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<document id="39FF8E2CB908CD5F86B4D3C1E3C61CC3" ID-DOI="10.5281/zenodo.10981864" ID-ISSN="2295-0214" ID-Zenodo-Dep="10981864" ID-ZooBank="9E295BCA-47CF-4991-930C-60E295516E51" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1713291721940" checkinUser="felipe" docAuthor="Balkenohl, Michael" docDate="2023" docId="175ACD40FFC14A6DFDB8FDA142BBFCA5" docLanguage="en" docName="BelgJEntomol.138.1-17.pdf" docOrigin="Belgian Journal of Entomology 138" docStyle="DocumentStyle:32E55E25D8523BC97666C7812278ECC1.2:BelgJEntomol.2022-.journal_article.type1" docStyleId="32E55E25D8523BC97666C7812278ECC1" docStyleName="BelgJEntomol.2022-.journal_article.type1" docStyleVersion="2" docTitle="Eoclivina machadoi" docType="treatment" docVersion="3" lastPageNumber="13" masterDocId="EB63B538FFCA4A61FFB5FFAE4353FF97" masterDocTitle="The genus Eoclivina (KULT, 1959) from Africa with the description of two new species (Coleoptera, Carabidae, Clivinini)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="12" updateTime="1713864061911" updateUser="tatiana">
<mods:mods id="6382356F609895C9D3D8132AFD1DF9BF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1138FB45A8799C4D0EA7194EF6ADDB5A">
<mods:title id="B4C0A16AB8A44A8F883B315416EF0EFE">The genus Eoclivina (KULT, 1959) from Africa with the description of two new species (Coleoptera, Carabidae, Clivinini)</mods:title>
</mods:titleInfo>
<mods:name id="C220F3BDCC372456FAA79D42C07BED30" type="personal">
<mods:role id="C840703B2B9CF96D6194BFDBDC8E04EE">
<mods:roleTerm id="D5C0A8C45F2AF42597F5362F9551D72F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D2414D54CE88652F9C480C84BB936ED">Balkenohl, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="8555D6C5D86EE5AE67113D7E9D13C565">text</mods:typeOfResource>
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<mods:title id="1ED731C016A318B8FD59AB83A9E5110A">Belgian Journal of Entomology</mods:title>
</mods:titleInfo>
<mods:part id="666D96230B4BFF0ACF39ADA4CE537EB1">
<mods:date id="1A13AC9FB18FED48D837C76BE998EB1A">2023</mods:date>
<mods:detail id="6AE979785B9E1DF2B837A0AED4D0F2E8" type="pubDate">
<mods:number id="553EE3FAA719B16E279439628D5E5AF4">2023-06-16</mods:number>
</mods:detail>
<mods:detail id="4B1FDAA035CC41A09FFB7681385A5827" type="volume">
<mods:number id="86BB2632BE43549AA725E9CD658B62BE">138</mods:number>
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<mods:start id="955623CD90BE7532F6EFEA285E8E3208">1</mods:start>
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<mods:classification id="2ABA04864464B701441D3763322892D3">journal article</mods:classification>
<mods:identifier id="EF63ED92829D152399A0ED2B5E2657BD" type="DOI">10.5281/zenodo.10981864</mods:identifier>
<mods:identifier id="11D12478CD2CD1DD180111972575DE08" type="ISSN">2295-0214</mods:identifier>
<mods:identifier id="2DE924FB0ED69BEA70A11AA612EFF73C" type="Zenodo-Dep">10981864</mods:identifier>
<mods:identifier id="21FEA653E0A9CD791E2EE60A8CAF3C60" type="ZooBank">9E295BCA-47CF-4991-930C-60E295516E51</mods:identifier>
</mods:mods>
<treatment id="175ACD40FFC14A6DFDB8FDA142BBFCA5" LSID="urn:lsid:plazi:treatment:175ACD40FFC14A6DFDB8FDA142BBFCA5" httpUri="http://treatment.plazi.org/id/175ACD40FFC14A6DFDB8FDA142BBFCA5" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="D7E92FDDFFC14A6AFDB8FDA14775FDBA" box="[525,1062,527,557]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="9F4C7C56FFC14A6AFDB8FDA14775FDBA" blockId="11.[525,1062,527,557]" box="[525,1062,527,557]" pageId="11" pageNumber="12">
<heading id="C404CB3AFFC14A6AFDB8FDA14775FDBA" bold="true" box="[525,1062,527,557]" centered="true" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="58F307D5FFC14A6AFDB8FDA14775FDBA" ID-CoL="5ZCCZ" authority="(BASILEWSKY, 1955)" baseAuthorityName="Basilewsky" baseAuthorityYear="1955" box="[525,1062,527,557]" class="Insecta" family="Carabidae" genus="Eoclivina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="machadoi">
<emphasis id="AD87A044FFC14A6AFDB8FDA14775FDBA" bold="true" box="[525,1062,527,557]" pageId="11" pageNumber="12">
