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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1191.110217" ID-Pensoft-Pub="1313-2970-1191-23" ID-Pensoft-UUID="D21A11D2AFBB5AD4B38BB1B9EB59704E" ID-ZooBank="863317785C0C448FBDAAF81AE79F4DB7" ModsDocID="1313-2970-1191-23" checkinTime="1707311028507" checkinUser="pensoft" docAuthor="Cabras, Analyn A., Buenavente, Perry Archival C. &amp; Medina, Milton Norman" docDate="2024" docId="17F2C49A4301512DBD569D3194D609AD" docLanguage="en" docName="ZooKeys 1191: 23-33" docOrigin="ZooKeys 1191" docPubDate="2024-02-06" docSource="http://dx.doi.org/10.3897/zookeys.1191.110217" docTitle="Enoplocyrtus angelalcalai Cabras 2024, sp. nov." docType="treatment" docUuid="D305F11A-D77C-4BB4-A2B3-E629F52C3971" docUuidSource="ZooBank" docVersion="1" id="D21A11D2AFBB5AD4B38BB1B9EB59704E" lastPageNumber="23" masterDocId="D21A11D2AFBB5AD4B38BB1B9EB59704E" masterDocTitle="New species in the genera Eumacrocyrtus Schultze, 1923 and Enoplocyrtus Yoshitake, 2017 from Luzon Island, Philippines (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini)" masterLastPageNumber="33" masterPageNumber="23" pageNumber="23" updateTime="1707311028507" updateUser="pensoft">
<mods:mods id="6DCF271AB68074765D3B66966C334AFE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="65628512CD41D3178F34BC5605D10355">
<mods:title id="F6E3043794D3D34B6DF500BC71125736">New species in the genera Eumacrocyrtus Schultze, 1923 and Enoplocyrtus Yoshitake, 2017 from Luzon Island, Philippines (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini)</mods:title>
</mods:titleInfo>
<mods:name id="F5751EDD5D32E97F63EA8E6A5059CB93" type="personal">
<mods:role id="71A95C729AD2ABE1D4E36C578716905B">
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<mods:namePart id="693EB16A5348F1C33D98CC7F10E59BA9">Cabras, Analyn A.</mods:namePart>
<mods:nameIdentifier id="EBB0D5139E55FF5BC86EE825DA8F10CC" type="ORCID">https://orcid.org/0000-0002-0980-1651</mods:nameIdentifier>
<mods:affiliation id="3F4F08D7F2CBFE11A2ADD0B1B6E86D11">Institute of Agriculture and Life Sciences, Davao Oriental State University, Dahican, City of Mati 8200, Philippines &amp; Terrestrial Invertebrate Research Laboratory, Davao Oriental State University, Dahican, City of Mati 8200, Philippines</mods:affiliation>
<mods:nameIdentifier id="1DBA92750F23BF3EC35DD6569EF4A61B" type="email">ann.cabras24@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="A9C89EF1DF94C505AA0EA45E35EB02DC" type="personal">
<mods:role id="9D7F096E7D72BE3AFC5255C2DA4E99F1">
<mods:roleTerm id="424F414D2AC5AB00EB481A29515FA631">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA31F795A003475C6EE22C039F00A845">Buenavente, Perry Archival C.</mods:namePart>
<mods:nameIdentifier id="207A572C1C07063AA8DF67DD1A3E8D6A" type="ORCID">https://orcid.org/0000-0001-8353-6019</mods:nameIdentifier>
<mods:affiliation id="1168CE71C8ABFC9E4DAF0C2BD4FB3CAA">Terrestrial Invertebrate Research Laboratory, Davao Oriental State University, Dahican, City of Mati 8200, Philippines</mods:affiliation>
</mods:name>
<mods:name id="B7D0DC3C309E99F7C5A20816FB97FD3A" type="personal">
<mods:role id="B868E553B56DEFCFD528C721E77521C0">
