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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7852FF9EFF013BD7AC85ECD9" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Ostinobolus Wesener &amp; Enghoff &amp; Sierwald 2009, gen. n." docType="treatment" docUuid="1AAC85EB-6256-450E-B461-707D235E61BA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="89" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="86" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791445" ID-GBIF-Taxon="164184219" ID-Zenodo-Dep="3791445" LSID="urn:lsid:zoobank.org:act:1AAC85EB-6256-450E-B461-707D235E61BA" httpUri="http://treatment.plazi.org/id/1772122E7852FF9EFF013BD7AC85ECD9" lastPageId="88" lastPageNumber="89" pageId="85" pageNumber="86">
<subSubSection box="[140,483,1187,1214]" pageId="85" pageNumber="86" type="nomenclature">
<paragraph blockId="85.[140,978,1187,1249]" box="[140,483,1187,1214]" pageId="85" pageNumber="86">
<heading box="[140,483,1187,1214]" level="2" pageId="85" pageNumber="86">
<emphasis bold="true" box="[140,483,1187,1214]" pageId="85" pageNumber="86">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,390,1187,1214]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[140,282,1187,1214]" inLineHeading="true" italics="true" pageId="85" pageNumber="86" reason="1">Ostinobolus</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[402,483,1187,1213]" pageId="85" pageNumber="86" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,978,1223,1249]" pageId="85" pageNumber="86" type="description">
<paragraph blockId="85.[140,978,1187,1249]" box="[140,978,1223,1249]" pageId="85" pageNumber="86">
<uri box="[140,978,1223,1249]" pageId="85" pageNumber="86">
urn:lsid:zoobank.org:act:
<uuid box="[420,978,1223,1249]" pageId="85" pageNumber="86">1A0ECD32-48CB-4631-AE39-06263DF7EB87</uuid>
</uri>
</paragraph>
</subSubSection>
<subSection lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86" type="multiple">
<subSubSection box="[140,559,1293,1320]" pageId="85" pageNumber="86" type="type_taxon">
<paragraph blockId="85.[140,559,1293,1320]" box="[140,559,1293,1320]" pageId="85" pageNumber="86">
<emphasis bold="true" box="[140,287,1293,1319]" pageId="85" pageNumber="86">
<typeStatus box="[140,198,1293,1319]" pageId="85" pageNumber="86">Type</typeStatus>
species
</emphasis>
:
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[302,491,1293,1319]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[302,491,1293,1319]" italics="true" pageId="85" pageNumber="86">Ostinobolus rufus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[498,559,1294,1320]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="85" pageNumber="86" type="reference_group">
<paragraph blockId="85.[140,419,1363,1496]" pageId="85" pageNumber="86">
<emphasis bold="true" box="[140,413,1363,1389]" pageId="85" pageNumber="86">
<emphasis box="[140,212,1363,1389]" inLineHeading="true" pageId="85" pageNumber="86" reason="1">Other</emphasis>
species included
</emphasis>
:
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,255,1399,1425]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[140,255,1399,1425]" italics="true" pageId="85" pageNumber="86">O. stellaris</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[263,324,1399,1425]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,282,1434,1460]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[140,282,1434,1460]" italics="true" pageId="85" pageNumber="86">O. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[290,350,1434,1460]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,310,1469,1496]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[140,310,1469,1496]" italics="true" pageId="85" pageNumber="86">O. subterraneus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[317,378,1470,1496]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86" type="diagnosis">
<paragraph blockId="85.[140,1108,1539,1672]" lastBlockId="87.[140,1108,166,1179]" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86">
<emphasis bold="true" box="[140,262,1539,1565]" inLineHeading="true" pageId="85" pageNumber="86" reason="1">Diagnosis</emphasis>
: the posterior gonopods of
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[587,717,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis box="[587,717,1539,1565]" italics="true" pageId="85" pageNumber="86">Ostinobolus</emphasis>
</taxonomicName>
with their apically swollen membranous area (
<emphasis box="[307,319,1575,1600]" italics="true" pageId="85" pageNumber="86">v</emphasis>
in
<figureCitation box="[362,461,1575,1601]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="85" pageNumber="86">Figs 48J</figureCitation>
,
