300 lines
34 KiB
XML
300 lines
34 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7842FF80FF013BB2AB08EE60" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Granitobolus endemicus Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="7EFB7119-B5B1-4D30-BE97-F9F709EDD92B" docUuidSource="ZooBank" docVersion="12" lastPageNumber="71" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="70" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>1</mods:start>
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||
<mods:end>128</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791431" ID-GBIF-Taxon="163747631" ID-Zenodo-Dep="3791431" LSID="urn:lsid:zoobank.org:act:7EFB7119-B5B1-4D30-BE97-F9F709EDD92B" httpUri="http://treatment.plazi.org/id/1772122E7842FF80FF013BB2AB08EE60" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
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||
<subSubSection box="[140,614,1222,1249]" pageId="69" pageNumber="70" type="nomenclature">
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||
<paragraph blockId="69.[140,978,1222,1285]" box="[140,614,1222,1249]" pageId="69" pageNumber="70">
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||
<heading box="[140,614,1222,1249]" fontSize="11" level="2" pageId="69" pageNumber="70" reason="3">
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<emphasis bold="true" box="[140,614,1222,1249]" pageId="69" pageNumber="70">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,537,1222,1249]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,428,1222,1249]" italics="true" pageId="69" pageNumber="70">Granitobolus endemicus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[549,614,1223,1249]" pageId="69" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[140,978,1258,1285]" pageId="69" pageNumber="70" type="description">
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||
<paragraph blockId="69.[140,978,1222,1285]" box="[140,978,1258,1285]" pageId="69" pageNumber="70">
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||
<uri box="[140,978,1258,1285]" pageId="69" pageNumber="70">
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||
urn:lsid:zoobank.org:act:
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||
<uuid box="[423,978,1258,1285]" pageId="69" pageNumber="70">7EFB7119-B5B1-4D30-BE97-F9F709EDD92B</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="69" pageNumber="70" type="materials_examined">
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<paragraph blockId="69.[140,1108,1328,1672]" pageId="69" pageNumber="70">
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<emphasis bold="true" box="[140,365,1328,1354]" pageId="69" pageNumber="70">
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<emphasis box="[140,238,1328,1354]" inLineHeading="true" pageId="69" pageNumber="70" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608818250" collectingDate="2007-05-21" collectionCode="FMMC" collectorName="Wesener" country="Madagascar" county="Province" latitude="-25.083683" location="Foret Petit Lavasoa" longLatPrecision="1" longitude="46.7685" municipality="Ambatotsirongorongo Mountain" pageId="69" pageNumber="70" specimenCode="W010" specimenCount="20" specimenCount-female="6" stateProvince="Toliara" typeStatus="holotype">
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1 ♂,
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<specimenCount box="[429,472,1328,1355]" pageId="69" pageNumber="70" type="female">6 ♀</specimenCount>
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,
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<specimenCount box="[486,582,1328,1354]" pageId="69" pageNumber="70" type="generic">14 imm.</specimenCount>
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<emphasis box="[589,688,1328,1354]" italics="true" pageId="69" pageNumber="70">
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<typeStatus box="[589,683,1328,1354]" pageId="69" pageNumber="70">Holotype</typeStatus>
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:
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</emphasis>
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1 ♂ (
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<quantity box="[755,836,1328,1355]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="69" pageNumber="70" unit="mm" value="35.0">35 mm</quantity>
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long),
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<collectionCode box="[916,1004,1328,1354]" pageId="69" pageNumber="70">FMMC</collectionCode>
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<specimenCode box="[1012,1082,1328,1354]" collectionCode="FMMC" pageId="69" pageNumber="70">W010</specimenCode>
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A,
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<collectingCountry box="[140,270,1363,1389]" name="Madagascar" pageId="69" pageNumber="70">Madagascar</collectingCountry>
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,
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<collectingCounty box="[280,374,1363,1389]" pageId="69" pageNumber="70">Province</collectingCounty>
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<collectingRegion box="[378,457,1363,1389]" country="Madagascar" name="Toliara" pageId="69" pageNumber="70">Toliara</collectingRegion>
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,
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<collectingMunicipality box="[466,829,1363,1389]" pageId="69" pageNumber="70">Ambatotsirongorongo Mountain</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:1772122E7842FF80FF013BB2AB08EE60:9A04F5E37842FF83FCCA3A20AB4CEF3F" box="[839,1045,1363,1390]" country="Madagascar" county="Province" latitude="-25.083683" longLatPrecision="1" longitude="46.7685" municipality="Ambatotsirongorongo Mountain" name="Foret Petit Lavasoa" pageId="69" pageNumber="70" stateProvince="Toliara">Forêt Petit Lavasoa</location>
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, rainforest,
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<geoCoordinate box="[213,356,1398,1425]" degrees="25" direction="south" minutes="05.021" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="-25.083683">25°05.021’ S</geoCoordinate>
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,
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<geoCoordinate box="[367,514,1399,1425]" degrees="46" direction="east" minutes="46.110" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="46.7685">46°46.110’ E</geoCoordinate>
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, leg.