<mods:roleTerm id="728ED2D32ECA9859FF0A969A3DBD45AD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36A01D2B1E7C008C7FC5993A99CC32FD">Medina, Milton Norman</mods:namePart>
<mods:nameIdentifier id="C228F2982EA2F7C8B597BC6CF11637F6" type="ORCID">https://orcid.org/0000-0001-6858-8048</mods:nameIdentifier>
<mods:affiliation id="5A795B4A021AEC4B04C64A0614884DC8">Institute of Agriculture and Life Sciences, Davao Oriental State University, Dahican, City of Mati 8200, Philippines &amp; Terrestrial Invertebrate Research Laboratory, Davao Oriental State University, Dahican, City of Mati 8200, Philippines</mods:affiliation>
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<mods:title id="6C0EBEDDD0EB86DFB504842A9E42A93D">ZooKeys</mods:title>
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<mods:part id="898CF9F8686EB58B96EC6364CCB8FC8B">
<mods:date id="382E7B2AF86A97807AB8D5AA934622EF">2024</mods:date>
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<mods:number id="E82911A46C12956AE6D8652D4FC005BD">2024-02-06</mods:number>
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<mods:detail id="7CE1EC935E51771359BBF4B3096C0EB8" type="volume">
<mods:number id="02423076B6F22AE9B3F84A4E8C3B2FCE">1191</mods:number>
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<mods:start id="8FFEA4D0BD32341DE6D23CE5877731BD">23</mods:start>
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<mods:url id="1B7AEFA07E1F867BAAB45839EAFA2DA0">http://dx.doi.org/10.3897/zookeys.1191.110217</mods:url>
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<mods:classification id="62C1C99CA67E44EF300EC864C074B5A1">journal article</mods:classification>
<mods:identifier id="243EF7215684E5D34C6289B05121CD0D" type="DOI">http://dx.doi.org/10.3897/zookeys.1191.110217</mods:identifier>
<mods:identifier id="CD23F24F8FD9DCB5EF844F082E823257" type="Pensoft-Pub">1313-2970-1191-23</mods:identifier>
<mods:identifier id="DDF0160379A4AA87304BF84BFE7EC38A" type="ZooBank">863317785C0C448FBDAAF81AE79F4DB7</mods:identifier>
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<subSection id="0FBCF994CFD71D0A2F4705554A14CBCC" pageId="0" pageNumber="23" type="taxonomy">
<treatment id="17F2C49A4301512DBD569D3194D609AD" LSID="urn:lsid:zoobank.org:act:D305F11A-D77C-4BB4-A2B3-E629F52C3971" httpUri="http://treatment.plazi.org/id/17F2C49A4301512DBD569D3194D609AD" lastPageNumber="23" pageId="0" pageNumber="23">
<subSubSection id="26C9F94F1CD8F0A441C59C7191B31C7A" pageId="0" pageNumber="23" type="nomenclature">
<paragraph id="E66C30B44B7AC9E0BA44E94CC9090D8C" pageId="0" pageNumber="23">
<taxonomicName id="F2F75C489950C6A9E7A50F658FF3CEC0" LSID="https://zoobank.org/D305F11A-D77C-4BB4-A2B3-E629F52C3971" authority="Cabras" authorityName="Cabras" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Enoplocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enoplocyrtus angelalcalai" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="angelalcalai" status="sp. nov.">Enoplocyrtus angelalcalai Cabras</taxonomicName>
<taxonomicNameLabel id="F56975C45177E67166F3A0D24D19C780" pageId="0" pageNumber="23">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="AE0A06C462D7FD5D9D540582B42370C2" pageId="0" pageNumber="23" type="description">