<figureCitation box="[475,520,1574,1601]" captionStart="Figure 50" captionStartId="92.[140,212,839,861]" captionTargetBox="[140,1108,165,812]" captionTargetId="figure@92.[140,1108,421,819]" captionTargetPageId="92" captionText="Figure 50. Ostinobolus stellaris sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view; F: right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; h = head; IL = incisura lateralis; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767342" httpUri="https://zenodo.org/record/3767342/files/figure.png" pageId="85" pageNumber="86">50F</figureCitation>
,
<figureCitation box="[534,589,1574,1601]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="85" pageNumber="86">51G</figureCitation>
, 52H) are unique among
<taxonomicName authorityName="Cook" authorityYear="1895" box="[899,1037,1575,1601]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
. Telopodites of anterior gonopod diverse, either inconspicuous (
<figureCitation box="[815,916,1610,1637]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="85" pageNumber="86">Fig. 51F</figureCitation>
), or on posterior side apically with a small, non-protruding process (
<figureCitation box="[722,831,1645,1671]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="85" pageNumber="86">Fig. 48H</figureCitation>
), or with a large retrorse process (
<figureCitation box="[242,354,166,193]" captionStart="Figure 52" captionStartId="95.[140,212,843,865]" captionTargetBox="[140,1108,165,817]" captionTargetId="figure@95.[140,1108,165,493]" captionTargetPageId="95" captionText="Figure 52. Ostinobolus subterraneus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; E = reduced eye; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767344" httpUri="https://zenodo.org/record/3767344/files/figure.png" pageId="87" pageNumber="88">Fig. 52G</figureCitation>
). Telopodites of posterior gonopod as wide as coxites, arranged face-to-face with one another. First leg pair longer than second (
<figureCitation box="[903,1012,202,229]" captionStart="Figure 52" captionStartId="95.[140,212,843,865]" captionTargetBox="[140,1108,165,817]" captionTargetId="figure@95.[140,1108,165,493]" captionTargetPageId="95" captionText="Figure 52. Ostinobolus subterraneus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; E = reduced eye; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767344" httpUri="https://zenodo.org/record/3767344/files/figure.png" pageId="87" pageNumber="88">Fig. 52A</figureCitation>
). Living specimens roll themselves into a double-spiral when disturbed. Natural colour varies between species, red to brown (
<figureCitation box="[497,637,272,298]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="87" pageNumber="88">Figs 48A, C</figureCitation>
,
<figureCitation box="[649,736,272,299]" captionStart="Figure 52" captionStartId="95.[140,212,843,865]" captionTargetBox="[140,1108,165,817]" captionTargetId="figure@95.[140,1108,165,493]" captionTargetPageId="95" captionText="Figure 52. Ostinobolus subterraneus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; E = reduced eye; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767344" httpUri="https://zenodo.org/record/3767344/files/figure.png" pageId="87" pageNumber="88">52A, C</figureCitation>
). Some
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[827,957,272,298]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[827,957,272,298]" italics="true" pageId="87" pageNumber="88">Ostinobolus</emphasis>
</taxonomicName>
species share the habitus and the colour pattern with species of the sympatric living
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[963,1100,307,333]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[963,1100,307,333]" italics="true" pageId="87" pageNumber="88">Alluviobolus</emphasis>
</taxonomicName>
. The latter, however, differs from the former in the presence of completely different anterior and posterior gonopods. Gnathochilarium with a subdivided mentum and a single sclerotized ledge on each stipites (
<figureCitation box="[678,784,413,439]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="87" pageNumber="88">Figs 48E</figureCitation>
,
<figureCitation box="[798,854,413,440]" captionStart="Figure 52" captionStartId="95.[140,212,843,865]" captionTargetBox="[140,1108,165,817]" captionTargetId="figure@95.[140,1108,165,493]" captionTargetPageId="95" captionText="Figure 52. Ostinobolus subterraneus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; E = reduced eye; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767344" httpUri="https://zenodo.org/record/3767344/files/figure.png" pageId="87" pageNumber="88">52D</figureCitation>
