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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7853FF93FF013E0DACBDEE05" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Alluviobolus antanosy Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n." docType="treatment" docUuid="1AAC85EB-6256-450E-B461-707D235E61BA" docUuidSource="ZooBank" docVersion="8" lastPageNumber="86" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="85" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791445" ID-GBIF-Taxon="164088184" ID-Zenodo-Dep="3791445" LSID="urn:lsid:zoobank.org:act:1AAC85EB-6256-450E-B461-707D235E61BA" httpUri="http://treatment.plazi.org/id/1772122E7853FF93FF013E0DACBDEE05" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<subSubSection box="[140,587,377,404]" pageId="84" pageNumber="85" type="nomenclature">
<paragraph blockId="84.[140,969,377,440]" box="[140,587,377,404]" pageId="84" pageNumber="85">
<heading box="[140,587,377,404]" fontSize="11" level="2" pageId="84" pageNumber="85" reason="3">
<emphasis bold="true" box="[140,587,377,404]" pageId="84" pageNumber="85">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,510,377,404]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis bold="true" box="[140,402,377,404]" italics="true" pageId="84" pageNumber="85">Alluviobolus antanosy</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[522,587,378,404]" pageId="84" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,969,413,440]" pageId="84" pageNumber="85" type="description">
<paragraph blockId="84.[140,969,377,440]" box="[140,969,413,440]" pageId="84" pageNumber="85">
<uri box="[140,969,413,440]" pageId="84" pageNumber="85">
urn:lsid:zoobank.org:act:
<uuid box="[417,969,413,440]" pageId="84" pageNumber="85">1AAC85EB-6256-450E-B461-707D235E61BA</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="materials_examined">
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[140,370,483,509]" pageId="84" pageNumber="85">
<emphasis box="[140,241,483,509]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Material</emphasis>
examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2612160309" collectingDate="2007-04-14" collectionCode="FMMC" collectorName="T. Wesener" country="Madagascar" county="Province" latitude="-25.086176" location="Grand Lavasoa" longLatPrecision="1" longitude="46.74887" municipality="Ambatotsirongorongo Mountain" pageId="84" pageNumber="85" specimenCode="W100" specimenCount="8" specimenCount-female="7" stateProvince="Toliara" typeStatus="holotype">
5 ♂,
<specimenCount box="[433,475,484,510]" pageId="84" pageNumber="85" type="female">7 ♀</specimenCount>
, 4 imm
<typeStatus box="[571,682,483,509]" pageId="84" pageNumber="85">Holotype</typeStatus>
: 1 ♂ (
<quantity box="[751,834,483,510]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="84" pageNumber="85" unit="mm" value="45.0">45 mm</quantity>
long),
<collectionCode box="[914,1004,484,510]" pageId="84" pageNumber="85">FMMC</collectionCode>
<specimenCode box="[1010,1082,484,510]" collectionCode="FMMC" pageId="84" pageNumber="85">W100</specimenCode>
A,
<collectingCountry box="[140,274,519,545]" name="Madagascar" pageId="84" pageNumber="85">Madagascar</collectingCountry>
,
<collectingCounty box="[296,394,519,545]" pageId="84" pageNumber="85">Province</collectingCounty>
<collectingRegion box="[410,492,519,545]" country="Madagascar" name="Toliara" pageId="84" pageNumber="85">Toliara</collectingRegion>
,
<collectingMunicipality box="[513,900,519,545]" pageId="84" pageNumber="85">Ambatotsirongorongo Mountain</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7853FF93FF013E0DACBDEE05:9A04F5E37853FF92FC143D73AB16E870" box="[921,1103,519,545]" country="Madagascar" county="Province" latitude="-25.086176" longLatPrecision="1" longitude="46.74887" municipality="Ambatotsirongorongo Mountain" name="Grand Lavasoa" pageId="84" pageNumber="85" stateProvince="Toliara">Grand Lavasoa</location>
,
<geoCoordinate box="[140,305,554,581]" degrees="25" direction="south" minutes="5" orientation="latitude" pageId="84" pageNumber="85" precision="1" seconds="10.23" value="-25.086176">25°5’10.23” S</geoCoordinate>
,
<geoCoordinate box="[319,502,554,581]" degrees="46" direction="east" minutes="44" orientation="longitude" pageId="84" pageNumber="85" precision="1" seconds="55.93" value="46.74887">46°44’55.93” E</geoCoordinate>
, rainforest, leg.