<paragraph id="20620EF2AB84E67D44DFD1D236738E84" pageId="0" pageNumber="23">
<figureCitation id="C0F89FBE89844F431B359233930A6DA1" captionStart="Figures 11–14" captionStartId="F4" captionText="Figures 11 - 14. Enoplocyrtus angelalcalai sp. nov. 11 male holotype, dorsal habitus 12 female paratype, dorsal habitus 13 male, lateral habitus 14 female, lateral habitus." figureDoi="10.3897/zookeys.1191.110217.figures11-14" httpUri="https://binary.pensoft.net/fig/980424" pageId="0" pageNumber="23">Figs 11-14</figureCitation>
<figureCitation id="C475CF64529DA1C5A0982CB88DD491E9" captionStart="Figures 15–20" captionStartId="F5" captionText="Figures 15 - 20. Male and female genitalia of Enoplocyrtus angelalcalai sp. nov. 15 median lobe in dorsal view 16 idem in lateral view 17 sternite IX in dorsal view 18 sternite VIII in ventral view 19 ovipositor in dorsal view 20 spermatheca." figureDoi="10.3897/zookeys.1191.110217.figures15-20" httpUri="https://binary.pensoft.net/fig/980425" pageId="0" pageNumber="23">, 15-20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D7EFB1C6D7B4FF3D693EB317F1D11C16" pageId="0" pageNumber="23" type="type material">
<paragraph id="706B864FEFD6A38B3E615BCB74C939DF" pageId="0" pageNumber="23">Type materials.</paragraph>
<paragraph id="1786621BD6C7DD0E02036064807A92F0" pageId="0" pageNumber="23">
<emphasis id="019515F917C0F99A1E6AE5721AD9464C" bold="true" italics="true" pageId="0" pageNumber="23">Holotype</emphasis>
(Figs
<figureCitation id="9830BFA547E2C7136FB7864D656F2DBC" captionStart="Figures 11–14" captionStartId="F4" captionText="Figures 11 - 14. Enoplocyrtus angelalcalai sp. nov. 11 male holotype, dorsal habitus 12 female paratype, dorsal habitus 13 male, lateral habitus 14 female, lateral habitus." figureDoi="10.3897/zookeys.1191.110217.figures11-14" httpUri="https://binary.pensoft.net/fig/980424" pageId="0" pageNumber="23">11</figureCitation>
,
<figureCitation id="685D101925E9F439AA3FADAEAF9661C4" captionStart="Figures 11–14" captionStartId="F4" captionText="Figures 11 - 14. Enoplocyrtus angelalcalai sp. nov. 11 male holotype, dorsal habitus 12 female paratype, dorsal habitus 13 male, lateral habitus 14 female, lateral habitus." figureDoi="10.3897/zookeys.1191.110217.figures11-14" httpUri="https://binary.pensoft.net/fig/980424" pageId="0" pageNumber="23">13</figureCitation>
), male: &quot;Philippines- Luzon Island, Mountain Province, Bontoc, Palapal / July, 1985/ leg. Samarita (typed on white card) // HOLOTYPE male /
<taxonomicName id="6974D293426302367C77DD62E21DD310" authorityName="Cabras" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Enoplocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enoplocyrtus angelalcalai" order="Vitales" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="angelalcalai">
<emphasis id="E4593EDC367A8B2712D796FD4CDD4FE4" italics="true" pageId="0" pageNumber="23">Enoplocyrtus angelalcalai</emphasis>
</taxonomicName>
sp. nov./ CABRAS, 2024 (typed on red card)&quot;(PNM 15218).
<emphasis id="E71EB0851ADDA286E7BE67DA1C037C65" bold="true" italics="true" pageId="0" pageNumber="23">Paratypes</emphasis>
1 ♀: same data as the holotype (PNM 15219).