), like in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[958,1100,413,439]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[958,1100,413,439]" italics="true" pageId="87" pageNumber="88">Flagellobolus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,287,448,474]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[140,287,448,474]" italics="true" pageId="87" pageNumber="88">Riotintobolus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[307,505,448,474]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[307,505,448,474]" italics="true" pageId="87" pageNumber="88">Pseudocentrobolus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[525,669,448,474]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[525,669,448,474]" italics="true" pageId="87" pageNumber="88">Granitobolus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[688,813,448,474]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[688,813,448,474]" italics="true" pageId="87" pageNumber="88">Caprobolus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[833,970,448,474]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[833,970,448,474]" italics="true" pageId="87" pageNumber="88">Alluviobolus</emphasis>
</taxonomicName>
. Species of this genus share fewer characters on the anterior gonopods than other genera of
<taxonomicName authorityName="Cook" authorityYear="1895" box="[140,278,519,545]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
. Especially the placement of the unusual
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[776,952,519,545]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[776,952,519,545]" italics="true" pageId="87" pageNumber="88">O. subterraneus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[963,1028,519,545]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
inside
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,270,554,580]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[140,270,554,580]" italics="true" pageId="87" pageNumber="88">Ostinobolus</emphasis>
</taxonomicName>
is tentative at best.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767338" ID-Zenodo-Dep="3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="86" pageNumber="87" startId="86.[140,212,1494,1516]" targetBox="[209,1039,164,1474]" targetPageId="86">
<paragraph blockId="86.[140,1108,1494,1676]" pageId="86" pageNumber="87">
<emphasis bold="true" box="[140,249,1494,1516]" pageId="86" pageNumber="87">Figure 48.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[256,411,1494,1515]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[256,411,1494,1515]" italics="true" pageId="86" pageNumber="87">Ostinobolus rufus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[417,469,1495,1516]" pageId="86" pageNumber="87" rank="species">sp. n.</taxonomicNameLabel>
, A, B, D, G–J male holotype; C, E, F male paratype.
<emphasis bold="true" box="[970,989,1494,1516]" pageId="86" pageNumber="87">A</emphasis>
head, lateral view
<emphasis bold="true" box="[189,206,1526,1548]" pageId="86" pageNumber="87">B</emphasis>
legs 1–7, ventral view
<emphasis bold="true" box="[424,443,1526,1548]" pageId="86" pageNumber="87">C</emphasis>
living specimen
<emphasis bold="true" box="[602,621,1526,1548]" pageId="86" pageNumber="87">D</emphasis>
telson
<emphasis bold="true" box="[691,706,1526,1548]" pageId="86" pageNumber="87">E</emphasis>
gnathochilarium
<emphasis bold="true" box="[875,889,1526,1548]" pageId="86" pageNumber="87">F</emphasis>
midbody ring with leg
<emphasis bold="true" box="[140,160,1558,1580]" pageId="86" pageNumber="87">G</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[466,486,1558,1580]" pageId="86" pageNumber="87">H</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[802,810,1558,1580]" pageId="86" pageNumber="87">I</emphasis>
left posterior gonopod, anterior view
<emphasis bold="true" box="[188,196,1590,1612]" pageId="86" pageNumber="87">Į</emphasis>
left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection pageId="87" pageNumber="88" type="distribution">
<paragraph blockId="87.[140,1108,166,1179]" pageId="87" pageNumber="88">
<emphasis bold="true" box="[185,485,589,615]" pageId="87" pageNumber="88">
<emphasis box="[185,336,589,615]" inLineHeading="true" pageId="87" pageNumber="88" reason="1">Distribution</emphasis>
and ecology
</emphasis>
: species of
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[610,737,589,615]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis box="[610,737,589,615]" italics="true" pageId="87" pageNumber="88">Ostinobolus</emphasis>
</taxonomicName>
could be recorded with four species from the rain- and subhumid forests in Southeastern
<collectingCountry box="[794,927,624,650]" name="Madagascar" pageId="87" pageNumber="88">Madagascar</collectingCountry>
(
<figureCitation box="[943,1028,624,650]" captionStart="Figure 49" captionStartId="87.[139,211,1654,1676]" captionTargetBox="[141,1112,1220,1629]" captionTargetId="figure@87.[139,1107,1228,1634]" captionTargetPageId="87" captionText="Figure 49. Distribution map of the genus Ostinobolus gen. n.." figureDoi="http://doi.org/10.5281/zenodo.3767340" httpUri="https://zenodo.org/record/3767340/files/figure.png" pageId="87" pageNumber="88">Fig. 49</figureCitation>