<collectorName box="[689,817,554,580]" pageId="84" pageNumber="85">T. Wesener</collectorName>
et al.,
<date box="[902,1035,554,580]" pageId="84" pageNumber="85" value="2007-04-14">
<collectingDate box="[902,1035,554,580]" pageId="84" pageNumber="85" value="2007-04-14">14.IV.2007</collectingDate>
</date>
.
<emphasis italics="true" pageId="84" pageNumber="85">
<typeStatus pageId="84" pageNumber="85">Paratypes</typeStatus>
:
</emphasis>
1 ♂,
<specimenCount box="[258,301,589,616]" pageId="84" pageNumber="85" type="generic">1 F.</specimenCount>
,
<collectionCode box="[311,401,589,615]" pageId="84" pageNumber="85">FMMC</collectionCode>
<specimenCode box="[407,479,589,615]" collectionCode="FMMC" pageId="84" pageNumber="85">W100</specimenCode>
A, same data as previous
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2612160310" collectingDate="2003-04-06" collectionCode="FMMC" collectorName="Sainte Luce, S &amp; T. Wesener" country="Madagascar" latitude="-24.783333" location="Province" longLatPrecision="1249" longitude="47.166668" pageId="84" pageNumber="85" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
1 ♂,
<collectionCode box="[817,910,589,615]" pageId="84" pageNumber="85">FMMC</collectionCode>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7853FF93FF013E0DACBDEE05:9A04F5E37853FF92FC153D39ACA3E836" box="[920,1018,589,615]" country="Madagascar" latitude="-24.783333" longLatPrecision="1249" longitude="47.166668" name="Province" pageId="84" pageNumber="85" stateProvince="Toliara">Province</location>
<collectingRegion box="[1022,1104,589,615]" country="Madagascar" name="Toliara" pageId="84" pageNumber="85">Toliara</collectingRegion>
,
<collectorName box="[140,298,624,651]" pageId="84" pageNumber="85">Sainte Luce, S</collectorName>
9, littoral forest on sand,
<geoCoordinate box="[574,672,624,650]" degrees="24" direction="south" minutes="47" orientation="latitude" pageId="84" pageNumber="85" precision="925" value="-24.783333">24°47’ S</geoCoordinate>
,
<geoCoordinate box="[682,782,624,651]" degrees="47" direction="east" minutes="10" orientation="longitude" pageId="84" pageNumber="85" precision="925" value="47.166668">47°10’ E</geoCoordinate>
, leg.
<collectorName box="[836,962,625,651]" pageId="84" pageNumber="85">T. Wesener</collectorName>
,
<date box="[971,1103,624,650]" pageId="84" pageNumber="85" value="2003-04-06">
<collectingDate box="[971,1103,624,650]" pageId="84" pageNumber="85" value="2003-04-06">06.IV.2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2612160307" collectingDate="1995-01-25" collectionCode="FMMC" collectorName="K. Emberton" country="Madagascar" county="Province" elevation="340" latitude="-24.436666" location="Mt. Mahermana" longLatPrecision="21" longitude="47.22028" municipality="Esetra" pageId="84" pageNumber="85" specimenCode="FMMC 13608" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
1 ♂,
<specimenCode box="[195,367,659,686]" collectionCode="FMMC" pageId="84" pageNumber="85">FMMC 13608</specimenCode>
,
<collectingCounty box="[378,476,659,685]" pageId="84" pageNumber="85">Province</collectingCounty>
<collectingRegion box="[480,562,659,685]" country="Madagascar" name="Toliara" pageId="84" pageNumber="85">Toliara</collectingRegion>
,
<collectingMunicipality box="[572,644,660,686]" pageId="84" pageNumber="85">Esetra</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7853FF93FF013E0DACBDEE05:9A04F5E37853FF92FD033DE0AC12E8FC" box="[654,843,659,686]" country="Madagascar" county="Province" latitude="-24.436666" longLatPrecision="21" longitude="47.22028" municipality="Esetra" name="Mt. Mahermana" pageId="84" pageNumber="85" stateProvince="Toliara">Mt. Mahermana</location>
,
<quantity box="[853,929,660,686]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="84" pageNumber="85" unit="m" value="340.0">
<elevation box="[853,929,660,686]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="84" pageNumber="85" unit="m" value="340.0">340 m</elevation>
</quantity>
, station 1, summit, 0° slope, frequent granite outcrops,
<geoCoordinate box="[598,738,695,721]" degrees="24" direction="south" minutes="26" orientation="latitude" pageId="84" pageNumber="85" precision="15" seconds="12" value="-24.436666">24°26’12” S</geoCoordinate>
,
<geoCoordinate box="[750,892,695,721]" degrees="47" direction="east" minutes="13" orientation="longitude" pageId="84" pageNumber="85" precision="15" seconds="13" value="47.22028">47°13’13” E</geoCoordinate>
, leg.