</paragraph>
<caption id="06F79A5C7443421603E667F5B12A9DFD" doi="10.3897/zookeys.1191.110217.figures11-14" httpUri="https://binary.pensoft.net/fig/980424" pageId="0" pageNumber="23" start="Figures 11–14" startId="F4">
<paragraph id="872E892DA1B74992D619A60BECDFCE04" pageId="0" pageNumber="23">
<emphasis id="34513F07866C199ECDAB2150219BBB5A" bold="true" pageId="0" pageNumber="23">Figures 11-14.</emphasis>
<taxonomicName id="97B3E86CCDA651E067334CA85B3B0D3E" authorityName="Cabras" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Enoplocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enoplocyrtus angelalcalai" order="Vitales" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="angelalcalai">
<emphasis id="5AD52AE87DBFC8653DFCE9C5ECE0ECCF" italics="true" pageId="0" pageNumber="23">Enoplocyrtus angelalcalai</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="4FA5BA1D2E79A8C7253C3D5F0E80A79B" bold="true" pageId="0" pageNumber="23">11</emphasis>
male holotype, dorsal habitus
<emphasis id="73B746C70212C688487690B2801C2A63" bold="true" pageId="0" pageNumber="23">12</emphasis>
female paratype, dorsal habitus
<emphasis id="0F39E65E8DCC20042EC6AE4A80603278" bold="true" pageId="0" pageNumber="23">13</emphasis>
male, lateral habitus
<emphasis id="EB96118EB098A586C0611979428ECDDC" bold="true" pageId="0" pageNumber="23">14</emphasis>
female, lateral habitus.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F22DCC54FE5EC8ACB0F9F06E379D59F0" pageId="0" pageNumber="23" type="description">
<paragraph id="CD304308AC3A965647C48C789A2A42C9" pageId="0" pageNumber="23">Description.</paragraph>
<paragraph id="844E8D23979503940D489D13C67306A0" pageId="0" pageNumber="23">
<emphasis id="EFA46B321B7773991F844EB6B4C4B7E7" bold="true" italics="true" pageId="0" pageNumber="23">Dimensions</emphasis>
: LB: 8.4 mm. LR: 1.1 mm. WR: 1.1 mm. LP: 2.5 mm. WP: 2.5 mm. LE: 5.8 mm. WE: 4.0 mm.
<emphasis id="BFAC3212ED28C655649F723C0E8B5245" italics="true" pageId="0" pageNumber="23">N</emphasis>
= 1. Integument dark brown. Body surface, rostrum, head, legs matte, and underside weakly lustrous.
<emphasis id="7C4F8F0DBD012708E756F15A22D65BAD" bold="true" italics="true" pageId="0" pageNumber="23">Head</emphasis>
dorsal surface weakly punctate and each puncture with subappressed white piliform scales; lateral side with weak rugae and sides below eye with sparse appressed bluish piliform scales; forehead between eyes flat with distinct but faint midline groove; eyes medium-sized, feebly convex, and moderately prominent on lateral outline of head.
<emphasis id="9861B44F3AA5591BB98442C84E6CE9B6" bold="true" italics="true" pageId="0" pageNumber="23">Rostrum</emphasis>
dorsal surface weakly rugose up to anterior two-thirds with appressed white piliform scales; apical third finely punctate with fine sparse setae; as long as wide (LR/WR:1.1 mm/1.1 mm); transverse groove at base absent; with midline groove distinct until middle and progressively fainter and indistinct towards apex; dorsal contour flat, slightly curved towards apex but without apical bulge; lateral sides at upper margin of antennal scrobe with subtriangular depression. Antennal scape slightly longer than funicle, scape reaching beyond the posterior margin of eye, with few sparse subappressed pubescence, and funicle with suberect brownish setae. Funicular segment I nearly four times as long as wide, slightly longer than II, funicular segment II nearly three times as long as wide, longer than segments III-V, segments III-V as long as wide, shorter and narrower than VI, segment VI as long as wide, longer and wider than segments III-V, segment VII as long as wide, wider and longer than segments VI; club lanceolate.
<emphasis id="CA89F56DD6E04B38B59A3CBFA8FA4C51" bold="true" italics="true" pageId="0" pageNumber="23">Prothorax</emphasis>
subcylindrical, significantly wider at posterior margin then narrowed towards anterior margin, as long as wide (LP/WP:2.5/2.5 mm), coarsely punctate and rugose with sparse appressed white piliform scales and lanceolate pale turquoise and pale blue scales, widest at base, dorsal contour flat.
