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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E782CFFF7FF013B62ACEEEE5F" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Riotintobolus Wesener &amp; Enghoff &amp; Sierwald 2009, gen. n." docType="treatment" docUuid="2D3A99A5-E680-4E8D-BFD4-B10BD240636D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="50" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="44" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
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<mods:detail type="volume">
<mods:number>19</mods:number>
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<mods:number>19</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791405" ID-GBIF-Taxon="164184226" ID-Zenodo-Dep="3791405" LSID="urn:lsid:zoobank.org:act:2D3A99A5-E680-4E8D-BFD4-B10BD240636D" httpUri="http://treatment.plazi.org/id/1772122E782CFFF7FF013B62ACEEEE5F" lastPageId="49" lastPageNumber="50" pageId="43" pageNumber="44">
<subSubSection box="[140,502,1046,1073]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph blockId="43.[140,991,1046,1109]" box="[140,502,1046,1073]" pageId="43" pageNumber="44">
<heading box="[140,502,1046,1073]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="3">
<emphasis bold="true" box="[140,502,1046,1073]" pageId="43" pageNumber="44">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,409,1046,1073]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[140,301,1046,1073]" italics="true" pageId="43" pageNumber="44">Riotintobolus</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[421,502,1047,1073]" pageId="43" pageNumber="44" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,991,1082,1109]" pageId="43" pageNumber="44" type="description">
<paragraph blockId="43.[140,991,1046,1109]" box="[140,991,1082,1109]" pageId="43" pageNumber="44">
<uri box="[140,991,1082,1109]" pageId="43" pageNumber="44">
urn:lsid:zoobank.org:act:
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2D3A99A5-E680-4E8D-BFD4-B10BD
<date box="[878,970,1082,1109]" pageId="43" pageNumber="44" value="36-06-24">240636</date>
D
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</uri>
</paragraph>
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<subSection lastPageId="49" lastPageNumber="50" pageId="43" pageNumber="44" type="multiple">
<subSubSection box="[140,657,1152,1179]" pageId="43" pageNumber="44" type="type_taxon">
<paragraph blockId="43.[140,657,1152,1179]" box="[140,657,1152,1179]" pageId="43" pageNumber="44">
<emphasis bold="true" box="[140,287,1152,1178]" pageId="43" pageNumber="44">
<typeStatus box="[140,198,1152,1178]" pageId="43" pageNumber="44">Type</typeStatus>
species
</emphasis>
:
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[302,588,1152,1178]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[302,588,1152,1178]" italics="true" pageId="43" pageNumber="44">Riotintobolus mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[596,657,1153,1179]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="reference_group">
<paragraph blockId="43.[140,419,1222,1355]" box="[140,419,1222,1249]" pageId="43" pageNumber="44">
<emphasis bold="true" box="[140,413,1222,1248]" pageId="43" pageNumber="44">
<emphasis box="[140,212,1222,1248]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Other</emphasis>
species included
</emphasis>
:
</paragraph>
<paragraph blockId="43.[140,419,1222,1355]" box="[140,329,1258,1284]" pageId="43" pageNumber="44">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,260,1259,1284]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[140,260,1259,1284]" italics="true" pageId="43" pageNumber="44">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[268,329,1258,1284]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="43.[140,419,1222,1355]" box="[140,307,1293,1320]" pageId="43" pageNumber="44">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,238,1293,1319]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[140,238,1293,1319]" italics="true" pageId="43" pageNumber="44">R. aridus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[246,307,1294,1320]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="43.[140,419,1222,1355]" box="[140,343,1328,1355]" pageId="43" pageNumber="44">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,274,1328,1354]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[140,274,1328,1354]" italics="true" pageId="43" pageNumber="44">R. anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[282,343,1329,1355]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44" type="diagnosis">
<paragraph blockId="43.[140,1108,1398,1672]" lastBlockId="45.[140,1108,167,1179]" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<emphasis bold="true" box="[140,255,1398,1424]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Diagnosis</emphasis>
: only Malagasy
<taxonomicName authorityName="Cook" authorityYear="1895" box="[434,560,1399,1425]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
genus where males in some species still have apodous rings in front of the telson. Body rings dorsally with a very wide stripe of flashy colour (
<figureCitation box="[226,330,1469,1495]" captionStart="Figure 24" captionStartId="44.[140,212,1590,1612]" captionTargetBox="[140,1108,447,1570]" captionTargetId="figure@44.[140,1108,446,1570]" captionTargetPageId="44" captionText="Figure 24. A Riotintobolus mandenensis sp. n., living specimens B pseudosteppe and burned shrubs dominating the area between the littoral forests of Mandena and Sainte Luce, SE Madagascar C Sainte Luce littoral forest, manioc plantation and destroyed trees inside conservation zone S8." figureDoi="http://doi.org/10.5281/zenodo.3767302" httpUri="https://zenodo.org/record/3767302/files/figure.png" pageId="43" pageNumber="44">Figs 24A</figureCitation>
,
<figureCitation box="[344,394,1469,1495]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="43" pageNumber="44">40A</figureCitation>
), a unique feature, although slim stripes occur in several genera of
<taxonomicName authorityName="Cook" authorityYear="1895" box="[170,299,1504,1530]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
. The posterior gonopods differ from other
<taxonomicName authorityName="Cook" authorityYear="1895" box="[777,903,1504,1530]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
in the presence of a unique, finger-shaped process located laterally on the telopodite (
<figureCitation box="[880,1044,1539,1566]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="43" pageNumber="44">Fig. 26M, x, y</figureCitation>