<collectorName box="[951,1103,695,721]" pageId="84" pageNumber="85">K. Emberton</collectorName>
,
<date box="[140,255,730,757]" pageId="84" pageNumber="85" value="1995-01-25">
<collectingDate box="[140,255,730,757]" pageId="84" pageNumber="85" value="1995-01-25">25.I.1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2612160306" collectingDate="2007-06-08" collectionCode="FMMC" collectorName="Andohahela I &amp; T. Wesener" country="Madagascar" latitude="-24.828167" location="Isaka-Ivondro Sud" longLatPrecision="1" longitude="46.866184" pageId="84" pageNumber="85" specimenCode="W049" specimenCount="2" specimenCount-female="2" stateProvince="Toliara" typeStatus="holotype">
1 ♂,
<specimenCount box="[330,375,730,756]" pageId="84" pageNumber="85" type="female">2 ♀</specimenCount>
,
<collectionCode box="[392,482,730,756]" pageId="84" pageNumber="85">FMMC</collectionCode>
<specimenCode box="[490,562,730,756]" collectionCode="FMMC" pageId="84" pageNumber="85">W049</specimenCode>
A,
<collectorName box="[597,754,730,756]" pageId="84" pageNumber="85">Andohahela I</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7853FF93FF013E0DACBDEE05:9A04F5E37853FF92FC8D3DAEAC8DE8A5" box="[768,980,730,756]" country="Madagascar" latitude="-24.828167" longLatPrecision="1" longitude="46.866184" name="Isaka-Ivondro Sud" pageId="84" pageNumber="85" stateProvince="Toliara">Isaka-Ivondro Sud</location>
, rainforest,
<geoCoordinate box="[140,291,765,791]" degrees="24" direction="south" minutes="49.690" orientation="latitude" pageId="84" pageNumber="85" precision="1" value="-24.828167">24°49.690’ S</geoCoordinate>
,
<geoCoordinate box="[304,458,765,792]" degrees="46" direction="east" minutes="51.971" orientation="longitude" pageId="84" pageNumber="85" precision="1" value="46.866184">46°51.971’ E</geoCoordinate>
, leg.
<collectorName box="[518,644,766,792]" pageId="84" pageNumber="85">T. Wesener</collectorName>
et al.,
<date box="[724,860,765,791]" pageId="84" pageNumber="85" value="2007-06-08">
<collectingDate box="[724,860,765,791]" pageId="84" pageNumber="85" value="2007-06-08">08.VI.2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2612160316" box="[872,1103,765,792]" collectionCode="ZMH" country="Madagascar" pageId="84" pageNumber="85" specimenCode="W049" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="holotype">
<specimenCount box="[872,917,765,792]" pageId="84" pageNumber="85" type="female">1 ♀</specimenCount>
,
<collectionCode box="[932,1001,766,792]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="84" pageNumber="85" type="Museum">ZMH</collectionCode>
<specimenCode box="[1010,1082,765,791]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="84" pageNumber="85" type="Museum">W049</specimenCode>
A
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2612160314" box="[140,385,800,827]" collectionCode="ZMH" country="Madagascar" pageId="84" pageNumber="85" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">same data as previous</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2612160308" box="[396,900,800,827]" collectionCode="ZMUC" country="Madagascar" pageId="84" pageNumber="85" specimenCode="W049" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="holotype">
<specimenCount box="[396,439,800,827]" pageId="84" pageNumber="85" type="female">1 ♀</specimenCount>
,
<collectionCode box="[454,542,800,826]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="84" pageNumber="85">ZMUC</collectionCode>
<specimenCode box="[549,621,800,826]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="84" pageNumber="85">W049</specimenCode>
A, same data as previous
</materialsCitation>
.