). Most species are restricted to the rainforest and littoral rainforest with the exception of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis italics="true" pageId="87" pageNumber="88">O. subterraneus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[281,341,695,721]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
which also occurs in the semi-dry forest of Petriky. Specimens of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis italics="true" pageId="87" pageNumber="88">O. rufus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[204,267,730,756]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
were always collected on the soil surface, with juveniles feeding
<quantity box="[1019,1108,730,756]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="87" pageNumber="88" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
deeper in the leaf litter. The same life style can be suggested for
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[843,957,765,792]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[843,957,765,792]" italics="true" pageId="87" pageNumber="88">O. stellaris</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[964,1024,766,792]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis italics="true" pageId="87" pageNumber="88">O. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[254,318,801,827]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
, which share a similar colour pattern and leg length with
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[977,1068,800,827]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[977,1068,800,827]" italics="true" pageId="87" pageNumber="88">O. rufus</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[170,341,835,862]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[170,341,835,862]" italics="true" pageId="87" pageNumber="88">O. subterraneus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[348,412,836,862]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
, however, differs from the other species of the genus in living deeply inside the root horizon of the soil.
</paragraph>
</subSubSection>
<subSubSection lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="description">
<paragraph blockId="87.[140,1108,166,1179]" pageId="87" pageNumber="88">
<emphasis bold="true" box="[185,335,905,931]" pageId="87" pageNumber="88">
<emphasis box="[185,330,905,931]" inLineHeading="true" pageId="87" pageNumber="88" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[342,406,906,932]" italics="true" pageId="87" pageNumber="88">Males</emphasis>
: length up to
<quantity box="[567,653,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="87" pageNumber="88" unit="mm" value="53.0">53 mm</quantity>
, diameter up to
<quantity box="[839,933,906,932]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="87" pageNumber="88" unit="mm" value="4.4">4.4 mm</quantity>
. 40–49 podous rings. Female length up to
<quantity box="[444,529,941,968]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="87" pageNumber="88" unit="mm" value="65.0">65 mm</quantity>
, diameter up to
<quantity box="[716,810,941,968]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="87" pageNumber="88" unit="mm" value="6.3">6.3 mm</quantity>
. 40–48 podous rings.
</paragraph>
<paragraph blockId="87.[140,1108,166,1179]" box="[185,795,976,1003]" pageId="87" pageNumber="88">
<emphasis box="[185,260,976,1002]" italics="true" pageId="87" pageNumber="88">Colour</emphasis>
highly species-specific, see species descriptions.
</paragraph>
<paragraph blockId="87.[140,1108,166,1179]" pageId="87" pageNumber="88">
<emphasis box="[185,244,1011,1037]" italics="true" pageId="87" pageNumber="88">Head</emphasis>
: each eye with circa 20–25 ocelli arranged in 4–6 vertical rows (
<figureCitation box="[996,1104,1011,1038]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="87" pageNumber="88">Figs 48A</figureCitation>
,
<figureCitation box="[140,190,1046,1073]" captionStart="Figure 50" captionStartId="92.[140,212,839,861]" captionTargetBox="[140,1108,165,812]" captionTargetId="figure@92.[140,1108,421,819]" captionTargetPageId="92" captionText="Figure 50. Ostinobolus stellaris sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view; F: right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; h = head; IL = incisura lateralis; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767342" httpUri="https://zenodo.org/record/3767342/files/figure.png" pageId="87" pageNumber="88">50A</figureCitation>
,
<figureCitation box="[201,252,1046,1073]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="87" pageNumber="88">51A</figureCitation>
).