), and the presence of a large, very thin membrane located apically like a flag (
<figureCitation box="[906,1037,1575,1601]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="43" pageNumber="44">Fig. 26M, z</figureCitation>
). Anal valves with extraordinary thick lips (absent in
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[630,759,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[630,759,1610,1636]" italics="true" pageId="43" pageNumber="44">R. anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[765,823,1610,1636]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
,
<figureCitation box="[835,931,1610,1636]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="43" pageNumber="44">Figs 26B</figureCitation>
,
<figureCitation box="[942,994,1610,1636]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="43" pageNumber="44">27G</figureCitation>
), a unique character for this genus. Living specimens of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[621,896,1645,1671]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[621,896,1645,1671]" italics="true" pageId="43" pageNumber="44">Riotintobolus mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[902,959,1646,1672]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
and
<emphasis italics="true" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">R. minu- tus</emphasis>
sp. n. often remain stiff like a stick instead of rolling into a spiral when disturbed, a unique behaviour for
<taxonomicName authorityName="Cook" authorityYear="1895" box="[406,535,202,228]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
. Telson with a well-developed, sharp-edged preanal process (absent in
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[346,479,237,263]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[346,479,237,263]" italics="true" pageId="44" pageNumber="45">R. anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[488,550,238,264]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
,
<figureCitation box="[565,666,237,263]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="44" pageNumber="45">Figs 26B</figureCitation>
,
<figureCitation box="[680,732,237,263]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="44" pageNumber="45">27G</figureCitation>
,
<figureCitation box="[746,795,237,263]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="44" pageNumber="45">28A</figureCitation>
), a feature only shared with
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,326,272,298]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[140,326,272,298]" italics="true" pageId="44" pageNumber="45">Pseudocentrobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[333,407,273,299]" pageId="44" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[461,596,272,298]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[461,596,272,298]" italics="true" pageId="44" pageNumber="45">Granitobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[603,677,273,299]" pageId="44" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
Collum not greatly enlarged, similar to
<taxonomicName authorityName="de Saussure &amp; Zehntner" authorityYear="1901" box="[140,265,308,334]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis box="[140,265,308,334]" italics="true" pageId="44" pageNumber="45">Spiromimus</emphasis>
</taxonomicName>
and other small-bodied genera of
<taxonomicName authorityName="Cook" authorityYear="1895" box="[645,774,307,333]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
. Gnathochilarium with a subdivided mentum and a single sclerotized ledge on each stipites (
<figureCitation box="[827,957,343,369]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="44" pageNumber="45">Figs 26F, G</figureCitation>
,
<figureCitation box="[968,1019,343,369]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="44" pageNumber="45">27C</figureCitation>
), like in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,272,378,404]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis box="[140,272,378,404]" italics="true" pageId="44" pageNumber="45">Flagellobolus</emphasis>
</taxonomicName>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[285,471,378,404]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[285,471,378,404]" italics="true" pageId="44" pageNumber="45">Pseudocentrobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[476,549,378,404]" pageId="44" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[562,697,378,404]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[562,697,378,404]" italics="true" pageId="44" pageNumber="45">Granitobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[703,776,378,404]" pageId="44" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[789,906,378,404]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[789,906,378,404]" italics="true" pageId="44" pageNumber="45">Caprobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[912,985,378,404]" pageId="44" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
,
<emphasis italics="true" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">Alluviobo- lus</emphasis>
gen. n.,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[262,384,167,193]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[262,384,167,193]" italics="true" pageId="45" pageNumber="46">Ostinobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[391,465,167,193]" pageId="45" pageNumber="46" rank="genus">gen. n.</taxonomicNameLabel>
Vulva simple, bivalve-like, similar to all other small-bodied genera of
<taxonomicName authorityName="Cook" authorityYear="1895" box="[247,373,202,228]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
from
<collectingCountry box="[440,570,202,228]" name="Madagascar" pageId="45" pageNumber="46">Madagascar</collectingCountry>
. Posterior plate apically overlapping anterior one. Anterior gonopods of a very general shape (
<figureCitation box="[609,709,237,264]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="45" pageNumber="46">Fig. 26K</figureCitation>
), similar to those of numerous other, probably not closely related genera, like
<taxonomicName authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" box="[580,908,272,298]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis box="[580,743,272,298]" italics="true" pageId="45" pageNumber="46">Aphistogoniulus</emphasis>
<bibRefCitation author="Silvestri" box="[751,908,272,298]" firstAuthor="Silvestri" pageId="45" pageNumber="46" pagination="651 - 653" refId="ref57740" refString="Silvestri F (1897) Systema Diplopodum. Annali del Museo civico di storia naturale di Genova 38: 651 - 653." type="journal article" year="1897">Silvestri, 1897</bibRefCitation>