<emphasis id="CEC5798D16FC404E85524911622136A7" bold="true" italics="true" pageId="0" pageNumber="23">Elytra</emphasis>
pyriform (LE/ WE:5.8/4.0 mm), moderately longer than wide; moderately longer and wider than prothorax (WE/ WP: 4.0/2.5 mm, LE/LP: 5.8/2.5 mm); elytral surface coarse and with granulately rugose intervals, sparsely covered with minute white appressed piliform scales and pale turquoise and pale blue lanceolate scales, dorsum weakly convex, lateral contour widest before middle.
<emphasis id="125AFDB157DF0F42A4267481F96B83A7" bold="true" pageId="0" pageNumber="23">
<emphasis id="CB5D90A7B396AE69914BBCDD2FDCA268" italics="true" pageId="0" pageNumber="23">Legs</emphasis>
.
</emphasis>
Mid and hind femora dark brown covered with white subappressed piliform scales. Mid tibiae covered with white subappressed piliform scales on outer edge and suberect brown setae along inner edge. Hind tibiae covered with subappressed white piliform scales on outer edge and suberect brown setae along inner edge; inner edge and part of outer edge coarsely granulated. Mid and hind tibiae mucronate. Tarsomeres pubescent. Coxae with sparse appressed pale blue and white piliform scales. Mesoventrite with sparse appressed pale blue lanceolate scales and white setae. Metaventrite and ventrite I weakly depressed on disc, mostly glabrous except distal ends with appressed pale blue and pale-yellow lanceolate scales and white piliform scales. Ventrite II with sparse pale blue and yellow lanceolate scales and white setae. Ventrite III to V with fine sparse brown setae.
</paragraph>
<paragraph id="A957F778950AEC2AB0E00ABD723B1B3D" pageId="0" pageNumber="23">
<emphasis id="51EFE242A4659F909E0C33A924BDFFFB" bold="true" pageId="0" pageNumber="23">Female</emphasis>
(PNM 15219). Dimensions: LB: 8.0 mm: LR: 1.1 mm: WR:1.1 mm.LP: 2.5 mm. WP: 2.5 mm. LE: 5.8 mm. WE: 4.1 mm.
<emphasis id="E8BA29B71ED629728A136D75A73FDAC4" italics="true" pageId="0" pageNumber="23">N</emphasis>
= 1. Habitus, as shown in Figs
<figureCitation id="07EEB2D1BEC62A2418CA12288FA16FB4" captionStart="Figures 11–14" captionStartId="F4" captionText="Figures 11 - 14. Enoplocyrtus angelalcalai sp. nov. 11 male holotype, dorsal habitus 12 female paratype, dorsal habitus 13 male, lateral habitus 14 female, lateral habitus." figureDoi="10.3897/zookeys.1191.110217.figures11-14" httpUri="https://binary.pensoft.net/fig/980424" pageId="0" pageNumber="23">12</figureCitation>
,
<figureCitation id="1638C7DB98F0D5FBB676137DA66FB5A8" captionStart="Figures 11–14" captionStartId="F4" captionText="Figures 11 - 14. Enoplocyrtus angelalcalai sp. nov. 11 male holotype, dorsal habitus 12 female paratype, dorsal habitus 13 male, lateral habitus 14 female, lateral habitus." figureDoi="10.3897/zookeys.1191.110217.figures11-14" httpUri="https://binary.pensoft.net/fig/980424" pageId="0" pageNumber="23">14</figureCitation>
. Females differ from males in the following: a) fore tibiae flat; b) pronotum narrower with more flat dorsal contour; c) elytra moderately wider, and slightly longer; d) ventrite I flat. Otherwise, it is similar to the male.