<emphasis id="AD87A044FFC14A6AFDB8FDA14045FDBB" bold="true" box="[525,790,527,556]" italics="true" pageId="11" pageNumber="12">Eoclivina machadoi</emphasis>
(
<bibRefCitation id="FB6201A7FFC14A6AFC9DFDBE474EFDBA" author="BASILEWSKY P." box="[808,1053,528,557]" pageId="11" pageNumber="12" pagination="93 - 137" refId="ref8356" refString="BASILEWSKY P., 1955. - Carabidae (Coleoptera) de l'Angola. Publicacoes culturais Companhia de Diamantes de Angola, Lisboa, 27: 93 - 137." type="journal article" year="1955">BASILEWSKY, 1955</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D7E92FDDFFC14A6AFD0AFDEA4027FDF6" box="[703,884,580,609]" pageId="11" pageNumber="12" type="description">
<paragraph id="9F4C7C56FFC14A6AFD0AFDEA4027FDF6" blockId="11.[703,884,580,609]" box="[703,884,580,609]" pageId="11" pageNumber="12">
<heading id="C404CB3AFFC14A6AFD0AFDEA4027FDF6" box="[703,884,580,609]" centered="true" fontSize="12" level="2" pageId="11" pageNumber="12" reason="7">
(
<figureCitation id="07C860D3FFC14A6AFD7DFDEA4048FDF6" box="[712,795,580,609]" captionStart="Figs 1-5" captionStartId="5.[189,235,1963,1987]" captionTargetBox="[260,1326,243,1919]" captionTargetId="figure-9@5.[210,1378,212,1963]" captionTargetPageId="5" captionText="Figs 1-5. Eoclivina species, habitus, dorsal view. 1, E. dumolinii (Putzeys, 1846), ♂, specimen from Velingara, Guinea. 2, E. basilewskyi sp. nov., holotype. 3, E. machadoi (Basilewsky, 1955), holotype. 4, E. burgeoni (Kult, 1959), holotype. 5, E. schmidli sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10981866" httpUri="https://zenodo.org/record/10981866/files/figure.png" pageId="11" pageNumber="12">Figs 3</figureCitation>
,
<figureCitation id="07C860D3FFC14A6AFC9CFDEA4068FDF6" box="[809,827,580,609]" captionStart="Figs 6-9" captionStartId="6.[189,235,1935,1959]" captionTargetBox="[213,1382,675,1851]" captionTargetId="figure-133@6.[189,1399,628,1919]" captionTargetPageId="6" captionText="Figs 6-9. Eoclivina species, male genitalia. A-B, aedeagus. A, dorsal view. B, lateral view. C-D, parameres, dorsal view. 6, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 7, E. basilewskyi sp. nov., holotype. 8, E. machadoi (Basilewsky, 1955), paratype from Riv. Chiumbe, Angola. 9, E. schmidli sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10981868" httpUri="https://zenodo.org/record/10981868/files/figure.png" pageId="11" pageNumber="12">8</figureCitation>
,
<figureCitation id="07C860D3FFC14A6AFCFCFDEA4039FDF6" box="[841,874,580,609]" captionStart="Figs 10-14" captionStartId="7.[189,235,1966,1990]" captionTargetBox="[228,1363,216,1876]" captionTargetId="figure-9@7.[204,1374,212,1967]" captionTargetPageId="7" captionText="Figs 10-14. Eoclivina species, females, gonocoxite and epipleurite, ventro-lateral view. 10, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 11, E. basilewskyi sp. nov., paratype. 12, E. machadoi (Basilewsky, 1955), paratype. 13, E. burgeoni (Kult, 1959), holotype. 14, E. schmidli sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.10981870" httpUri="https://zenodo.org/record/10981870/files/figure.png" pageId="11" pageNumber="12">12</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D7E92FDDFFC14A6AFF08FDD64759FCB5" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="9F4C7C56FFC14A6AFF08FDD6418DFD02" blockId="11.[189,734,632,661]" box="[189,734,632,661]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="BFE35B78FFC14A6AFF08FDD6418DFD02" box="[189,734,632,661]" pageId="11" pageNumber="12">
<treatmentCitation id="1E525A47FFC14A6AFF08FDD6418BFD02" author="BASILEWSKY P." box="[189,728,632,661]" page="101" pageId="11" pageNumber="12" year="1955">
<taxonomicName id="58F307D5FFC14A6AFF08FDD6418BFD02" ID-CoL="5ZCCZ" authority="BASILEWSKY, 1955: 101" authorityName="BASILEWSKY" authorityPageNumber="101" authorityYear="1955" box="[189,728,632,661]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="machadoi">
<emphasis id="AD87A044FFC14A6AFF08FDD642F2FD02" box="[189,417,632,661]" italics="true" pageId="11" pageNumber="12">Clivina machadoi</emphasis>