</taxonomicName>
(Wesener et al. in press). Telopodite of posterior gonopod with a torsion: the sperm canal runs basally along the mesal margin, but is apically rotated, so that the opening is located at the lateral margin (
<figureCitation box="[189,297,378,405]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="45" pageNumber="46">Fig. 26M</figureCitation>
). A torsion also exists in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[565,725,378,404]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis box="[565,725,378,404]" italics="true" pageId="45" pageNumber="46">Zehntnerobolus</emphasis>
</taxonomicName>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[780,908,378,404]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[780,908,378,404]" italics="true" pageId="45" pageNumber="46">Alluviobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[915,989,378,404]" pageId="45" pageNumber="46" rank="genus">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767302" ID-Zenodo-Dep="3767302" httpUri="https://zenodo.org/record/3767302/files/figure.png" pageId="44" pageNumber="45" startId="44.[140,212,1590,1612]" targetBox="[140,1108,447,1570]" targetPageId="44">
<paragraph blockId="44.[140,1108,1590,1676]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[140,282,1590,1612]" pageId="44" pageNumber="45">Figure 24. A</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[291,528,1590,1611]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[291,528,1590,1611]" italics="true" pageId="44" pageNumber="45">Riotintobolus mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[538,592,1591,1612]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
, living specimens
<emphasis bold="true" box="[769,786,1590,1612]" pageId="44" pageNumber="45">B</emphasis>
pseudosteppe and burned shrubs dominating the area between the littoral forests of Mandena and Sainte Luce, SE Madagascar
<emphasis bold="true" box="[1025,1044,1622,1644]" pageId="44" pageNumber="45">C</emphasis>
Sainte Luce littoral forest, manioc plantation and destroyed trees inside conservation zone S8.
</paragraph>
</caption>
<subSubSection pageId="45" pageNumber="46" type="distribution">
<paragraph blockId="45.[140,1108,167,1179]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[185,492,413,439]" pageId="45" pageNumber="46">
<emphasis box="[185,336,413,439]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">Distribution</emphasis>
and ecology
</emphasis>
: two species,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[656,946,413,439]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[656,946,413,439]" italics="true" pageId="45" pageNumber="46">Riotintobolus mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[956,1020,414,440]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis italics="true" pageId="45" pageNumber="46">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[238,300,449,475]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
were collected in the littoral rainforest. The two other species occur in the spiny forest.
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[333,433,483,509]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[333,433,483,509]" italics="true" pageId="45" pageNumber="46">R. aridus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[442,505,484,510]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
was recorded from the spiny forest of Angavo, circa
<quantity box="[140,215,519,545]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="46" unit="km" value="20.0">20 km</quantity>
East of Antanimora, as well as Antafoky (
<figureCitation box="[698,782,518,545]" captionStart="Figure 25" captionStartId="45.[140,212,1654,1676]" captionTargetBox="[162,1133,1244,1638]" captionTargetId="figure@45.[162,1130,1244,1634]" captionTargetPageId="45" captionText="Figure 25. Distribution of the genus Riotintobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820816" httpUri="https://zenodo.org/record/3820816/files/figure.png" pageId="45" pageNumber="46">Fig. 25</figureCitation>
), while
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[874,1010,519,545]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[874,1010,519,545]" italics="true" pageId="45" pageNumber="46">R. anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1019,1081,519,545]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
is recorded from the driest parts of
<collectingCountry box="[518,651,554,580]" name="Madagascar" pageId="45" pageNumber="46">Madagascar</collectingCountry>
(
<bibRefCitation author="Battistini" box="[666,841,554,580]" firstAuthor="Battistini" pageId="45" pageNumber="46" pagination="1 - 27" refId="ref56767" refString="Battistini R (1972) Madagascar relief and main types of landscape. In: Battistini R, Richard- Vindard G (Eds) Biogeography and Ecology in Madagascar. Monographiae Biologicae 21. The Hague, 1 - 27." type="book chapter" year="1972">Battistini 1972</bibRefCitation>
,
<bibRefCitation author="Moat &amp; Smith" box="[854,1095,554,580]" firstAuthor="Moat" pageId="45" pageNumber="46" refId="ref57536" refString="Moat J, Smith P (2007) Atlas of the Vegetation of Madagascar. Kew Publishing, London, 124 pp." type="book" year="2007">Moat &amp; Smith 2007</bibRefCitation>
), the Mahafaly plateau and the Cap Sainte Marie (
<figureCitation box="[704,791,589,616]" captionStart="Figure 25" captionStartId="45.[140,212,1654,1676]" captionTargetBox="[162,1133,1244,1638]" captionTargetId="figure@45.[162,1130,1244,1634]" captionTargetPageId="45" captionText="Figure 25. Distribution of the genus Riotintobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820816" httpUri="https://zenodo.org/record/3820816/files/figure.png" pageId="45" pageNumber="46">Fig. 25</figureCitation>
). The genus
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[938,1081,589,615]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis box="[938,1081,589,615]" italics="true" pageId="45" pageNumber="46">Riotintobolus</emphasis>
</taxonomicName>
is the first example of a relationship between spiny forest and littoral rainforest species (
<figureCitation box="[148,232,659,686]" captionStart="Figure 25" captionStartId="45.[140,212,1654,1676]" captionTargetBox="[162,1133,1244,1638]" captionTargetId="figure@45.[162,1130,1244,1634]" captionTargetPageId="45" captionText="Figure 25. Distribution of the genus Riotintobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820816" httpUri="https://zenodo.org/record/3820816/files/figure.png" pageId="45" pageNumber="46">Fig. 25</figureCitation>
).