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<collectorName box="[567,660,1399,1425]" pageId="69" pageNumber="70">Wesener</collectorName>
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et al.,
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<date box="[730,849,1399,1425]" pageId="69" pageNumber="70" value="2007-05-21">
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<collectingDate box="[730,849,1399,1425]" pageId="69" pageNumber="70" value="2007-05-21">21.V.2007</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2608818301" collectingDate="2007-05-21" collectionCode="FMMC" collectorName="Wesener" country="Madagascar" county="Province" latitude="-25.083683" location="Foret Petit Lavasoa" longLatPrecision="1" longitude="46.7685" municipality="Ambatotsirongorongo Mountain" pageId="69" pageNumber="70" specimenCode="W010" specimenCount="9" specimenCount-female="3" stateProvince="Toliara" typeStatus="paratype">
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<emphasis box="[858,961,1399,1424]" italics="true" pageId="69" pageNumber="70">
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<typeStatus box="[858,956,1399,1424]" pageId="69" pageNumber="70">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount box="[967,1008,1399,1425]" pageId="69" pageNumber="70" type="female">3 ♀</specimenCount>
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,
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<specimenCount box="[1020,1103,1399,1425]" pageId="69" pageNumber="70" type="generic">6 imm.</specimenCount>
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,
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<collectionCode box="[140,228,1434,1460]" pageId="69" pageNumber="70">FMMC</collectionCode>
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<specimenCode box="[235,305,1434,1460]" collectionCode="FMMC" pageId="69" pageNumber="70">W010</specimenCode>
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A, same data as holotype
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818348" box="[589,1064,1434,1460]" collectingDate="2007-05-21" collectionCode="ZMH" collectorName="Wesener" country="Madagascar" county="Province" latitude="-25.083683" location="Foret Petit Lavasoa" longLatPrecision="1" longitude="46.7685" municipality="Ambatotsirongorongo Mountain" pageId="69" pageNumber="70" specimenCode="W010" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
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<specimenCount box="[589,631,1434,1460]" pageId="69" pageNumber="70" type="female">1 ♀</specimenCount>
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,
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<collectionCode box="[645,713,1434,1460]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="69" pageNumber="70" type="Museum">ZMH</collectionCode>
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||
<specimenCode box="[719,790,1434,1460]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="69" pageNumber="70" type="Museum">W010</specimenCode>
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A, same data as holotype
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph blockId="69.[140,1108,1328,1672]" pageId="69" pageNumber="70">
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<materialsCitation ID-GBIF-Occurrence="2608818305" location="University" pageId="69" pageNumber="70" specimenCount="9" specimenCount-female="2" stateProvince="Antananarivo">
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||
<emphasis bold="true" box="[185,500,1469,1495]" pageId="69" pageNumber="70">