<bibRefCitation id="FB6201A7FFC14A6AFE1CFDD6418BFD02" author="BASILEWSKY P." box="[425,728,632,661]" pageId="11" pageNumber="12" pagination="93 - 137" refId="ref8356" refString="BASILEWSKY P., 1955. - Carabidae (Coleoptera) de l'Angola. Publicacoes culturais Companhia de Diamantes de Angola, Lisboa, 27: 93 - 137." type="journal article" year="1955">BASILEWSKY, 1955: 101</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="9F4C7C56FFC14A6AFF08FD02427BFD7A" blockId="11.[189,1398,684,749]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="BFE35B78FFC14A6AFF08FD02427BFD7A" pageId="11" pageNumber="12">
<taxonomicName id="58F307D5FFC14A6AFF08FD024013FD5E" ID-CoL="5ZCCZ" authority="(BASILEWSKY, 1955)" baseAuthorityName="BASILEWSKY" baseAuthorityYear="1955" box="[189,832,684,713]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="machadoi" subGenus="Eoclivina">
<emphasis id="AD87A044FFC14A6AFF08FD024164FD5E" box="[189,567,684,713]" italics="true" pageId="11" pageNumber="12">Clivina (Eoclivina) machadoi</emphasis>
(
<bibRefCitation id="FB6201A7FFC14A6AFDFCFD024064FD5E" author="BASILEWSKY P." box="[585,823,684,713]" pageId="11" pageNumber="12" pagination="93 - 137" refId="ref8356" refString="BASILEWSKY P., 1955. - Carabidae (Coleoptera) de l'Angola. Publicacoes culturais Companhia de Diamantes de Angola, Lisboa, 27: 93 - 137." type="journal article" year="1955">BASILEWSKY, 1955</bibRefCitation>
)
</taxonomicName>
,
<treatmentCitation id="1E525A47FFC14A6AFCF9FD024770FD5E" author="KULT K." box="[844,1059,684,713]" page="215" pageId="11" pageNumber="12" year="1959">
<bibRefCitation id="FB6201A7FFC14A6AFCF9FD024770FD5E" author="KULT K." box="[844,1059,684,713]" pageId="11" pageNumber="12" pagination="172 - 225" refId="ref8778" refString="KULT K., 1959. - Revision of the African Species of the old Genus Clivina LATR. (Col. Carabidae). Revue de Zoologie et de Botanique Africaines, 60: 172 - 225." type="journal article" year="1959">KULT, 1959: 215</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1E525A47FFC14A6AFB9AFD024672FD5E" author="LORENZ W." box="[1071,1313,684,713]" page="145" pageId="11" pageNumber="12" year="2005">
<bibRefCitation id="FB6201A7FFC14A6AFB9AFD024672FD5E" author="LORENZ W." box="[1071,1313,684,713]" pageId="11" pageNumber="12" refId="ref8817" refString="LORENZ W., 2005. - Systematic list of extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition 2005. Tutzing, printed by the author. 530 pp." type="book" year="2005">LORENZ 2005: 145</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1E525A47FFC14A6AFA9BFD024398FD7A" author="LORENZ W." page="5" pageId="11" pageNumber="12" year="2023">
<bibRefCitation id="FB6201A7FFC14A6AFA9BFD024398FD7A" author="LORENZ W." pageId="11" pageNumber="12" pagination="2023" refId="ref8866" refString="LORENZ W., 2023. - CarabCat: Global database of ground beetles (version February 2023). In: Species 2000 &amp; IT IS catalogue of life, 2021 Annual Checklist (Roskov Y., Ower G., Orrell T., Nicolson D., Bailly N., Kirk P. M., Bourgoin T., DeWalt R. E., Decock W., Nieukerken E. van, Zarucchi J., Penev L., eds). Species 2000. Naturalis, Leiden, The Netherlands. www. catalogueoflife. org / annual-checklist / 2023. (date accessed March 2023)." type="book chapter" year="2023">2023: 5</bibRefCitation>
</treatmentCitation>
ZCCZ;
</treatmentCitationGroup>
</paragraph>
<paragraph id="9F4C7C56FFC14A6AFF08FCAA4759FCB5" blockId="11.[189,1034,772,802]" box="[189,1034,772,802]" pageId="11" pageNumber="12">
<taxonomicName id="58F307D5FFC14A6AFF08FCAA419BFCB6" ID-CoL="5ZCCZ" authority="(BASILEWSKY, 1955)" baseAuthorityName="Basilewsky" baseAuthorityYear="1955" box="[189,712,772,802]" class="Insecta" family="Carabidae" genus="Eoclivina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="machadoi">
<emphasis id="AD87A044FFC14A6AFF08FCAA42EDFCB6" box="[189,446,772,801]" italics="true" pageId="11" pageNumber="12">Eoclivina machadoi</emphasis>
(
<bibRefCitation id="FB6201A7FFC14A6AFE65FCAB41ECFCB6" author="BASILEWSKY P." box="[464,703,772,802]" pageId="11" pageNumber="12" pagination="93 - 137" refId="ref8356" refString="BASILEWSKY P., 1955. - Carabidae (Coleoptera) de l'Angola. Publicacoes culturais Companhia de Diamantes de Angola, Lisboa, 27: 93 - 137." type="journal article" year="1955">BASILEWSKY, 1955</bibRefCitation>
)
</taxonomicName>
, BALKENOHL, 2022: 108.