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[252,419,659,685]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[252,419,659,685]" italics="true" pageId="45" pageNumber="46">R. mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[426,487,660,686]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[543,663,660,685]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[543,663,660,685]" italics="true" pageId="45" pageNumber="46">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[670,731,660,686]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
are microendemic,
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species">
<emphasis italics="true" pageId="45" pageNumber="46">R. mandenensis</emphasis>
</taxonomicName>
is most probably restricted to the littoral rainforest of Mandena, while
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[985,1107,695,720]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species">
<emphasis box="[985,1107,695,720]" italics="true" pageId="45" pageNumber="46">R. minutus</emphasis>
</taxonomicName>
could only be found in Sainte Luce (
<figureCitation box="[549,633,730,757]" captionStart="Figure 25" captionStartId="45.[140,212,1654,1676]" captionTargetBox="[162,1133,1244,1638]" captionTargetId="figure@45.[162,1130,1244,1634]" captionTargetPageId="45" captionText="Figure 25. Distribution of the genus Riotintobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820816" httpUri="https://zenodo.org/record/3820816/files/figure.png" pageId="45" pageNumber="46">Fig. 25</figureCitation>
). Both species from the littoral forest were collected inside the wet leaf litter, mainly on the bottom of the thin layer of leaves slightly above the root horizon. In both species the eyes are partly reduced, featuring only 1220 fused ocelli (
<figureCitation box="[416,555,835,862]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="45" pageNumber="46">Figs 26A, C</figureCitation>
,
<figureCitation box="[566,617,836,862]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="45" pageNumber="46">27E</figureCitation>
). Legs are also shorter than in the other two congeneric species (reaching only 0.7 times the diameter of body rings,
<figureCitation box="[985,1095,871,897]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="45" pageNumber="46">Fig. 27D</figureCitation>
). No explanation can be given for the unique and highly unusual defence behaviour of
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,308,941,967]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species">
<emphasis box="[140,308,941,967]" italics="true" pageId="45" pageNumber="46">R. mandenensis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[366,487,942,967]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species">
<emphasis box="[366,487,942,967]" italics="true" pageId="45" pageNumber="46">R. minutus</emphasis>
</taxonomicName>
. Both species turn stiff and motionless like a stick and rarely curl into a spiral like most other species of the
<taxonomicName authorityName="Cook" authorityYear="1895" box="[737,871,976,1002]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46" type="description">
<paragraph blockId="45.[140,1108,167,1179]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[185,335,1011,1037]" pageId="45" pageNumber="46">
<emphasis box="[185,330,1011,1037]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[342,406,1011,1037]" italics="true" pageId="45" pageNumber="46">Males</emphasis>
: length up to
<quantity box="[567,653,1011,1038]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="45" pageNumber="46" unit="mm" value="45.0">45 mm</quantity>
, diameter up to
<quantity box="[839,933,1012,1038]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="45" pageNumber="46" unit="mm" value="4.3">4.3 mm</quantity>
. 3845 podous and up to two additional apodous rings. Females: length up to
<quantity box="[845,930,1046,1073]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="45" pageNumber="46" unit="mm" value="51.0">51 mm</quantity>
, diameter up to
<quantity box="[140,234,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="45" pageNumber="46" unit="mm" value="4.6">4.6 mm</quantity>
. 3945 podous and up to two additional apodous rings.