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||
<typeStatus box="[185,495,1469,1495]" isOtherTypeStatus="true" pageId="69" pageNumber="70">
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||
<emphasis box="[185,257,1469,1495]" inLineHeading="true" pageId="69" pageNumber="70" reason="1">Other</emphasis>
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||
material examined
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</typeStatus>
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||
:
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</emphasis>
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||
<specimenCount box="[511,558,1469,1496]" pageId="69" pageNumber="70" type="female">2 ♀</specimenCount>
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,
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<specimenCount box="[576,666,1469,1496]" pageId="69" pageNumber="70" type="generic">7 imm.</specimenCount>
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||
, W010A, same data as type material, but stored as vouchers at
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||
<location LSID="urn:lsid:plazi:treatment:1772122E7842FF80FF013BB2AB08EE60:9A04F5E37842FF83FE243A94AD44EFAB" box="[425,541,1504,1530]" name="University" pageId="69" pageNumber="70" stateProvince="Antananarivo">University</location>
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||
<collectingRegion box="[549,705,1504,1530]" country="Madagascar" name="Antananarivo" pageId="69" pageNumber="70">Antananarivo</collectingRegion>
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818267" collectingDate="2007-06-14" collectionCode="FMMC" collectorName="Wesener & Schutte" latitude="-25.086176" location="Ambatotsirongorongo Mountain" longLatPrecision="1" longitude="46.74887" pageId="69" pageNumber="70" specimenCode="W099" specimenCount="1" stateProvince="Antananarivo">
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||
<specimenCount box="[717,803,1504,1530]" pageId="69" pageNumber="70" type="generic">1 imm.</specimenCount>
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,
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||
<collectionCode box="[815,905,1504,1530]" pageId="69" pageNumber="70">FMMC</collectionCode>
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||
<specimenCode box="[911,987,1504,1530]" collectionCode="FMMC" pageId="69" pageNumber="70">W099</specimenCode>
|
||
,
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||
<location LSID="urn:lsid:plazi:treatment:1772122E7842FF80FF013BB2AB08EE60:9A04F5E37842FF83FC6B3A94AEF8EC4C" latitude="-25.086176" longLatPrecision="1" longitude="46.74887" name="Ambatotsirongorongo Mountain" pageId="69" pageNumber="70" stateProvince="Antananarivo">Ambatotsirongorongo Mountain</location>
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, Forêt Grand Lavasoa,
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||
<geoCoordinate box="[683,846,1539,1566]" degrees="25" direction="south" minutes="5" orientation="latitude" pageId="69" pageNumber="70" precision="1" seconds="10.23" value="-25.086176">25°5’10.23” S</geoCoordinate>
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,
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||
<geoCoordinate box="[859,1040,1539,1566]" degrees="46" direction="east" minutes="44" orientation="longitude" pageId="69" pageNumber="70" precision="1" seconds="55.93" value="46.74887">46°44’55.93” E</geoCoordinate>
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, rainforest, leg.