</paragraph>
</subSubSection>
<subSubSection id="D7E92FDDFFC14A6AFF08FC9742CAFAC5" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="9F4C7C56FFC14A6AFF08FC974087FC7E" blockId="11.[189,1399,824,1001]" pageId="11" pageNumber="12">
<materialsCitation id="2F9B760BFFC14A6AFF08FC974087FC7E" collectingDate="1948-09" collectionCode="MRACT" country="Angola" determinerName="P. Basilewsky" location="Dundo Foret" municipality="Luachimo" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
TYPE MATERIAL.
<typeStatus id="4048C2F4FFC14A6AFE3AFC964158FCC2" box="[399,523,824,853]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
:
<specimenCount id="89F5B7DFFFC14A6AFDACFC974162FCC2" box="[537,561,825,853]" count="1" pageId="11" pageNumber="12" type="female">♀</specimenCount>
, with white black printed and handwritten in black ink label and data: “
<collectingCountry id="E7E43CC6FFC14A6AFEB8FCF3423CFCED" box="[269,367,861,890]" name="Angola" pageId="11" pageNumber="12">Angola</collectingCountry>
:
<location id="9A2C2A8DFFC14A6AFE33FCF3417BFCED" LSID="urn:lsid:plazi:treatment:175ACD40FFC14A6DFDB8FDA142BBFCA5:9A2C2A8DFFC14A6AFE33FCF3417BFCED" box="[390,552,861,890]" country="Angola" municipality="Luachimo" name="Dundo Foret" pageId="11" pageNumber="12">Dundo Forêt</location>
(
<collectingMunicipality id="7F28E62CFFC14A6AFD82FCF341EEFCED" box="[567,701,861,890]" pageId="11" pageNumber="12">Luachimo</collectingMunicipality>
)
<date id="EB4D5A96FFC14A6AFD79FCF0406EFCED" box="[716,829,861,890]" pageId="11" pageNumber="12" value="1948-09">
<collectingDate id="FB09A37EFFC14A6AFD79FCF0406EFCED" box="[716,829,861,890]" pageId="11" pageNumber="12" value="1948-09">IX. 1948</collectingDate>
</date>
A.deBarrosMachado” / handwritten in black ink: “11 30.4” / red, black framed and printed: “HOLOTYPUS” / white, handwritten in black ink and printed: “
<taxonomicName id="58F307D5FFC14A6AFE12FC0941C5FC53" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[423,662,935,964]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">Clivina machadoi</taxonomicName>
,
<taxonomicNameLabel id="B6B41D3FFFC14A6AFD12FC0941B0FC54" box="[679,739,935,963]" pageId="11" pageNumber="12" rank="species">n.sp.</taxonomicNameLabel>
<determinerName id="6A909092FFC14A6AFD5AFC0940F5FC53" box="[751,934,935,964]" pageId="11" pageNumber="12">P. Basilewsky</determinerName>
det., 1953” / white, black printed: “RMCA ENT 000019856” and a bar code (
<figureCitation id="07C860D3FFC14A6AFD52FC624060FC7E" box="[743,819,972,1001]" captionStart="Figs 1-5" captionStartId="5.[189,235,1963,1987]" captionTargetBox="[260,1326,243,1919]" captionTargetId="figure-9@5.[210,1378,212,1963]" captionTargetPageId="5" captionText="Figs 1-5. Eoclivina species, habitus, dorsal view. 1, E. dumolinii (Putzeys, 1846), ♂, specimen from Velingara, Guinea. 2, E. basilewskyi sp. nov., holotype. 3, E. machadoi (Basilewsky, 1955), holotype. 4, E. burgeoni (Kult, 1959), holotype. 5, E. schmidli sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10981866" httpUri="https://zenodo.org/record/10981866/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
) (
<collectionCode id="F9E2E493FFC14A6AFCFBFC62409BFC7E" box="[846,968,972,1001]" pageId="11" pageNumber="12">MRACT</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="9F4C7C56FFC14A6AFF08FBAE4652FB1C" blockId="11.[189,1399,1023,1163]" pageId="11" pageNumber="12">