</paragraph>
<paragraph blockId="45.[140,1108,167,1179]" pageId="45" pageNumber="46">
<emphasis box="[185,260,1117,1143]" italics="true" pageId="45" pageNumber="46">Colour</emphasis>
highly species-specific, ventrally always with a distinct wide stripe, see species for accurate descriptions.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820816" ID-Zenodo-Dep="3820816" box="[140,685,1654,1676]" httpUri="https://zenodo.org/record/3820816/files/figure.png" pageId="45" pageNumber="46" startId="45.[140,212,1654,1676]" targetBox="[162,1133,1244,1638]" targetPageId="45">
<paragraph box="[140,685,1654,1676]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[140,251,1654,1676]" pageId="45" pageNumber="46">Figure 25.</emphasis>
Distribution of the genus
<emphasis box="[498,615,1654,1675]" italics="true" pageId="45" pageNumber="46">Riotintobolus</emphasis>
gen. n.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767304" ID-Zenodo-Dep="3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="46" pageNumber="47" startId="46.[140,212,1398,1420]" targetBox="[184,1064,165,1374]" targetPageId="46">
<paragraph blockId="46.[140,1108,1398,1676]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[140,252,1398,1420]" pageId="46" pageNumber="47">Figure 26.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[259,495,1398,1419]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[259,495,1398,1419]" italics="true" pageId="46" pageNumber="47">Riotintobolus mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[502,555,1399,1420]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
, A, B, IM male holotype, CH female paratype.
<emphasis bold="true" box="[1032,1051,1398,1420]" pageId="46" pageNumber="47">A</emphasis>
head, lateral view
<emphasis bold="true" box="[252,269,1430,1452]" pageId="46" pageNumber="47">B</emphasis>
telson
<emphasis bold="true" box="[338,357,1430,1452]" pageId="46" pageNumber="47">C</emphasis>
head, detail of incisura lateralis
<emphasis bold="true" box="[661,680,1430,1452]" pageId="46" pageNumber="47">D</emphasis>
head, frontal view
<emphasis bold="true" box="[862,877,1430,1452]" pageId="46" pageNumber="47">E</emphasis>
second leg with vulva
<emphasis bold="true" box="[1093,1107,1430,1452]" pageId="46" pageNumber="47">F</emphasis>
gnathochilarium, overview
<emphasis bold="true" box="[393,413,1462,1484]" pageId="46" pageNumber="47">G</emphasis>
gnathochilarium, detail of mentum
<emphasis bold="true" box="[752,772,1462,1484]" pageId="46" pageNumber="47">H</emphasis>
left mandible, mesal view
<emphasis bold="true" box="[1020,1028,1462,1484]" pageId="46" pageNumber="47">I</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[363,371,1494,1516]" pageId="46" pageNumber="47">Į</emphasis>
left posterior gonopod, anterior view
<emphasis bold="true" box="[726,743,1494,1516]" pageId="46" pageNumber="47">K</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[1062,1077,1494,1516]" pageId="46" pageNumber="47">L</emphasis>
telopodite of left posterior gonopod, posterior view
<emphasis bold="true" box="[599,620,1526,1548]" pageId="46" pageNumber="47">M</emphasis>
left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767306" ID-Zenodo-Dep="3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="47" pageNumber="48" startId="47.[140,212,1494,1516]" targetBox="[181,1066,165,1466]" targetPageId="47">
<paragraph blockId="47.[140,1108,1494,1676]" pageId="47" pageNumber="48">
<emphasis bold="true" box="[140,250,1494,1516]" pageId="47" pageNumber="48">Figure 27.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[255,450,1494,1516]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[255,450,1494,1516]" italics="true" pageId="47" pageNumber="48">Riotintobolus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[455,506,1495,1516]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
,
<emphasis bold="true" box="[515,561,1494,1516]" pageId="47" pageNumber="48">AE</emphasis>
female paratype(s),
<emphasis bold="true" box="[744,788,1494,1516]" pageId="47" pageNumber="48">FK</emphasis>
male holotype.