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||
<collectorName box="[263,358,1575,1601]" pageId="69" pageNumber="70">Wesener</collectorName>
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||
et
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||
<collectorName box="[394,483,1575,1601]" pageId="69" pageNumber="70">Schütte</collectorName>
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||
,
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||
<date box="[494,630,1575,1601]" pageId="69" pageNumber="70" value="2007-06-14">
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||
<collectingDate box="[494,630,1575,1601]" pageId="69" pageNumber="70" value="2007-06-14">14.VI.2007</collectingDate>
|
||
</date>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="69" pageNumber="70" type="diagnosis">
|
||
<paragraph blockId="69.[140,1108,1328,1672]" pageId="69" pageNumber="70">
|
||
<emphasis bold="true" box="[185,451,1609,1635]" pageId="69" pageNumber="70">
|
||
<emphasis box="[185,324,1609,1635]" inLineHeading="true" pageId="69" pageNumber="70" reason="1">Differential</emphasis>
|
||
diagnosis:
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||
</emphasis>
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||
see key for a separation between
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[830,977,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species">
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||
<emphasis box="[830,977,1610,1636]" italics="true" pageId="69" pageNumber="70">G. endemicus</emphasis>
|
||
</taxonomicName>
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||
and
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis italics="true" pageId="69" pageNumber="70">G. andohahelensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[280,341,1646,1672]" pageId="69" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="70" pageNumber="71" type="description">
|
||
<paragraph blockId="70.[140,1108,166,1073]" pageId="70" pageNumber="71">
|
||
<emphasis bold="true" box="[185,335,166,192]" pageId="70" pageNumber="71">
|
||
<emphasis box="[185,330,166,192]" inLineHeading="true" pageId="70" pageNumber="71" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[344,496,167,192]" italics="true" pageId="70" pageNumber="71">Measurements</emphasis>
|
||
: male with 37 rings, circa
|
||
<quantity box="[805,889,166,193]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="70" pageNumber="71" unit="mm" value="35.0">35 mm</quantity>
|
||
long (broken),
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="70" pageNumber="71" unit="mm" value="3.5">3.5 mm</quantity>
|
||
wide. Females with 37 or 38 rings,
|
||
<quantity box="[591,718,202,228]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="4.8" metricValueMin="4.2" pageId="70" pageNumber="71" unit="mm" value="45.0" valueMax="48.0" valueMin="42.0">42–48 mm</quantity>
|
||
long,
|
||
<quantity box="[790,931,202,228]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.9" metricValueMin="3.8" pageId="70" pageNumber="71" unit="mm" value="4.35" valueMax="4.9" valueMin="3.8">3.8–4.9 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph blockId="70.[140,1108,166,1073]" pageId="70" pageNumber="71">
|
||
<emphasis box="[185,300,237,263]" italics="true" pageId="70" pageNumber="71">Coloration</emphasis>
|
||
affected by alcohol. Anterior part of head and appendages red (
|
||
<figureCitation captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Figs 37D, G</figureCitation>
|
||
). Mesozona and anterior part of metazona dark grey, posterior part of metazona light brown to dark orange (
|
||
<figureCitation box="[515,598,307,334]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Figs 37</figureCitation>
|
||
A–C).
|
||
<emphasis box="[675,778,308,334]" italics="true" pageId="70" pageNumber="71">Antennae</emphasis>
|
||
very short, in males protruding back to body ring 3 (
|
||
<figureCitation box="[420,526,343,369]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37D</figureCitation>
|
||
), in females to ring 2.
|
||
<emphasis box="[782,942,343,369]" italics="true" pageId="70" pageNumber="71">Preanal process</emphasis>
|
||
well-rounded, longer than anal valves (
|
||
<figureCitation box="[412,512,378,404]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="70.[140,1108,166,1073]" pageId="70" pageNumber="71">
|
||
<emphasis box="[185,368,413,439]" italics="true" pageId="70" pageNumber="71">Anterior gonopod</emphasis>
|
||
sternite elongated into a short, wide and well-rounded lobe (
|
||
<figureCitation captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37L</figureCitation>
|
||
). Mesal margin of coxite at midpoint with a projection towards sternite (
|
||
<figureCitation box="[990,1095,448,474]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37N</figureCitation>