<materialsCitation id="2F9B760BFFC14A6AFF08FBAE4061FB8A" box="[189,818,1023,1053]" collectingDate="1948-09" collectionCode="MRACT" country="Angola" determinerName="P. Basilewsky" location="Dundo Foret" municipality="Luachimo" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="4048C2F4FFC14A6AFF08FBAE426CFB8B" box="[189,319,1024,1052]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="89F5B7DFFFC14A6AFEFEFBAE422EFB8B" box="[331,381,1024,1053]" count="1" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
, same data as holotype (
<collectionCode id="F9E2E493FFC14A6AFD1AFBAE407AFB8A" box="[687,809,1024,1053]" pageId="11" pageNumber="12">MRACT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2F9B760BFFC14A6AFC8AFBAE47AEFB1C" collectingDate="1949-05" collectionCode="MRACT" collectorName="E. de Dundo" country="Angola" determinerName="P. Basilewsky" location="Angola" pageId="11" pageNumber="12" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="89F5B7DFFFC14A6AFC8AFBAE4022FB8B" box="[831,881,1024,1053]" count="1" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
,
<specimenCount id="89F5B7DFFFC14A6AFCCAFBAE40E3FB8B" box="[895,944,1024,1053]" count="1" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
, with white black printed label and data: “
<collectingCountry id="E7E43CC6FFC14A6AFEBAFB8A4226FBD6" box="[271,373,1060,1089]" name="Angola" pageId="11" pageNumber="12">Angola</collectingCountry>
: Riv. Chiumbe,
<quantity id="580BD1B3FFC14A6AFDF1FB8A41C6FBD6" box="[580,661,1060,1089]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="12" unit="km" value="40.0">40 km</quantity>
<collectorName id="32061980FFC14A6AFD28FB8B406CFBD6" box="[669,831,1060,1089]" pageId="11" pageNumber="12">E. de Dundo</collectorName>
<date id="EB4D5A96FFC14A6AFCF2FB8A40F6FBD6" box="[839,933,1060,1089]" pageId="11" pageNumber="12" value="1949-05">
<collectingDate id="FB09A37EFFC14A6AFCF2FB8A40F6FBD6" box="[839,933,1060,1089]" pageId="11" pageNumber="12" value="1949-05">V-1949</collectingDate>
</date>
A.deBarrosMachado” / handwritten in black ink: “1493.18” / red, black framed and printed: “
<typeStatus id="4048C2F4FFC14A6AFC3EFBE7471BFBF1" box="[907,1096,1097,1126]" pageId="11" pageNumber="12" type="paratype">PARATYPUS</typeStatus>
” / white, handwritten in black ink and printed: “
<taxonomicName id="58F307D5FFC14A6AFE5DFBC04182FB1C" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[488,721,1134,1163]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">Clivina machadoi</taxonomicName>
,
<taxonomicNameLabel id="B6B41D3FFFC14A6AFD6BFBC14049FB1C" box="[734,794,1135,1163]" pageId="11" pageNumber="12" rank="species">n.sp.</taxonomicNameLabel>
<determinerName id="6A909092FFC14A6AFC97FBC14087FB1C" box="[802,980,1134,1163]" pageId="11" pageNumber="12">P. Basilewsky</determinerName>
det., 1954” (
<collectionCode id="F9E2E493FFC14A6AFBCFFBC047A7FB1C" box="[1146,1268,1134,1163]" pageId="11" pageNumber="12">MRACT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="9F4C7C56FFC14A6AFF08FB0C42CAFAC5" blockId="11.[189,1399,1186,1362]" pageId="11" pageNumber="12">
<materialsCitation id="2F9B760BFFC14A6AFF08FB0C40E4FB73" collectionCode="MRAC" collectorName="R. Cambuacala &amp; Dundo A. de Barros Machado" country="Angola" elevation="720" latitude="-7.2" location="Angola" longLatPrecision="782" longitude="20.52" pageId="11" pageNumber="12" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
ADDITIONNAL MATERIAL.
<collectingCountry id="E7E43CC6FFC14A6AFDB3FB0C41C2FB28" box="[518,657,1186,1215]" name="Angola" pageId="11" pageNumber="12">ANGOLA</collectingCountry>
.