<emphasis bold="true" box="[934,953,1494,1516]" pageId="47" pageNumber="48">A</emphasis>
two living specimens
<emphasis bold="true" box="[196,213,1526,1548]" pageId="47" pageNumber="48">B</emphasis>
habitus
<emphasis bold="true" box="[295,314,1526,1548]" pageId="47" pageNumber="48">C</emphasis>
gnathochilarium, overview;
<emphasis bold="true" box="[583,602,1526,1548]" pageId="47" pageNumber="48">D</emphasis>
midbody ring with leg pair
<emphasis bold="true" box="[871,886,1526,1548]" pageId="47" pageNumber="48">E</emphasis>
head, detail of incisura lateralis
<emphasis bold="true" box="[217,231,1558,1580]" pageId="47" pageNumber="48">F</emphasis>
legs 15, ventral view
<emphasis bold="true" box="[451,471,1558,1580]" pageId="47" pageNumber="48">G</emphasis>
telson
<emphasis bold="true" box="[541,561,1558,1580]" pageId="47" pageNumber="48">H</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[876,884,1558,1580]" pageId="47" pageNumber="48">I</emphasis>
left posterior gonopod, anterior view
<emphasis bold="true" box="[265,273,1590,1612]" pageId="47" pageNumber="48">Į</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[585,602,1590,1612]" pageId="47" pageNumber="48">K</emphasis>
left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,244,167,193]" italics="true" pageId="48" pageNumber="49">Head</emphasis>
: each eye either with circa 2834 (
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[643,742,167,193]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species">
<emphasis box="[643,742,167,193]" italics="true" pageId="48" pageNumber="49">R. aridus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[801,936,167,193]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species">
<emphasis box="[801,936,167,193]" italics="true" pageId="48" pageNumber="49">R. anomalus</emphasis>
</taxonomicName>
), or 1220 (
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species">
<emphasis italics="true" pageId="48" pageNumber="49">R. mandenensis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[340,463,203,228]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species">
<emphasis box="[340,463,203,228]" italics="true" pageId="48" pageNumber="49">R. minutus</emphasis>
</taxonomicName>
) partly fused ocelli arranged in 35 vertical rows (
<figureCitation captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Figs 26C</figureCitation>
,
<figureCitation box="[204,255,237,263]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="48" pageNumber="49">27E</figureCitation>
). Labrum with standard three irregular teeth and a row of 1012 stout marginal setae. Clypeus with two (rarely three) setiferous foveolae on each side (
<figureCitation box="[988,1094,272,299]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26D</figureCitation>
). Antennae short or of medium length, reaching back to ring 2 or 5 (
<figureCitation box="[924,1030,307,333]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Figs 26A</figureCitation>
,
<figureCitation box="[1044,1095,308,334]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="48" pageNumber="49">27B</figureCitation>
). Relative lengths of antennomeres: 1&lt;&lt;2&gt;3=4=5&lt;6 (
<figureCitation box="[740,849,343,369]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Figs 26A</figureCitation>
,
<figureCitation box="[865,915,343,369]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="48" pageNumber="49">28B</figureCitation>
,
<figureCitation box="[931,983,343,369]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="48" pageNumber="49">29B</figureCitation>
). Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 latero-apically with a field of four rows, antennomere 6 with a field of two rows of sensilla basiconica.
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,367,483,509]" italics="true" pageId="48" pageNumber="49">Gnathochilarium</emphasis>
unusual (
<figureCitation box="[474,571,483,510]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26F</figureCitation>
). Lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae (
<figureCitation box="[886,986,519,546]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26F</figureCitation>
). Mentum basally subdivided by a wide suture (
<figureCitation box="[536,640,554,580]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Figs 26G</figureCitation>
,
<figureCitation box="[652,704,554,580]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="48" pageNumber="49">27C</figureCitation>
). Stipites each towards mentum with a large sclerotized ledge (
<figureCitation box="[407,507,589,616]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26G</figureCitation>
) Palpi of gnathochilarium with numerous sensilla. Hypopharyngeal crest with a field of spine-like structures. Central pads of endochilarium separated by a step into two levels, each carrying sensory cones, number of cones not counted.
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,291,695,721]" italics="true" pageId="48" pageNumber="49">Mandible</emphasis>
: external tooth simple, rounded; mesal tooth with three shallow cusps (
<figureCitation box="[148,257,730,757]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26H</figureCitation>
). Four or five pectinate lamellae. Molar plate with few (five) transverse furrows, anterior two furrows enlarged, posterior furrows minute.
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" box="[185,948,800,827]" pageId="48" pageNumber="49">
<emphasis box="[185,274,800,826]" italics="true" pageId="48" pageNumber="49">Collum:</emphasis>
smooth, laterally not protruding as far as ring 2 (
<figureCitation box="[832,935,800,827]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26A</figureCitation>
).
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,306,835,861]" italics="true" pageId="48" pageNumber="49">Body rings:</emphasis>
dorsally and laterally smooth, meso- and metazona ventrally with numerous transverse impressions. Ozopores starting at ring 6, touching suture between mesozona and metazona (
<figureCitation box="[431,534,906,932]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="48" pageNumber="49">Fig. 27B</figureCitation>
).
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,250,941,967]" italics="true" pageId="48" pageNumber="49">Telson</emphasis>
: preanal process protruding, sharp-edged (except for
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[836,968,941,967]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[836,968,941,967]" italics="true" pageId="48" pageNumber="49">R. anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[974,1035,942,968]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
). Anal valves with well-visible lips and micropunctation (
<figureCitation box="[695,796,976,1003]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26B</figureCitation>
). Deep groove present anteriorly to sharp-edged lips in
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[449,613,1011,1037]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[449,613,1011,1037]" italics="true" pageId="48" pageNumber="49">R. mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[619,678,1012,1038]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[732,850,1012,1037]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[732,850,1012,1037]" italics="true" pageId="48" pageNumber="49">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[856,915,1012,1038]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[928,1030,1011,1037]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Figs 26B</figureCitation>
,
<figureCitation box="[1040,1095,1012,1038]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="48" pageNumber="49">27G</figureCitation>
).