</paragraph>
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[185,501,835,861]" pageId="84" pageNumber="85">
<emphasis box="[185,257,835,861]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Other</emphasis>
material examined:
</emphasis>
<specimenCount box="[513,560,836,862]" pageId="84" pageNumber="85" type="female">2 ♀</specimenCount>
,
<specimenCount box="[579,670,835,862]" pageId="84" pageNumber="85" type="generic">4 imm.</specimenCount>
, W049A, same data as previous, but stored as vouchers at University
<collectingRegion box="[503,660,871,897]" country="Madagascar" name="Antananarivo" pageId="84" pageNumber="85">Antananarivo</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="diagnosis">
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[185,453,905,931]" pageId="84" pageNumber="85">
<emphasis box="[185,324,905,931]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Differential</emphasis>
diagnosis:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[463,593,906,932]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species">
<emphasis box="[463,593,906,932]" italics="true" pageId="84" pageNumber="85">A. antanosy</emphasis>
</taxonomicName>
differs from all congeners in the absence of male coxal processes (
<figureCitation box="[391,497,941,967]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47B</figureCitation>
). While the anterior gonopods are very similar in all species of the
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[305,439,976,1002]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis box="[305,439,976,1002]" italics="true" pageId="84" pageNumber="85">Alluviobolus</emphasis>
</taxonomicName>
(
<figureCitation box="[456,602,976,1003]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Figs 47D, F</figureCitation>
), are the posterior gonopods in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[978,1107,977,1003]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species">
<emphasis box="[978,1107,977,1003]" italics="true" pageId="84" pageNumber="85">A. antanosy</emphasis>
</taxonomicName>
rectangular apically (
<figureCitation box="[378,486,1011,1038]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47G</figureCitation>
), compared with acute telopodite tips in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[964,1108,1011,1038]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species">
<emphasis box="[964,1108,1011,1038]" italics="true" pageId="84" pageNumber="85">A. laticlavius</emphasis>
</taxonomicName>
(
<figureCitation box="[148,253,1047,1073]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="84" pageNumber="85">Fig. 44H</figureCitation>
) and in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[349,482,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species">
<emphasis box="[349,482,1047,1073]" italics="true" pageId="84" pageNumber="85">A. tsimelahy</emphasis>
</taxonomicName>
(
<figureCitation box="[497,599,1047,1074]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="84" pageNumber="85">Fig. 46L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="description">
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[185,335,1081,1107]" pageId="84" pageNumber="85">
<emphasis box="[185,330,1081,1107]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[345,493,1083,1108]" italics="true" pageId="84" pageNumber="85">Measurements</emphasis>
: males with 42–44 body rings, circa
<quantity box="[915,1042,1082,1108]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="4.7" metricValueMin="4.3" pageId="84" pageNumber="85" unit="mm" value="45.0" valueMax="47.0" valueMin="43.0">43–47 mm</quantity>
long,
<quantity box="[140,276,1117,1144]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="84" pageNumber="85" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2–3.8 mm</quantity>
wide. Females with 43 or 44 body rings,
<quantity box="[717,839,1117,1144]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" metricValueMax="6.2" metricValueMin="5.4" pageId="84" pageNumber="85" unit="mm" value="58.0" valueMax="62.0" valueMin="54.0">54–62 mm</quantity>
long, 5.0–
<quantity box="[957,1042,1117,1144]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="84" pageNumber="85" unit="mm" value="5.5">5.5 mm</quantity>
wide.
</paragraph>
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis box="[185,300,1152,1178]" italics="true" pageId="84" pageNumber="85">Coloration</emphasis>
on mesozonites dark grey, metazonites red (
<figureCitation box="[815,923,1152,1178]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47C</figureCitation>
). Rings dorsally with a slender orange stripe. Head, legs, and posterior part of telson red (
<figureCitation box="[954,1038,1187,1214]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Figs 47</figureCitation>
A–C). Antennae and anterior part of telson dark grey (
<figureCitation box="[689,831,1223,1249]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Figs 47A, C</figureCitation>
).
<emphasis box="[854,897,1223,1248]" italics="true" pageId="84" pageNumber="85">Eyes</emphasis>
with 22–25 ocelli arranged in 4 or 5 rows (
<figureCitation box="[433,539,1258,1284]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47A</figureCitation>
).
<emphasis box="[561,664,1258,1284]" italics="true" pageId="84" pageNumber="85">Antennae</emphasis>
protruding back to ring 5 (
<figureCitation box="[989,1095,1258,1284]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47A</figureCitation>
).
<emphasis box="[140,355,1293,1319]" italics="true" pageId="84" pageNumber="85">Male coxal processes</emphasis>
absent, coxa 3 of slightly conical shape (
<figureCitation box="[821,925,1293,1319]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47B</figureCitation>
).