<specimenCount id="89F5B7DFFFC14A6AFD28FB0C419CFB29" box="[669,719,1186,1215]" count="1" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
,
<specimenCount id="89F5B7DFFFC14A6AFD68FB0C405EFB29" box="[733,781,1186,1215]" count="1" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
,
<location id="9A2C2A8DFFC14A6AFCAEFB0C402DFB28" LSID="urn:lsid:plazi:treatment:175ACD40FFC14A6DFDB8FDA142BBFCA5:9A2C2A8DFFC14A6AFCAEFB0C402DFB28" box="[795,894,1186,1215]" country="Angola" latitude="-7.2" longLatPrecision="782" longitude="20.52" name="Angola" pageId="11" pageNumber="12">Angola</location>
: 15908.6
<geoCoordinate id="FAC71A91FFC14A6AFC4FFB0C471DFB28" box="[1018,1102,1186,1215]" degrees="7.20" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-7.2">7.20 S</geoCoordinate>
,
<geoCoordinate id="FAC71A91FFC14A6AFBE9FB0C4790FB29" box="[1116,1219,1186,1215]" degrees="20.52" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="20.52">20.52 E</geoCoordinate>
; alt.
<quantity id="580BD1B3FFC14A6AFAB5FB0C4602FB29" box="[1280,1361,1186,1215]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="11" pageNumber="12" unit="m" value="720.0">
<elevation id="14DE9B65FFC14A6AFAB5FB0C4602FB29" box="[1280,1361,1186,1215]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="11" pageNumber="12" unit="m" value="720.0">720 m</elevation>
</quantity>
<collectorName id="32061980FFC14A6AFAECFB0C420CFB73" pageId="11" pageNumber="12">R. Cambuacala</collectorName>
, env.
<collectorName id="32061980FFC14A6AFE19FB694067FB73" box="[428,820,1223,1252]" pageId="11" pageNumber="12">Dundo A. de Barros Machado</collectorName>
(
<collectionCode id="F9E2E493FFC14A6AFCFDFB6940FDFB73" box="[840,942,1223,1252]" country="Belgium" name="Musée Royal de l’Afrique Centrale" pageId="11" pageNumber="12" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2F9B760BFFC14A6AFC73FB694620FB73" box="[966,1395,1223,1252]" collectionCode="MRAC" country="Angola" pageId="11" pageNumber="12" specimenCount="2" specimenCount-male="2">
<specimenCount id="89F5B7DFFFC14A6AFC73FB694740FB74" box="[966,1043,1223,1252]" count="2" pageId="11" pageNumber="12" type="male">2 ♂♂</specimenCount>
, ANG. 14614.14 (
<collectionCode id="F9E2E493FFC14A6AFAB1FB694639FB73" box="[1284,1386,1223,1252]" country="Belgium" name="Musée Royal de l’Afrique Centrale" pageId="11" pageNumber="12" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="2F9B760BFFC14A6AFF08FB4242C7FAC5" collectingDate="2010-05-11" collectionCode="RBINS" collectorName="P. Grootaert" country="Democratic Republic of the Congo" latitude="2.0424917" location="DEMOCRATIC REPUBLIC OF THE CONGO" longLatPrecision="1" longitude="22.79058" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1">
<collectingCountry id="E7E43CC6FFC14A6AFF08FB424069FA9E" box="[189,826,1260,1289]" name="Democratic Republic of the Congo" pageId="11" pageNumber="12">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
.
<specimenCount id="89F5B7DFFFC14A6AFCFEFB4240D2FA9F" box="[843,897,1260,1289]" count="1" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
,
<collectorName id="32061980FFC14A6AFC23FB454670FA9F" box="[918,1315,1259,1289]" pageId="11" pageNumber="12">Coll. I.R.Sc.N.B. Congo D.R.</collectorName>
<collectorName id="32061980FFC14A6AFA84FB424625FA9F" box="[1329,1398,1260,1288]" pageId="11" pageNumber="12">Kona</collectorName>
<geoCoordinate id="FAC71A91FFC14A6AFF08FABE4226FABA" box="[189,373,1296,1325]" degrees="2" direction="north" minutes="02" orientation="latitude" pageId="11" pageNumber="12" precision="1" seconds="32.97" value="2.0424917">2°02’32.97”N</geoCoordinate>
,
<geoCoordinate id="FAC71A91FFC14A6AFE3CFABE4119FABA" box="[393,586,1296,1325]" degrees="22" direction="east" minutes="47" orientation="longitude" pageId="11" pageNumber="12" precision="1" seconds="26.09" value="22.79058">22°47’26.09”E</geoCoordinate>
<date id="EB4D5A96FFC14A6AFDEFFABE418AFABA" box="[602,729,1296,1325]" pageId="11" pageNumber="12" value="2010-05-11">
<collectingDate id="FB09A37EFFC14A6AFDEFFABE418AFABA" box="[602,729,1296,1325]" pageId="11" pageNumber="12" value="2010-05-11">11.v.2010</collectingDate>
</date>
,
<collectingMethod id="46B20441FFC14A6AFD5BFABF4769FABA" box="[750,1082,1296,1325]" pageId="11" pageNumber="12">swamp forest Light trap</collectingMethod>
, leg.