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,230,1047,1072]" italics="true" pageId="48" pageNumber="49">Legs</emphasis>
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 410 ventral/mesal setae. Length of midbody legs species-specific (
<figureCitation box="[287,395,1117,1143]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="48" pageNumber="49">Figs 27D</figureCitation>
,
<figureCitation box="[408,460,1117,1143]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="48" pageNumber="49">28C</figureCitation>
,
<figureCitation box="[472,525,1117,1143]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="48" pageNumber="49">29C</figureCitation>
). Each podomere with an apical ventral seta. Coxae 3 and beyond of cylindrical shape (
<figureCitation box="[546,654,1152,1178]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="48" pageNumber="49">Figs 28C</figureCitation>
,
<figureCitation box="[668,722,1152,1178]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="48" pageNumber="49">29C</figureCitation>
). Tarsus with a stout dorso-apical seta and three pairs of ventral setae. Tibia short, half as long as tarsus (
<figureCitation box="[923,1029,1187,1214]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="48" pageNumber="49">Figs 28C</figureCitation>
,
<figureCitation box="[1041,1094,1188,1214]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="48" pageNumber="49">29C</figureCitation>
).
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,427,1223,1249]" italics="true" pageId="48" pageNumber="49">Male sexual characters</emphasis>
: male legs in some species with tarsal pads. Male coxae 37 unmodified (
<figureCitation box="[287,388,1258,1284]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="48" pageNumber="49">Fig. 27F</figureCitation>
).
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" pageId="48" pageNumber="49">
<emphasis box="[185,384,1293,1319]" italics="true" pageId="48" pageNumber="49">Anterior gonopods</emphasis>
: median sternal projection triangular, broadly rounded (
<figureCitation captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26I</figureCitation>
). Sternite longer than coxite, but slightly shorter than mesal process of coxite (
<figureCitation captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26I</figureCitation>
). Process of coxite long, but shorter than telopodite (
<figureCitation box="[790,885,1363,1390]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26I</figureCitation>
). Telopodite always with a large, triangular retrorse process. Retrorse process laterally and apically extending beyond telopodite and coxite (
<figureCitation box="[527,633,1434,1461]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26K</figureCitation>
).
</paragraph>
<paragraph blockId="48.[140,1108,167,1672]" lastBlockId="49.[140,1108,167,1038]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis box="[185,391,1469,1495]" italics="true" pageId="48" pageNumber="49">Posterior gonopods</emphasis>
coxite and telopodite clearly separated (
<figureCitation box="[878,1028,1469,1496]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Figs 26J, M</figureCitation>
). Sternite sclerotized and well-visible (
<figureCitation box="[531,649,1504,1531]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26M</figureCitation>
). Coxite mesally with a single groove, coxite protruding into a short stem towards telopodite. Telopodite slightly shorter than coxite (
<figureCitation box="[292,410,1575,1602]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26M</figureCitation>
). Sperm canal discharging apically and laterally instead of mesally because of a torsion (
<figureCitation box="[473,614,1610,1636]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Figs 26J, M</figureCitation>
). Telopodites laterally always with a fingershaped process (
<figureCitation box="[331,508,1645,1672]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="48" pageNumber="49">Fig. 26M, x, y</figureCitation>
), apically with a large, thin membrane (
<figureCitation captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">Fig. 26M, z</figureCitation>
). Width of membrane apically larger than basally. Sperm canal first running along inner margin of coxite and sternite, than apically running through the telopodite before discharging laterally on finger-shaped process and often also apically into thin membrane (
<figureCitation box="[279,393,272,299]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="49" pageNumber="50">Fig. 26M</figureCitation>
).
</paragraph>
<paragraph blockId="49.[140,1108,167,1038]" pageId="49" pageNumber="50">
<emphasis box="[185,457,307,333]" italics="true" pageId="49" pageNumber="50">Female sexual characters:</emphasis>
vulva simple, with a small, poorly sclerotized operculum at base, bivalve-like. Posterior valve apically overlapping anterior valve. Both valves smooth, lacking sensory cones. Towards opening basally on each valve with two or three rows of setae.
