" pageNumber="145">
<collectingRegion country="Iran" name="Lorestan">Lorestan Province</collectingRegion>
</emphasis>
:
<specimenCount type="female">1♀</specimenCount>
(NMP),
<collectingMunicipality>Pol-e Tang</collectingMunicipality>
,
<locationDeviation location="Andimeshk">
<quantity metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
NW of Andimeshk
</locationDeviation>
,
<geoCoordinate degrees="32" direction="north" minutes="51" orientation="latitude" precision="925" value="32.85">32°51'N</geoCoordinate>
,
<geoCoordinate degrees="47" direction="east" minutes="56" orientation="longitude" precision="925" value="47.933334">47°56'E</geoCoordinate>
, near the
<location LSID="urn:lsid:plazi:treatment:3D04C9134C3357AF827C326761DF8161:71B04B1A5985BF7FCAC1BD1C1C6158A6" country="Iran" latitude="32.85" longLatPrecision="1208" longitude="47.933334" municipality="Pol-e Tang" name="river Saimareh" stateProvince="Lorestan Province">river Saimareh</location>
,
<collectingDate value="1977-04-11">11.04.1977</collectingDate>
(
<collectorName>
B.
<normalizedToken originalValue="Pražan">Prazan</normalizedToken>
</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to Ellipi, an ancient kingdom located on the western side of the Zagros Mountains, between Babylonia at the west, Media at the north-east, Mannae at the north and Elam at the south.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
The new species differs from
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName>
sp. nov., the only other
<taxonomicName authorityName="Levy" authorityYear="1990" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pax</emphasis>
</taxonomicName>
species known from Iran, by the cymbium lacking a horn-like outgrowth (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
present), by having a deep lateral fold of the cymbium (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
lacking) and the epigyne with a rectangular median plate (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
triangular). The male of
<taxonomicName lsidName="P. ellipita" pageId="0" pageNumber="145" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">P. ellipita</emphasis>
</taxonomicName>
sp. nov. differs from the rest of the species by having the RTA shorter than the tibia (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer), while the female differs by the trilobate posterior margin of the epigyne (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
solid) and copulatory openings located posteriorly (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
anteriorly).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5B</figureCitation>
. Total length 5.07. Carapace 2.43 long, 1.27 wide at pars cephalica, 1.65 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.11, ALE: 0.10, PME: 0.08, PLE: 0.10, PME-PME: 0.09. Carapace, sternum, labium, chelicerae and maxillae reddish brown, without any pattern. Chelicera with 2 promarginal teeth. Legs yellow, with few spines and without annulations. Abdomen dark brown, dorsally with large scutum covering almost the entire abdomen. Spinnerets pale, uniform in color. Measurements of legs: I: 5.49 (1.48, 0.62, 1.22, 1.10, 1.07), II: 4.88 (1.29, 0.60, 1.04, 1.05, 0.90), III: 4.74 (1.27, 0.61, 0.90, 1.19, 0.77), IV: 6.71 (1.80, 0.68, 1.39, 1.80, 1.04).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 5.</emphasis>
Dorsal habitus of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A, B</emphasis>
) and
<taxonomicName lsidName="P. leila" pageId="0" pageNumber="145" rank="species" species="leila">
<emphasis italics="true" pageId="0" pageNumber="145">P. leila</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
females
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis>
males.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male palp of Pax ellipita sp. nov. A ventral B, C retrolateral D, E retrodorsal and prolateral. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145">6A-E</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM images of the male palp of Pax ellipita sp. nov. A retrolateral B anteroretrolateral C proventral D posterodorsal E ventral. Abbreviations: E 1, E 2 - pro- and retrolateral cymbial extensions, Eb - embolar base, Em - embolus, Cf - cymbial fold, Co - conductor, Ma - median apophysis, Ra - retrolateral apophysis, Va - ventral apophysis. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145">7A-E</figureCitation>
. Femur slightly shorter than cymbium, 3.3 times longer than wide; patella globular; tibia wider than long with ventral apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
) and short RTA bifurcated at the tip (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
); cymbium with large fold (
<emphasis italics="true" pageId="0" pageNumber="145">Cf</emphasis>
) and 2 extensions, pro- and retrolateral (
<emphasis italics="true" pageId="0" pageNumber="145">E1</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="145">E2</emphasis>
); bulb with large lamella, as long as bulb, tapering to the tip; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) long, about 5 times longer than wide; conductor (
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
) heavily sclerotized; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
) long, originates at 6
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
position.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure6" httpUri="https://binary.pensoft.net/fig/536883" pageId="0" pageNumber="145" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 6.</emphasis>
Male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, C</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">D, E</emphasis>
retrodorsal and prolateral. Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure7" httpUri="https://binary.pensoft.net/fig/536884" pageId="0" pageNumber="145" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 7.</emphasis>
SEM images of the male palp of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Pax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pax ellipita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="ellipita">
<emphasis italics="true" pageId="0" pageNumber="145">Pax ellipita</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
anteroretrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
proventral
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
posterodorsal
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
ventral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">E1</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="145">E2</emphasis>
- pro- and retrolateral cymbial extensions,
<emphasis italics="true" pageId="0" pageNumber="145">Eb</emphasis>
- embolar base,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Cf</emphasis>
- cymbial fold,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus of Pax ellipita sp. nov. (A, B) and P. leila sp. nov. (C, D) A, D females B, C males." figureDoi="10.3897/zookeys.1035.65767.figure5" httpUri="https://binary.pensoft.net/fig/536882" pageId="0" pageNumber="145">5A</figureCitation>
. Total length 5.33. Carapace 2.18 long, 1.08 wide at pars cephalica, 1.51 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.06, ALE: 0.09, PME: 0.07, PLE: 0.09, PME-PME: 0.08. Coloration as in male, with lighter abdomen lacking a scutum. Measurements of legs: I: 5.36 (1.52, 0.60, 1.15, 1.09, 1.00), II: 4.76 (1.32, 0.61, 0.99, 1.04, 0.80), III: 4.53 (1.23, 0.60, 0.89, 1.11, 0.70), IV: 5.78 (1.42, 0.65, 1.38, 1.53, 0.80).
</paragraph>
<paragraph pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epigynes of Pax ellipita sp. nov. (A-E) and P. leila sp. nov. (F-J) A, F intact, ventral B, J posterior C, G macerated, ventral D, E, H, I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure10" httpUri="https://binary.pensoft.net/fig/536887" pageId="0" pageNumber="145">10A-E</figureCitation>
. Trilobate, wider than long; median plate rectangular (with subparallel lateral margins), ca. 3.6 times thinner than lateral lobe; receptacles oval, wider than long, touching each other.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comments">
<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
<paragraph pageId="0" pageNumber="145">Because the female specimen was not collected together with the male, its assignment to this species is tentative and shall be confirmed when both sexes are collected together.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
Known from the listed localities in Kermanshah and Lorestan provinces, western Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>