</paragraph>
</subSubSection>
<subSubSection id="1CC49AAA5E2F30F4782CF58200C02B13" pageId="0" pageNumber="23" type="diagnosis">
<paragraph id="DEB2852CDA205D2EDA22DD073E87833C" pageId="0" pageNumber="23">Diagnosis.</paragraph>
<paragraph id="19866696B6DA332DF9835BD4E8B8276B" pageId="0" pageNumber="23">
The new species is easily distinguished from the only known species,
<taxonomicName id="59393332D25510B49F3BE864892C3EEB" authorityName="Yoshitake" authorityYear="2017" class="Insecta" family="Curculionidae" genus="Enoplocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enoplocyrtus marusan" order="Vitales" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="marusan">
<emphasis id="84412CDA86558DFE39B729F227082EC4" italics="true" pageId="0" pageNumber="23">Enoplocyrtus marusan</emphasis>
</taxonomicName>
Yoshitake, 2017, based on the following morphological characteristics: a) coarsely punctate and rugose pronotum, b) elytral surface coarse and with granulately rugose intervals, c) integument dark brown, d) pronotum and elytra sparsely covered with appressed white piliform scales and pale turquoise and pale blue lanceolate scales, and e) differently shaped aedeagus (Figs
<figureCitation id="3B0455D16D3E7CC59DC6C82604D9E818" captionStart="Figures 15–20" captionStartId="F5" captionText="Figures 15 - 20. Male and female genitalia of Enoplocyrtus angelalcalai sp. nov. 15 median lobe in dorsal view 16 idem in lateral view 17 sternite IX in dorsal view 18 sternite VIII in ventral view 19 ovipositor in dorsal view 20 spermatheca." figureDoi="10.3897/zookeys.1191.110217.figures15-20" httpUri="https://binary.pensoft.net/fig/980425" pageId="0" pageNumber="23">15</figureCitation>
,
<figureCitation id="79CFC76A4D227D48468B108E8C3E2F5F" captionStart="Figures 15–20" captionStartId="F5" captionText="Figures 15 - 20. Male and female genitalia of Enoplocyrtus angelalcalai sp. nov. 15 median lobe in dorsal view 16 idem in lateral view 17 sternite IX in dorsal view 18 sternite VIII in ventral view 19 ovipositor in dorsal view 20 spermatheca." figureDoi="10.3897/zookeys.1191.110217.figures15-20" httpUri="https://binary.pensoft.net/fig/980425" pageId="0" pageNumber="23">16</figureCitation>
).
</paragraph>
<caption id="EACA281C6DBC2F730623FE9A134B6BEF" doi="10.3897/zookeys.1191.110217.figures15-20" httpUri="https://binary.pensoft.net/fig/980425" pageId="0" pageNumber="23" start="Figures 15–20" startId="F5">
<paragraph id="F825BD383F834EE480CE0162C4522355" pageId="0" pageNumber="23">
<emphasis id="DBD2192807DCA1A754615909022B76E8" bold="true" pageId="0" pageNumber="23">Figures 15-20.</emphasis>
Male and female genitalia of
<taxonomicName id="46BEB857BFB2D065DAF2D2A7AFE86D7E" authorityName="Cabras" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Enoplocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enoplocyrtus angelalcalai" order="Vitales" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="angelalcalai">
<emphasis id="F86B6DAD9427160FBFC0A447DE155DD1" italics="true" pageId="0" pageNumber="23">Enoplocyrtus angelalcalai</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="FE9827407D5459F5252BA39D0EDD1871" bold="true" pageId="0" pageNumber="23">15</emphasis>
median lobe in dorsal view
<emphasis id="7F351DA99E050956D68BF875F2FBDE28" bold="true" pageId="0" pageNumber="23">16</emphasis>
idem in lateral view
<emphasis id="9F18106A220792265B6256A34B15F0DB" bold="true" pageId="0" pageNumber="23">17</emphasis>
sternite IX in dorsal view
<emphasis id="468699FD3DC5DE60B31B86E5B88AE9E6" bold="true" pageId="0" pageNumber="23">18</emphasis>
sternite VIII in ventral view
<emphasis id="56CCE5677704C321DD34D51C22048686" bold="true" pageId="0" pageNumber="23">19</emphasis>
ovipositor in dorsal view
<emphasis id="97AAF9263B233E549529C8B95A392683" bold="true" pageId="0" pageNumber="23">20</emphasis>
spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="64533445156222D9E31A810707F2C80D" pageId="0" pageNumber="23" type="etymology">
<paragraph id="DACC363FFAB66E6C6E059D78CC0F0495" pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph id="26A623818112FF9C76B6A786C66E81BB" pageId="0" pageNumber="23">
The name &quot;
<emphasis id="DE5ED9AD0021059164D268D106F5E07B" italics="true" pageId="0" pageNumber="23">angelalcalai</emphasis>
&quot; is dedicated to National Scientist Angel Chua Alcala (1929-2023, Dumaguete, Negros Oriental, Philippines) for his contributions to advancing the
<normalizedToken id="00972CBE5BABF2C60CEB238654E539CA" originalValue="Philippines’">Philippines'</normalizedToken>
herpetological and marine research and conservation. Additionally for inspiring the authors to work on biodiversity research and conservation in the Philippines.