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="50" type="etymology">
<paragraph blockId="49.[140,1108,167,1038]" pageId="49" pageNumber="50">
<emphasis bold="true" box="[185,315,448,474]" inLineHeading="true" pageId="49" pageNumber="50" reason="1">Etymology</emphasis>
:
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[332,475,448,474]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis box="[332,475,448,474]" italics="true" pageId="49" pageNumber="50">Riotintobolus</emphasis>
</taxonomicName>
, masculine, refers to the mining company Rio Tintoº. Rio Tintos subsidiary, Qit Madgascar Minearlsº, now owns the littoral rainforests of Mandena and Sainte Luce (
<bibRefCitation author="Vincelette" box="[447,694,519,545]" etAl="et al." firstAuthor="Vincelette" pageId="49" pageNumber="50" pagination="1535 - 1537" refId="ref57793" refString="Vincelette M, Randrihasipara L, Ramanamanjato J-B, Lowry (II) PP, Ganzhorn JU (2003) Mining and environmental conservation: The case of QIT Madagascar Minerals in the Southeast. In: Goodman SM, Benstead JP (Eds) The Natural History of Madagascar. University of Chicago Press, Chicago, pp. 1535 - 1537." type="book chapter" year="2003">Vincelette et al. 2003</bibRefCitation>
). This encompasses all the remaining distribution area of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[368,656,554,580]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[368,656,554,580]" italics="true" pageId="49" pageNumber="50">Riotintobolus mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[665,730,554,580]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
, which only occurs in Mandena, and of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[218,338,590,615]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[218,338,590,615]" italics="true" pageId="49" pageNumber="50">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[345,407,590,616]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
, a species only recorded from Sainte Luce. Conservation zones will be the only remaining natural habitat after a large-scale mining action is undertaken by Riotinto in the coming years (Ganzhorn et al. 2003,
<bibRefCitation author="Bollen and Donati" firstAuthor="Bollen" pageId="49" pageNumber="50" pagination="57 - 66" refId="ref56808" refString="Bollen A, Donati G (2006) Conservation status of the littoral forest of south-eastern Madagascar: a review. Oryx 40 (1): 57 - 66." type="journal article" year="2006">Bollen and Donati 2006</bibRefCitation>
). These conservation zones will be directly managed by Qit
<collectingCountry box="[869,1002,695,721]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
Minerals in order to conserve the biodiversity of the forests. The proposed conservation zones are 160 ha (but divided by a swamp) for the littoral forest of Mandena, and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis italics="true" pageId="49" pageNumber="50">Riotintobolus mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[321,382,801,827]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
The conservation zone for
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[690,811,801,826]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[690,811,801,826]" italics="true" pageId="49" pageNumber="50">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[819,882,801,827]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
, the S9 fragment of Sainte Luce is circa 200 ha large. The area between both forests is already degraded to pseudosteppe (
<figureCitation box="[306,408,871,897]" captionStart="Figure 24" captionStartId="44.[140,212,1590,1612]" captionTargetBox="[140,1108,447,1570]" captionTargetId="figure@44.[140,1108,446,1570]" captionTargetPageId="44" captionText="Figure 24. A Riotintobolus mandenensis sp. n., living specimens B pseudosteppe and burned shrubs dominating the area between the littoral forests of Mandena and Sainte Luce, SE Madagascar C Sainte Luce littoral forest, manioc plantation and destroyed trees inside conservation zone S8." figureDoi="http://doi.org/10.5281/zenodo.3767302" httpUri="https://zenodo.org/record/3767302/files/figure.png" pageId="49" pageNumber="50">Fig. 24B</figureCitation>
). The conservation zones, at least of the Sainte Luce fragment, are still under human pressure (
<figureCitation box="[506,613,906,932]" captionStart="Figure 24" captionStartId="44.[140,212,1590,1612]" captionTargetBox="[140,1108,447,1570]" captionTargetId="figure@44.[140,1108,446,1570]" captionTargetPageId="44" captionText="Figure 24. A Riotintobolus mandenensis sp. n., living specimens B pseudosteppe and burned shrubs dominating the area between the littoral forests of Mandena and Sainte Luce, SE Madagascar C Sainte Luce littoral forest, manioc plantation and destroyed trees inside conservation zone S8." figureDoi="http://doi.org/10.5281/zenodo.3767302" httpUri="https://zenodo.org/record/3767302/files/figure.png" pageId="49" pageNumber="50">Fig. 24C</figureCitation>
). Since Rio Tintoº owns and is responsible for the only remaining habitat where two
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[611,754,941,967]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[611,754,941,967]" italics="true" pageId="49" pageNumber="50">Riotintobolus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[761,842,941,967]" pageId="49" pageNumber="50">
<taxonomicNameLabel box="[761,842,941,967]" pageId="49" pageNumber="50" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
species exist, the whole genus is named after the company. The author hopes that this move will increase the probability of the survival of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[468,636,1011,1037]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[468,636,1011,1037]" italics="true" pageId="49" pageNumber="50">R. mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[643,704,1012,1038]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[762,882,1012,1037]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[762,882,1012,1037]" italics="true" pageId="49" pageNumber="50">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[890,951,1012,1038]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>