<collectorName id="32061980FFC14A6AFB39FABF467EFABA" box="[1164,1325,1296,1325]" pageId="11" pageNumber="12">
P.
<collectorName id="32061980FFC14A6AFB07FABE4622FABA" box="[1202,1393,1296,1325]" pageId="11" pageNumber="12">Grootaert I.G.</collectorName>
</collectorName>
: 33.593 (
<collectionCode id="F9E2E493FFC14A6AFE93FA9B42D8FAC5" box="[294,395,1333,1362]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="12" type="Museum">RBINS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D7E92FDDFFC14A6DFF08FAC4421FFD4C" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="9F4C7C56FFC14A6AFF08FAC44036F845" blockId="11.[189,1399,1385,2003]" pageId="11" pageNumber="12">
EXTENDED DIAGNOSIS. A medium sized piceous species with the following measurements: Body length
<quantity id="580BD1B3FFC14A6AFED3FA204141FA3D" box="[358,530,1422,1451]" metricMagnitude="-3" metricUnit="m" metricValue="5.3149999999999995" metricValueMax="5.53" metricValueMin="5.1" pageId="11" pageNumber="12" unit="mm" value="5.3149999999999995" valueMax="5.53" valueMin="5.1">5.1–5.53 mm</quantity>
(x =
<quantity id="580BD1B3FFC14A6AFDE8FA20418BFA3C" box="[605,728,1422,1451]" metricMagnitude="-3" metricUnit="m" metricValue="5.39" pageId="11" pageNumber="12" unit="mm" value="5.39">5.39 mm</quantity>
*), width
<quantity id="580BD1B3FFC14A6AFCE4FA20475EFA3D" box="[849,1037,1422,1451]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" metricValueMax="1.61" metricValueMin="1.51" pageId="11" pageNumber="12" unit="mm" value="1.56" valueMax="1.61" valueMin="1.51">1.51–1.61 mm</quantity>
(x =
<quantity id="580BD1B3FFC14A6AFBE2FA204781FA3C" box="[1111,1234,1422,1451]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="11" pageNumber="12" unit="mm" value="1.58">1.58 mm</quantity>
*), ratio L/W of pronotum 0.95–0.99 (x = 0.97*), ratio L/W of elytra 1.82–1.87 (x = 1.84*); (*n = 9). The elytron is unicolor. Wing of the clypeus relatively wide but nearly fused with lateral lobe of the clypeus. Eye convex and projecting laterally. Pronotum with lateral margin slightly attenuating anteriorly with straight parts anterior middle, on the disk with scattered small to moderate sized punctures, punctures larger towards base, proepisternum just not visible in dorsal view. Elytra with the shape long-oval with straight and diverging part of the lateral margins, with maximum width slightly behind middle, intervals with different pattern of reticulation among the sexes: In males, interval eight reticulated, all the others are smooth and glossy. In females, the reticulation covers intervals one, and six to eight. In addition, the base is reticulated as well as around a fifth of the apex. Intervals two to five smooth on the disk. Row of umbilical setigerous punctures of the lateral channel interrupted posterior humerus and at middle by one puncture. Abdominal sternite in males with minute punctures, smooth and glossy at middle, laterally with some smaller punctures, sternite of females distinctly and completely covered with transverse reticulation. Intermediate tibia with six tubercles at the inner side and eight at the lateral side, tubercles furnished with strong setae.
</paragraph>
<paragraph id="9F4C7C56FFC14A6DFF08F844473AFE81" blockId="11.[189,555,2026,2055]" lastBlockId="12.[189,1398,212,278]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="AD87A044FFC14A6AFF08F8444210F790" box="[189,323,2026,2055]" italics="true" pageId="11" pageNumber="12">Hind wing</emphasis>
: Fully developed.
<emphasis id="AD87A044FFC64A6DFF08FF7A422EFF66" box="[189,381,212,241]" italics="true" pageId="12" pageNumber="13">Male genitalia</emphasis>
(
<figureCitation id="07C860D3FFC64A6DFE26FF7A42B5FF66" box="[403,486,212,241]" captionStart="Figs 6-9" captionStartId="6.[189,235,1935,1959]" captionTargetBox="[213,1382,675,1851]" captionTargetId="figure-133@6.[189,1399,628,1919]" captionTargetPageId="6" captionText="Figs 6-9. Eoclivina species, male genitalia. A-B, aedeagus. A, dorsal view. B, lateral view. C-D, parameres, dorsal view. 6, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 7, E. basilewskyi sp. nov., holotype. 8, E. machadoi (Basilewsky, 1955), paratype from Riv. Chiumbe, Angola. 9, E. schmidli sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10981868" httpUri="https://zenodo.org/record/10981868/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
): Median lobe wide in lateral view, slightly arcuate, with small and rounded spatula. Endophallus with longitudinal group of bristles apically.