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[271,439,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[271,439,1047,1073]" italics="true" pageId="87" pageNumber="88">O. subterraneus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[445,504,1047,1073]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
with only 6–8 decolorized and fused ocelli (
<figureCitation box="[993,1095,1046,1073]" captionStart="Figure 52" captionStartId="95.[140,212,843,865]" captionTargetBox="[140,1108,165,817]" captionTargetId="figure@95.[140,1108,165,493]" captionTargetPageId="95" captionText="Figure 52. Ostinobolus subterraneus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; E = reduced eye; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767344" httpUri="https://zenodo.org/record/3767344/files/figure.png" pageId="87" pageNumber="88">Fig. 52A</figureCitation>
). Labrum with standard three irregular teeth and a row of 10–12 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennae of medium length, reaching back to ring 3–5 (
<figureCitation box="[402,507,1152,1179]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="87" pageNumber="88">Fig. 51B</figureCitation>
). Relative lengths of antennomeres: 1&lt;&lt;2=3=4=5&lt;6
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767340" ID-Zenodo-Dep="3767340" box="[139,723,1654,1676]" httpUri="https://zenodo.org/record/3767340/files/figure.png" pageId="87" pageNumber="88" startId="87.[139,211,1654,1676]" targetBox="[141,1112,1220,1629]" targetPageId="87">
<paragraph blockId="87.[139,723,1654,1676]" box="[139,723,1654,1676]" pageId="87" pageNumber="88">
<emphasis bold="true" box="[139,250,1654,1676]" pageId="87" pageNumber="88">Figure 49.</emphasis>
Distribution map of the genus
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[544,648,1654,1675]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus" status="gen. n">
<emphasis box="[544,648,1654,1675]" italics="true" pageId="87" pageNumber="88">Ostinobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[653,712,1655,1676]" pageId="87" pageNumber="88" rank="genus">gen. n</taxonomicNameLabel>
..
</paragraph>
</caption>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
(
<figureCitation box="[148,253,167,193]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48A</figureCitation>
). Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 apically with four rows, antennomere 6 with two rows of sensilla basiconica.
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,373,272,298]" italics="true" pageId="88" pageNumber="89">Gnathochilarium</emphasis>
unusual (
<figureCitation box="[491,596,272,298]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48E</figureCitation>
). Lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Mentum basally subdivided by a conspicuous suture (
<figureCitation box="[564,669,343,369]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Figs 48E</figureCitation>
,
<figureCitation box="[683,739,342,369]" captionStart="Figure 52" captionStartId="95.[140,212,843,865]" captionTargetBox="[140,1108,165,817]" captionTargetId="figure@95.[140,1108,165,493]" captionTargetPageId="95" captionText="Figure 52. Ostinobolus subterraneus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; E = reduced eye; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767344" httpUri="https://zenodo.org/record/3767344/files/figure.png" pageId="88" pageNumber="89">52D</figureCitation>
). Stipites towards mentum with a wide sclerotized ledge (
<figureCitation box="[432,538,378,404]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48E</figureCitation>
). Hypopharyngeal crest with a field of spine-like structures. Number of sensilla on central pads not counted.
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,291,448,474]" italics="true" pageId="88" pageNumber="89">Mandible</emphasis>
: external tooth simple, rounded; mesal tooth with three cusps. Six pectinate lamellae. Molar plate with few (6 or 7) transverse furrows, anterior three furrows enlarged, posterior furrows minute.
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" box="[185,1012,554,580]" pageId="88" pageNumber="89">
<emphasis box="[185,274,554,580]" italics="true" pageId="88" pageNumber="89">Collum:</emphasis>
smooth, laterally not protruding as far as body ring 2 (
<figureCitation box="[896,999,554,580]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48A</figureCitation>
).