|
||
). Telopodite on posterior side apically with a short, rounded process (
|
||
<figureCitation box="[869,973,483,510]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37N</figureCitation>
|
||
). Telopodite process not reaching mesal margin, laterally curved, slightly longer than coxite (
|
||
<figureCitation box="[992,1095,519,545]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="70.[140,1108,166,1073]" pageId="70" pageNumber="71">
|
||
<emphasis box="[185,382,554,580]" italics="true" pageId="70" pageNumber="71">Posterior gonopods</emphasis>
|
||
: telopodite apex erect, covered by a sclerite (
|
||
<figureCitation box="[890,1002,554,580]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37M</figureCitation>
|
||
). A small membranous area present between apex and beak (
|
||
<figureCitation box="[712,820,589,615]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="70" pageNumber="71">Fig. 37O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="70.[140,1108,166,1073]" pageId="70" pageNumber="71">
|
||
<emphasis bold="true" box="[185,455,624,650]" pageId="70" pageNumber="71">
|
||
<emphasis box="[185,333,624,650]" inLineHeading="true" pageId="70" pageNumber="71" reason="1">Intraspecific</emphasis>
|
||
variation:
|
||
</emphasis>
|
||
too few specimens, especially mature males, are known to evaluate the intraspecific variation. However, obviously mature females collected at the same spot range from
|
||
<quantity box="[388,536,694,721]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.8" metricValueMin="3.5" pageId="70" pageNumber="71" unit="mm" value="41.5" valueMax="48.0" valueMin="35.0">35 to 48 mm</quantity>
|
||
of length, most likely representing different stadia, or even generations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="70" pageNumber="71" type="distribution">
|
||
<paragraph blockId="70.[140,1108,166,1073]" pageId="70" pageNumber="71">
|
||
<emphasis bold="true" box="[185,496,765,791]" pageId="70" pageNumber="71">
|
||
<emphasis box="[185,336,765,791]" inLineHeading="true" pageId="70" pageNumber="71" reason="1">Distribution</emphasis>
|
||
and ecology:
|
||
</emphasis>
|
||
this species is currently known only from, and probably microendemic to, the isolated rainforests of the Ambatotsirongorongo Mountain in Southeast
|
||
<collectingCountry box="[289,422,835,861]" name="Madagascar" pageId="70" pageNumber="71">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation box="[438,522,835,862]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="70" pageNumber="71">Fig. 30</figureCitation>
|
||
), covering altogether less than
|
||
<quantity box="[875,955,833,863]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="km" value="10.0">10 km</quantity>
|
||
². The Mountain is now isolated from all other natural forests. The forest cover of the Ambatotsirongorongo Mountains decreased considerably in the last 50 years (
|
||
<bibRefCitation author="Ramanamanjato" etAl="et al." firstAuthor="Ramanamanjato" pageId="70" pageNumber="71" pagination="1791 - 1807" refId="ref57589" refString="Ramanamanjato J-B, McIntyre PB, Nussbaum RA (2002) Reptile, amphibian, and lemur diversity of the Malahelo Forest, a biogeographical transition zone in southeastern Madagascar. Biodiversity and Conservation 11: 1791 - 1807." type="journal article" year="2002">Ramanamanjato et al. 2002</bibRefCitation>
|
||
) and covers now only a few hundred hectares (
|
||
<bibRefCitation author="Moat and Smith" box="[788,1045,941,967]" firstAuthor="Moat" pageId="70" pageNumber="71" refId="ref57536" refString="Moat J, Smith P (2007) Atlas of the Vegetation of Madagascar. Kew Publishing, London, 124 pp." type="book" year="2007">Moat and Smith 2007</bibRefCitation>
|
||
). The nearest remaining forest is the worldwide unique, and also endangered, transitional littoral forest of Petriky, 5 kilometers north of Ambatotsirongorongo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[185,1105,1047,1073]" pageId="70" pageNumber="71" type="etymology">
|
||
<paragraph blockId="70.[140,1108,166,1073]" box="[185,1105,1047,1073]" pageId="70" pageNumber="71">
|
||
<emphasis bold="true" box="[185,323,1047,1073]" pageId="70" pageNumber="71">
|
||
<emphasis box="[185,318,1047,1073]" inLineHeading="true" pageId="70" pageNumber="71" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[330,441,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[330,441,1047,1073]" italics="true" pageId="70" pageNumber="71">endemicus</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the small area of distribution (
|
||
<figureCitation box="[1008,1092,1047,1073]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="70" pageNumber="71">Fig. 30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |