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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1008.61040" ID-GBIF-Dataset="6883cbae-4fcb-4a05-ab30-f0673672c13d" ID-PMC="PMC7790813" ID-Pensoft-Pub="1313-2970-1008-61" ID-Pensoft-UUID="F43798A25D4A53C8B58BACF7281D0CBD" ID-PubMed="33505188" ID-ZooBank="E17BB2E712A94F4295956082848B8FC9" ModsDocID="1313-2970-1008-61" checkinTime="1609463508641" checkinUser="pensoft" docAuthor="Tian, Mingyi &amp; He, Li" docDate="2020" docId="EE95CFE4E6B55F048E3EF4D17ED7ECE9" docLanguage="en" docName="ZooKeys 1008: 61-91" docOrigin="ZooKeys 1008" docPubDate="2020-12-31" docSource="http://dx.doi.org/10.3897/zookeys.1008.61040" docTitle="Uenoaphaenops Tian &amp; He 2020, gen. nov." docType="treatment" docUuid="20CF5C96-DFC5-419F-97D8-924D18433C3F" docUuidSource="ZooBank" docVersion="4" id="F43798A25D4A53C8B58BACF7281D0CBD" lastPageNumber="61" masterDocId="F43798A25D4A53C8B58BACF7281D0CBD" masterDocTitle="A contribution to the knowledge of cavernicolous ground beetles from Sichuan Province, southwestern China (Coleoptera, Carabidae, Trechini, Platynini)" masterLastPageNumber="91" masterPageNumber="61" pageNumber="61" updateTime="1668149672560" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A contribution to the knowledge of cavernicolous ground beetles from Sichuan Province, southwestern China (Coleoptera, Carabidae, Trechini, Platynini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Mingyi</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2823-7619</mods:nameIdentifier>
<mods:affiliation>Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, China</mods:affiliation>
<mods:nameIdentifier type="email">mytian168@aliyun.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>He, Li</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4597-3442</mods:nameIdentifier>
<mods:affiliation>Sichuan Cave Exploration Team, No. 66, 5 th Shuangcheng Road, Chenghua District, Chengdu, 610051, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1008</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>91</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1008.61040</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1008.61040</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1008-61</mods:identifier>
<mods:identifier type="ZooBank">E17BB2E712A94F4295956082848B8FC9</mods:identifier>
<mods:identifier type="Pensoft-UUID">F43798A25D4A53C8B58BACF7281D0CBD</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="176352580" LSID="urn:lsid:zoobank.org:act:20CF5C96-DFC5-419F-97D8-924D18433C3F" httpUri="http://treatment.plazi.org/id/EE95CFE4E6B55F048E3EF4D17ED7ECE9" lastPageNumber="61" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="http://zoobank.org/20CF5C96-DFC5-419F-97D8-924D18433C3F" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">Uenoaphaenops</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="61">gen. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map of Sichuan Province showing the locations of the related caves." figureDoi="10.3897/zookeys.1008.61040.figure2" httpUri="https://binary.pensoft.net/fig/492963" pageId="0" pageNumber="61">
Chinese name:
<normalizedToken originalValue="上野穴步甲属">上野穴步甲属</normalizedToken>
Figs 2
</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Uenoaphaenops fani (Ueno, 2003), comb. nov., female." figureDoi="10.3897/zookeys.1008.61040.figure3" httpUri="https://binary.pensoft.net/fig/492964" pageId="0" pageNumber="61">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cave trechine beetles A ventral head of Uenoaphaenops fani (Ueno, 2003), comb. nov., female B ventral head of Chu pheggomisetoides gen. nov. &amp; sp. nov., female C-E elytral chaetotaxy of Uenoaphaenops fani (Ueno, 2003), comb. nov., female, Chu pheggomisetoides gen. nov. &amp; sp. nov., female paratype, and Boreaphaenops liyuani sp. nov., female holotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.61040.figure4" httpUri="https://binary.pensoft.net/fig/492965" pageId="0" pageNumber="61">, 4A, C</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of cave trechine species, lateral and dorsal views A, B Uenoaphaenops fani (Ueno, 2003), comb. nov. C, D Chu pheggomisetoides gen. nov. &amp; sp. nov." figureDoi="10.3897/zookeys.1008.61040.figure5" httpUri="https://binary.pensoft.net/fig/492966" pageId="0" pageNumber="61">, 5A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="type species">
<paragraph pageId="0" pageNumber="61">Type species.</paragraph>
<paragraph pageId="0" pageNumber="61">
<taxonomicName authorityName="Ueno" authorityYear="2003" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 2003 from the cave
<normalizedToken originalValue="Hua’er">Hua'er</normalizedToken>
Dong, Gulin, southeastern Sichuan).
</paragraph>
<caption doi="10.3897/zookeys.1008.61040.figure3" httpUri="https://binary.pensoft.net/fig/492964" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
Habitus of
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov., female.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Generic characteristics.</paragraph>
<paragraph pageId="0" pageNumber="61">
Medium-sized, aphaenopsian and depigmented; body moderately elongate, wholly pubescent. Head strongly elongate, much longer than wide, nearly parallel-sided; neck weakly-marked, ring-shaped; two pairs of supraorbital setiferous pores present; frontal furrows incomplete, parallel-sided in most part though briefly divergent posteriorly; frons and vertex convex; right mandibular tooth bidentate; mentum and submentum completely fused; mentum bisetose, and covered with short setae on basal area of mental tooth, and along the site of labial suture; base largely concave, uni-foveate, tooth short and simple at tip; submentum with a row of 12 setae; labial palpomere 2 much longer than 3 (1.3 times), bisetose on inner margin, without additional setae; maxillary palpomere 3 much longer than 4 (1.4 times) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cave trechine beetles A ventral head of Uenoaphaenops fani (Ueno, 2003), comb. nov., female B ventral head of Chu pheggomisetoides gen. nov. &amp; sp. nov., female C-E elytral chaetotaxy of Uenoaphaenops fani (Ueno, 2003), comb. nov., female, Chu pheggomisetoides gen. nov. &amp; sp. nov., female paratype, and Boreaphaenops liyuani sp. nov., female holotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.61040.figure4" httpUri="https://binary.pensoft.net/fig/492965" pageId="0" pageNumber="61">4A</figureCitation>
); antennae long and thin, extending to about 1/4 of elytra from apex; one pair of suborbital pores present. Prothorax distinctly expanded, propleura visible from above; pronotum elongate, as long as head excluding mandibles, wider than head, widest at about 1/3 from front, lateral margins sinuate before hind angles which is nearly rectangular, base slightly narrower than front; two pairs of latero-marginal setae present, disc moderately convex. Elytra ovate, shoulders obtuse, almost rounded, distinctly serrate at prehumeral part, while ciliate on other parts; disc extraordinarily convex though depressed near base; striae reduced though traceable; presence of two dorsal pores along the 3rd striae and the preapical; prehumeral set of the marginal umbilicate pores not aggregated, the 5th pore much closer to 4th than to 6th. Protarsi not modified in male; tibiae without longitudinal sulci.
<taxonomicName genus="Ventrites" lsidName="Ventrites" pageId="0" pageNumber="61" rank="genus">Ventrites</taxonomicName>
IV-VI each with pair of paramedian setae; ventrite VII quadrisetose in both sexes.
</paragraph>
<caption doi="10.3897/zookeys.1008.61040.figure4" httpUri="https://binary.pensoft.net/fig/492965" pageId="0" pageNumber="61" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
Cave trechine beetles
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
ventral head of
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov., female
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
ventral head of
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Chu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chu pheggomisetoides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pheggomisetoides">
<emphasis italics="true" pageId="0" pageNumber="61">Chu pheggomisetoides</emphasis>
</taxonomicName>
gen. nov. &amp; sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="61">C-E</emphasis>
elytral chaetotaxy of
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov., female,
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Chu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chu pheggomisetoides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pheggomisetoides">
<emphasis italics="true" pageId="0" pageNumber="61">Chu pheggomisetoides</emphasis>
</taxonomicName>
gen. nov. &amp; sp. nov., female paratype, and
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Boreaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boreaphaenops liyuani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="liyuani">
<emphasis italics="true" pageId="0" pageNumber="61">Boreaphaenops liyuani</emphasis>
</taxonomicName>
sp. nov., female holotype. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of cave trechine species, lateral and dorsal views A, B Uenoaphaenops fani (Ueno, 2003), comb. nov. C, D Chu pheggomisetoides gen. nov. &amp; sp. nov." figureDoi="10.3897/zookeys.1008.61040.figure5" httpUri="https://binary.pensoft.net/fig/492966" pageId="0" pageNumber="61">5A, B</figureCitation>
). Aedeagus very short and small, but thick, weakly sclerotized. The median lobe slightly arcuate at median portion, but strongly sinuate before apex which is obtuse, with a large round opening; base quite large, without a sagittal aileron; inner sac provided with a thick and long copulatory piece, which is about 2/5 as long as aedeagus; in dorsal view, apical lobe gradually contracted towards apex which is broadly rounded; parameres well-developed, but much shorter than the median lobe, truncate at apical margin, each armed with four long setae at apex.