</paragraph>
<paragraph id="9F4C7C56FFC64A6DFF08FE82401EFE07" blockId="12.[189,1398,300,400]" pageId="12" pageNumber="13">
<emphasis id="AD87A044FFC64A6DFF08FE8241D3FEDE" box="[189,640,300,329]" italics="true" pageId="12" pageNumber="13">Female gonocoxite and epipleurite</emphasis>
(
<figureCitation id="07C860D3FFC64A6DFD26FE8241A7FEDE" box="[659,756,300,329]" captionStart="Figs 10-14" captionStartId="7.[189,235,1966,1990]" captionTargetBox="[228,1363,216,1876]" captionTargetId="figure-9@7.[204,1374,212,1967]" captionTargetPageId="7" captionText="Figs 10-14. Eoclivina species, females, gonocoxite and epipleurite, ventro-lateral view. 10, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 11, E. basilewskyi sp. nov., paratype. 12, E. machadoi (Basilewsky, 1955), paratype. 13, E. burgeoni (Kult, 1959), holotype. 14, E. schmidli sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.10981870" httpUri="https://zenodo.org/record/10981870/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
): Gonocoxite foliform, widened, with four robust nematiform setae positioned at the median side, apically with one long seta. Epipleurite quadrangular, asetose, with widened angulated rod.
</paragraph>
<paragraph id="9F4C7C56FFC64A6DFF08FE084792FE70" blockId="12.[189,1399,422,487]" pageId="12" pageNumber="13">
<emphasis id="AD87A044FFC64A6DFF08FE0841C4FE54" box="[189,663,422,451]" italics="true" pageId="12" pageNumber="13">Sexual dimorphism at the integument</emphasis>
: Males and females differ considerably in the reticulation on the intervals of the elytron and the abdominal sternites (see diagnosis above).
</paragraph>
<paragraph id="9F4C7C56FFC64A6DFF08FE53464AFDF6" blockId="12.[189,1399,509,609]" pageId="12" pageNumber="13">VARIATION. On the disk of the pronotum, the punctures are more or less dense and vary in its distinctness. In some of the females, the reticulation on the intervals of the elytron is not strongly expressed on interval six. In others, it extends slightly to interval two and five.</paragraph>
<paragraph id="9F4C7C56FFC64A6DFF08FDD9421FFD4C" blockId="12.[189,1398,631,731]" pageId="12" pageNumber="13">
The distinguishing character from the similar species
<taxonomicName id="58F307D5FFC64A6DFCC9FDD64732FD03" authority="KULT" authorityName="Kult" authorityYear="1959" box="[892,1121,631,660]" class="Insecta" family="Carabidae" genus="Eoclivina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="burgeoni">
<emphasis id="AD87A044FFC64A6DFCC9FDD64740FD03" box="[892,1043,631,660]" italics="true" pageId="12" pageNumber="13">E. burgeoni</emphasis>
KULT
</taxonomicName>
is the reticulation on the intervals of the elytron. In the female
<typeStatus id="4048C2F4FFC64A6DFD62FD354014FD2F" box="[727,839,667,696]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
of
<taxonomicName id="58F307D5FFC64A6DFCC4FD32475BFD2F" authorityName="Kult" authorityYear="1959" box="[881,1032,667,696]" class="Insecta" family="Carabidae" genus="Eoclivina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="burgeoni">
<emphasis id="AD87A044FFC64A6DFCC4FD32475BFD2F" box="[881,1032,667,696]" italics="true" pageId="12" pageNumber="13">E. burgeoni</emphasis>
</taxonomicName>
, all intervals are completely reticulated.
</paragraph>
</subSubSection>
<subSubSection id="D7E92FDDFFC64A6DFF08FD5F42BBFCA5" pageId="12" pageNumber="13" type="distribution">
<paragraph id="9F4C7C56FFC64A6DFF08FD5F42BBFCA5" blockId="12.[189,1398,753,818]" pageId="12" pageNumber="13">
DISTRIBUTION. The species is found in the North and West of
<collectingCountry id="E7E43CC6FFC64A6DFC51FD5F4716FC99" box="[996,1093,753,782]" name="Angola" pageId="12" pageNumber="13">Angola</collectingCountry>
, and in
<collectingCountry id="E7E43CC6FFC64A6DFB05FD5F42B0FCA5" name="Democratic Republic of the Congo" pageId="12" pageNumber="13">the Democratic Republic of the Congo</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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