<emphasis box="[946,1107,1293,1319]" italics="true" pageId="84" pageNumber="85">Preanal process</emphasis>
absent (
<figureCitation box="[227,329,1328,1354]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47C</figureCitation>
)
</paragraph>
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis box="[185,373,1363,1390]" italics="true" pageId="84" pageNumber="85">Anterior gonopod</emphasis>
sternite elevated into a broad triangular lobe. Tip well-rounded (
<figureCitation box="[148,255,1399,1425]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47D</figureCitation>
). Coxite and telopodite inconspicuous (
<figureCitation box="[707,807,1399,1425]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47F</figureCitation>
).
</paragraph>
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis box="[185,381,1434,1460]" italics="true" pageId="84" pageNumber="85">Posterior gonopods</emphasis>
: telopodite membranous fringe absent (
<figureCitation box="[831,933,1434,1460]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47E</figureCitation>
). Apical sclerite (
<emphasis box="[149,161,1470,1495]" italics="true" pageId="84" pageNumber="85">x</emphasis>
in
<figureCitation box="[205,310,1469,1495]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47G</figureCitation>
) wide, rectangular. Tip broadly rounded, basally elongated to a thin spine (
<figureCitation box="[214,317,1504,1530]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="84" pageNumber="85">Fig. 47E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="distribution">
<paragraph blockId="84.[140,1108,483,1672]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[185,502,1539,1565]" pageId="84" pageNumber="85">
<emphasis box="[185,336,1539,1565]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Distribution</emphasis>
and ecology
</emphasis>
: widely distributed in rainforests of Southeastern
<collectingCountry box="[140,275,1575,1601]" name="Madagascar" pageId="84" pageNumber="85">Madagascar</collectingCountry>
.
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[289,418,1575,1601]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species">
<emphasis box="[289,418,1575,1601]" italics="true" pageId="84" pageNumber="85">A. antanosy</emphasis>
</taxonomicName>
was collected in the Andohahela and Vohimena mountain chain as well as in the lowland littoral rainforest of Sainte Luce and the isolated Ambatotsirongorongo Mountain (
<figureCitation box="[486,570,1645,1671]" captionStart="Figure 43" captionStartId="77.[139,211,1099,1121]" captionTargetBox="[139,1111,687,1080]" captionTargetId="figure@77.[139,1108,688,1080]" captionTargetPageId="77" captionText="Figure 43. Distribution map of the genus Alluviobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820825" httpUri="https://zenodo.org/record/3820825/files/figure.png" pageId="84" pageNumber="85">Fig. 43</figureCitation>
). All specimens were found on the soil surface.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767336" ID-Zenodo-Dep="3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="85" pageNumber="86" startId="85.[140,212,825,847]" targetBox="[140,1108,165,794]" targetPageId="85">
<paragraph blockId="85.[140,1108,825,975]" pageId="85" pageNumber="86">
<emphasis bold="true" box="[140,252,825,847]" pageId="85" pageNumber="86">Figure 47.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[258,451,825,847]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[258,451,825,847]" italics="true" pageId="85" pageNumber="86">Alluviobolus antanosy</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[458,510,826,847]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
, male holotype.
<emphasis bold="true" box="[665,684,825,847]" pageId="85" pageNumber="86">A</emphasis>
head, lateral view
<emphasis bold="true" box="[858,875,825,847]" pageId="85" pageNumber="86">B</emphasis>
legs 1–7, ventral view
<emphasis bold="true" box="[1089,1108,825,847]" pageId="85" pageNumber="86">C</emphasis>
telson
<emphasis bold="true" box="[203,222,857,879]" pageId="85" pageNumber="86">D</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[537,552,857,879]" pageId="85" pageNumber="86">E</emphasis>
left posterior gonopod, anterior view
<emphasis bold="true" box="[916,930,857,879]" pageId="85" pageNumber="86">F</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[278,298,889,911]" pageId="85" pageNumber="86">G</emphasis>
left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection pageId="85" pageNumber="86" type="etymology">
<paragraph blockId="85.[140,1107,1047,1109]" pageId="85" pageNumber="86">
<emphasis bold="true" box="[185,323,1047,1073]" pageId="85" pageNumber="86">
<emphasis box="[185,318,1047,1073]" inLineHeading="true" pageId="85" pageNumber="86" reason="1">Etymology</emphasis>
:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[333,427,1047,1072]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species">
<emphasis box="[333,427,1047,1072]" italics="true" pageId="85" pageNumber="86">antanosy</emphasis>
</taxonomicName>
, noun in apposition, after the Malagasy Antanosy people, who live around Tolagnaro (Fort Dauphin) where this species was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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