</paragraph>
<caption doi="10.3897/zookeys.1008.61040.figure5" httpUri="https://binary.pensoft.net/fig/492966" pageId="0" pageNumber="61" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 5.</emphasis>
Male genitalia of cave trechine species, lateral and dorsal views
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov.
<emphasis bold="true" pageId="0" pageNumber="61">C, D</emphasis>
<taxonomicName authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Chu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chu pheggomisetoides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pheggomisetoides">
<emphasis italics="true" pageId="0" pageNumber="61">Chu pheggomisetoides</emphasis>
</taxonomicName>
gen. nov. &amp; sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="remarks">
<paragraph pageId="0" pageNumber="61">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="61">
This peculiar species was put into the genus
<taxonomicName authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 2000 due to the lack of a male at that time. However,
<bibRefCitation author="Ueno, SI" journalOrPublisher="Journal of the Speleological Society of Japan" pageId="0" pageNumber="61" pagination="1 - 11" refId="B15" refString="Ueno, SI, 2003. Two new cave trechines of the genus Qianotrechus (Coleoptera, Trechinae). Journal of the Speleological Society of Japan 28: 1 - 11" title="Two new cave trechines of the genus Qianotrechus (Coleoptera, Trechinae)." volume="28" year="2003">
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
(2003)
</bibRefCitation>
pointed out the following characteristics of this species which are not present in the congeners of
<taxonomicName authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
: body wholly pubescent; humeral margins of elytra strongly serrated; and the 5th pore of marginal umbilicate series forwardly and inwardly shifted, widely distant from the 6th pore. Hence, he mentioned that the above peculiarities may suggest a generic separation of this species from the Guizhou genus. Our examination of male individuals provided further evidence to support his opinion. First, protarsi are not modified in the male of
<taxonomicName authorityName="Ueno" authorityYear="2003" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
, while the 1st and 2nd protarsomeres are spurred inwards at the apices in all other
<taxonomicName authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
. Second, ventrite VII is quadrisetose in both sexes in
<taxonomicName authorityName="Ueno" authorityYear="2003" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
, vs. bisetose in males of other
<taxonomicName authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
. Third, the aedeagus of
<taxonomicName authorityName="Ueno" authorityYear="2003" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
is very small and stout, not the same type as in other
<taxonomicName authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
species, which are always large and elongate (
<bibRefCitation author="Ueno, SI" journalOrPublisher="Journal of the Speleological Society of Japan" pageId="0" pageNumber="61" pagination="1 - 38" refId="B13" refString="Ueno, SI, 2000. New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China. Journal of the Speleological Society of Japan 25: 1 - 38" title="New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China." volume="25" year="2000">
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
2000
</bibRefCitation>
,
<bibRefCitation author="Ueno, SI" journalOrPublisher="Journal of the Speleological Society of Japan" pageId="0" pageNumber="61" pagination="1 - 11" refId="B15" refString="Ueno, SI, 2003. Two new cave trechines of the genus Qianotrechus (Coleoptera, Trechinae). Journal of the Speleological Society of Japan 28: 1 - 11" title="Two new cave trechines of the genus Qianotrechus (Coleoptera, Trechinae)." volume="28" year="2003">2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="etymology">
<paragraph pageId="0" pageNumber="61">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="61">
&quot;
<emphasis italics="true" pageId="0" pageNumber="61">Ueno</emphasis>
&quot;+ &quot;-
<emphasis italics="true" pageId="0" pageNumber="61">aphaenops</emphasis>
&quot;. Dedicated to the late Dr Shun-Ichi
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
who made a great contribution to the knowledge of Chinese subterranean ground beetles. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="generic range">
<paragraph pageId="0" pageNumber="61">Generic range.</paragraph>
<paragraph pageId="0" pageNumber="61">
China (Sichuan) (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map of Sichuan Province showing the locations of the related caves." figureDoi="10.3897/zookeys.1008.61040.figure2" httpUri="https://binary.pensoft.net/fig/492963" pageId="0" pageNumber="61">2</figureCitation>
). A monospecific genus only recorded from the cave
<normalizedToken originalValue="Hua’er">Hua'er</normalizedToken>
Dong, Gulin County, southeastern Sichuan.
</paragraph>
</subSubSection>
</treatment>
</document>
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