</paragraph>
</subSubSection>
<subSubSection id="50DF2FF489D44E77D99975762C22744D" pageId="0" pageNumber="23" type="distribution">
<paragraph id="0EDFD11F61498CD04E5277E26CE5F2D9" pageId="0" pageNumber="23">Distribution.</paragraph>
<paragraph id="F233962416ECC6FD5E6578928FA2DC66" pageId="0" pageNumber="23">
<taxonomicName id="A307E3DF96AAD18F1554FA35EE45FFB6" authorityName="Cabras" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Enoplocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enoplocyrtus angelalcalai" order="Vitales" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="angelalcalai">
<emphasis id="633B8963B8D267C0FFA47181DCA09445" italics="true" pageId="0" pageNumber="23">Enoplocyrtus angelalcalai</emphasis>
</taxonomicName>
sp. nov. is known only from the type locality in Palapag, Bontoc, Mountain Province (Fig.
<figureCitation id="4A51FB0A7F17BDFE447617F38F354AE4" captionStart="Figure 21" captionStartId="F6" captionText="Figure 21. Distribution of E. robertfoxi sp. nov. (yellow), E. canlaonensis Schultze, 1923 (red) and Enoplocyrtus angelalcalai sp. nov. (blue) in the Philippines." figureDoi="10.3897/zookeys.1191.110217.figure21" httpUri="https://binary.pensoft.net/fig/980426" pageId="0" pageNumber="23">21</figureCitation>
).
</paragraph>
<caption id="43A68FBDEAEF044838D9A51B3E0D82FD" doi="10.3897/zookeys.1191.110217.figure21" httpUri="https://binary.pensoft.net/fig/980426" pageId="0" pageNumber="23" start="Figure 21" startId="F6">
<paragraph id="1D49AFADE9BE7118298603F259491ADC" pageId="0" pageNumber="23">
<emphasis id="7746B6FB50FC51874F8FA37FB3F47598" bold="true" pageId="0" pageNumber="23">Figure 21.</emphasis>
Distribution of
<taxonomicName id="F01045D73C07C8A3EA19E8FE38206397" lsidName="E. robertfoxi" pageId="0" pageNumber="23" rank="species" species="robertfoxi">
<emphasis id="8757EC5AB734CCBAE87C5BACD2AF2DB8" italics="true" pageId="0" pageNumber="23">E. robertfoxi</emphasis>
</taxonomicName>
sp. nov. (yellow),
<taxonomicName id="AC80544A772D75808884B989C26FD8D2" lsidName="E. canlaonensis" pageId="0" pageNumber="23" rank="species" species="canlaonensis">
<emphasis id="8233BD5F9D239BA2A1E7B9872B77B306" italics="true" pageId="0" pageNumber="23">E. canlaonensis</emphasis>
</taxonomicName>
Schultze, 1923 (red) and
<taxonomicName id="C02F81FB0C1B1EE7C9B03CD88A43FB50" authorityName="Cabras" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Enoplocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enoplocyrtus angelalcalai" order="Vitales" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="angelalcalai">
<emphasis id="2476FCCEE2E08A8B4B86E3B1DFDBD781" italics="true" pageId="0" pageNumber="23">Enoplocyrtus angelalcalai</emphasis>
</taxonomicName>
sp. nov. (blue) in the Philippines.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>
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