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,306,589,615]" italics="true" pageId="88" pageNumber="89">Body rings:</emphasis>
dorsally and laterally smooth, meso- and metazona ventrally with numerous transverse impressions. Ozopores starting at ring 6, touching suture between mesozona and metazona (
<figureCitation box="[431,536,659,686]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48C</figureCitation>
).
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,250,695,721]" italics="true" pageId="88" pageNumber="89">Telson</emphasis>
: preanal process absent. Anal valves with neither lips, nor grooves, nor micropunctation, but posteriorly elongated. Subanal scale inconspicuous (
<figureCitation box="[865,970,730,756]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Figs 48D</figureCitation>
,
<figureCitation box="[980,1032,730,757]" captionStart="Figure 50" captionStartId="92.[140,212,839,861]" captionTargetBox="[140,1108,165,812]" captionTargetId="figure@92.[140,1108,421,819]" captionTargetPageId="92" captionText="Figure 50. Ostinobolus stellaris sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view; F: right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; h = head; IL = incisura lateralis; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767342" httpUri="https://zenodo.org/record/3767342/files/figure.png" pageId="88" pageNumber="89">50C</figureCitation>
,
<figureCitation box="[1042,1094,730,757]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="88" pageNumber="89">51C</figureCitation>
).
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,230,766,791]" italics="true" pageId="88" pageNumber="89">Legs</emphasis>
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. First leg pair longer than second (
<figureCitation box="[235,341,835,862]" captionStart="Figure 52" captionStartId="95.[140,212,843,865]" captionTargetBox="[140,1108,165,817]" captionTargetId="figure@95.[140,1108,165,493]" captionTargetPageId="95" captionText="Figure 52. Ostinobolus subterraneus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; E = reduced eye; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767344" httpUri="https://zenodo.org/record/3767344/files/figure.png" pageId="88" pageNumber="89">Fig. 52A</figureCitation>
). Tarsus 2 with only small claw, but 3–6 ventral spines. Length of midbody legs (with exception of
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[511,681,871,897]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species">
<emphasis box="[511,681,871,897]" italics="true" pageId="88" pageNumber="89">O. subterraneus</emphasis>
</taxonomicName>
where leg length is reduced) circa 1.2 times body diameter in males (
<figureCitation box="[484,581,906,932]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48F</figureCitation>
) and 0.8 times body diameter in females. Each podomere with an apical, ventral seta. Coxae 3 and beyond of cylindrical shape (
<figureCitation captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48F</figureCitation>
). Tarsus with a stout dorso-apical seta and five pairs of ventral setae.
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,425,1011,1037]" italics="true" pageId="88" pageNumber="89">Male sexual characters</emphasis>
: coxae 3 and
<quantity box="[578,623,1011,1038]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="88" pageNumber="89" unit="in" value="4.0">4 in</quantity>
some species elongated into short cylindrical cone (
<figureCitation box="[246,350,1047,1073]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Figs 48B</figureCitation>
,
<figureCitation box="[362,412,1046,1073]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="88" pageNumber="89">51B</figureCitation>
).
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,378,1082,1108]" italics="true" pageId="88" pageNumber="89">Anterior gonopods</emphasis>
: median sternal projection triangular, with a wide, well-rounded tip (
<figureCitation box="[188,296,1117,1143]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48G</figureCitation>
). Sternite in most species longer than coxite. Coxite without a process, in anterior view well-visible, in posterior view only laterally present (
<figureCitation box="[922,1032,1152,1178]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Figs 48H</figureCitation>
,
<figureCitation box="[1046,1095,1152,1179]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="88" pageNumber="89">51F</figureCitation>
). Telopodite diverse inside the genus, see species-specific descriptions.
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,467,1223,1249]" italics="true" pageId="88" pageNumber="89">Posterior gonopods in situ</emphasis>
almost completely covered by anterior gonopod coxite and telopodite. Posterior gonopods coxite and telopodite completely fused, but suture partly visible (
<figureCitation box="[306,404,1293,1319]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48J</figureCitation>
). Sternite sclerotized and well-visible (
<figureCitation box="[852,950,1293,1319]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48J</figureCitation>
). Coxite protruding into a short stem towards telopodite, basally with a single spermatic groove. Telopodite as long and as wide as coxite (
<figureCitation box="[620,717,1363,1390]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Fig. 48J</figureCitation>
), the former arranged face-to-face with one another. Shape of telopodite species-specific, but apically always with a large, swollen membranous area (
<emphasis box="[456,468,1435,1460]" italics="true" pageId="88" pageNumber="89">v</emphasis>
in
<figureCitation box="[511,608,1434,1460]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Figs 48J</figureCitation>
,
<figureCitation box="[622,666,1434,1461]" captionStart="Figure 50" captionStartId="92.[140,212,839,861]" captionTargetBox="[140,1108,165,812]" captionTargetId="figure@92.[140,1108,421,819]" captionTargetPageId="92" captionText="Figure 50. Ostinobolus stellaris sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view; F: right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; h = head; IL = incisura lateralis; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767342" httpUri="https://zenodo.org/record/3767342/files/figure.png" pageId="88" pageNumber="89">50F</figureCitation>
,
<figureCitation box="[680,734,1434,1461]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="88" pageNumber="89">51G</figureCitation>
, 52H). Sperm canal discharging mesally, basal to membranous area (
<emphasis box="[545,564,1470,1495]" italics="true" pageId="88" pageNumber="89">w</emphasis>
in
<figureCitation box="[602,697,1469,1495]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="88" pageNumber="89">Figs 48J</figureCitation>
,
<figureCitation box="[709,753,1469,1496]" captionStart="Figure 50" captionStartId="92.[140,212,839,861]" captionTargetBox="[140,1108,165,812]" captionTargetId="figure@92.[140,1108,421,819]" captionTargetPageId="92" captionText="Figure 50. Ostinobolus stellaris sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view; F: right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; h = head; IL = incisura lateralis; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767342" httpUri="https://zenodo.org/record/3767342/files/figure.png" pageId="88" pageNumber="89">50F</figureCitation>
,
<figureCitation box="[764,818,1469,1496]" captionStart="Figure 51" captionStartId="93.[140,212,1558,1580]" captionTargetBox="[140,1108,976,1532]" captionTargetPageId="93" captionText="Figure 51. Ostinobolus montanus sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E right posterior gonopod, anterior view F anterior gonopod, posterior view G right posterior gonopod, posterior view. av = anal valves Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820833" httpUri="https://zenodo.org/record/3820833/files/figure.png" pageId="88" pageNumber="89">51G</figureCitation>
, 52H).
</paragraph>
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis box="[185,452,1504,1530]" italics="true" pageId="88" pageNumber="89">Female sexual characters:</emphasis>
vulva simple, bivalve-like, with a small, poorly sclerotized operculum at base. Posterior valve apically overlapping anterior one, both smooth, sensory cones absent. Towards opening, basally on each valve are two or three rows of setae.
</paragraph>
</subSubSection>
</subSection>
<subSubSection pageId="88" pageNumber="89" type="etymology">
<paragraph blockId="88.[140,1108,167,1672]" pageId="88" pageNumber="89">
<emphasis bold="true" box="[185,315,1610,1636]" inLineHeading="true" pageId="88" pageNumber="89" reason="1">Etymology</emphasis>
:
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[330,457,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis box="[330,457,1610,1636]" italics="true" pageId="88" pageNumber="89">Ostinobolus</emphasis>
</taxonomicName>
, masculine, consists of
<emphasis box="[725,797,1610,1636]" italics="true" pageId="88" pageNumber="89">Ostino</emphasis>
- referring to the superficial resemblance of the posterior gonopods to a human femur bone, and
<emphasis box="[917,982,1645,1671]" italics="true" pageId="88" pageNumber="89">-bolus</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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