477 lines
60 KiB
XML
477 lines
60 KiB
XML
<document id="570689016E29E705FB5ED58EFE3F928D" ID-CLB-Dataset="31484" ID-DOI="10.11646/zootaxa.4083.4.5" ID-GBIF-Dataset="e5e50995-178e-4a41-b636-7f54f566d5a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050911" ID-ZooBank="5FF5AC63-D26C-4456-84ED-B9DD218C100D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510812312033" checkinUser="plazi" docAuthor="Last, Peter R., White, William T. & Séret, Bernard" docDate="2016" docId="5F0687AAB361FF98FF48CEC9FA1DEC2E" docLanguage="en" docName="zootaxa.4083.4.5.pdf" docOrigin="Zootaxa 4083 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neotrygon kuhlii" docType="treatment" docVersion="8" lastPageNumber="540" masterDocId="A33FFFD2B363FF90FFDFCA3CFF93ED6F" masterDocTitle="Taxonomic status of maskrays of the Neotrygon kuhlii species complex (Myliobatoidei: Dasyatidae) with the description of three new species from the Indo-West Pacific" masterLastPageNumber="561" masterPageNumber="533" pageNumber="535" updateTime="1698496900406" updateUser="ExternalLinkService">
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<mods:title id="AE5989B0F3F575F7A9F40B0FA98494C2">Taxonomic status of maskrays of the Neotrygon kuhlii species complex (Myliobatoidei: Dasyatidae) with the description of three new species from the Indo-West Pacific</mods:title>
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<mods:namePart id="0360A62B2750DC4C39C60B6CD10B890F">Last, Peter R.</mods:namePart>
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<mods:namePart id="7ED0F8D3E3A76149DCE7E30FA22EF85C">White, William T.</mods:namePart>
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<mods:namePart id="439B99420830A2618D98F6165FA04A36">Séret, Bernard</mods:namePart>
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<mods:title id="1167E1C7634E56629A8EFB451136D6BE">Zootaxa</mods:title>
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<mods:part id="B267C436F3EF0F19DE47078C4599761C">
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<mods:date id="A07EBE3772B69F84D23B895181D3C143">2016</mods:date>
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<mods:number id="143B96CE834B48A28741F78B663822F5">2016-02-23</mods:number>
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<mods:number id="EE03155D21C4875E1E1C2DE5936D4FC6">4083</mods:number>
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<treatment id="5F0687AAB361FF98FF48CEC9FA1DEC2E" ID-DOI="http://doi.org/10.5281/zenodo.6079663" ID-GBIF-Taxon="137307641" ID-Zenodo-Dep="6079663" LSID="urn:lsid:plazi:treatment:5F0687AAB361FF98FF48CEC9FA1DEC2E" httpUri="http://treatment.plazi.org/id/5F0687AAB361FF98FF48CEC9FA1DEC2E" lastPageId="8" lastPageNumber="540" pageId="2" pageNumber="535">
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<subSubSection id="9FB56537B361FF92FF48CEC9FD9BE85E" pageId="2" pageNumber="535" type="nomenclature">
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<paragraph id="D71036BCB361FF92FF48CEC9FD1CE87F" blockId="2.[151,655,1269,1329]" box="[151,655,1269,1296]" pageId="2" pageNumber="535">
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<heading id="8C5881D0B361FF92FF48CEC9FD1CE87F" bold="true" box="[151,655,1269,1296]" fontSize="11" level="1" pageId="2" pageNumber="535" reason="1">
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<taxonomicName id="10AF4D3FB361FF92FF48CEC9FD1CE87F" ID-CoL="46W32" authority="(Muller & Henle, 1841)" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[151,655,1269,1296]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="2" pageNumber="535" phylum="Chordata" rank="species" species="kuhlii">
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<emphasis id="E5DBEAAEB361FF92FF48CEC9FD1CE87F" bold="true" box="[151,655,1269,1296]" pageId="2" pageNumber="535">
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<emphasis id="E5DBEAAEB361FF92FF48CEC9FEF6E860" bold="true" box="[151,357,1269,1295]" italics="true" pageId="2" pageNumber="535">Neotrygon kuhlii</emphasis>
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(
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<bibRefCitation id="B33E4B4DB361FF92FEABCECAFD14E87F" author="Muller" box="[372,647,1270,1296]" pageId="2" pageNumber="535" refString="Muller, J., & Henle, F. G. J. (1841) Systematische Beschreibung der Plagiostomen. Berlin, pp. 103 - 200." type="book chapter" year="1841">Müller & Henle, 1841</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="D71036BCB361FF92FF48CF24FD9BE85E" blockId="2.[151,655,1269,1329]" box="[151,520,1304,1329]" pageId="2" pageNumber="535">
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(
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<figureCitation id="4F942A39B361FF92FF40CF24FF7FE85E" box="[159,236,1304,1329]" captionStart="FIGURE 1" captionStartId="2.[151,250,1167,1189]" captionText="FIGURE 1. Original illustration of Trygon kuhlii in Müller & Henle (1841)." pageId="2" pageNumber="535">Figs. 1</figureCitation>
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,
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<figureCitation id="4F942A39B361FF92FF25CF25FE99E85E" box="[250,266,1305,1329]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[349,1334,217,1920]" captionTargetId="figure@4.[151,1436,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal view of Neotrygon kuhlii types from Vanikoro, Solomon Islands (preserved): (A) lectotype (MNHN A 2440, 1 of 2, female 154 mm DW); (B) paralectotype (MNHN A 2440, 1 of 2, female 162.7 mm DW)." httpUri="https://zenodo.org/record/1051776/files/figure.png" pageId="2" pageNumber="535">2</figureCitation>
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,
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<figureCitation id="4F942A39B361FF92FEC9CF25FEB5E85E" box="[278,294,1305,1329]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[176,1394,212,1911]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Freshly caught specimen of Neotrygon kuhlii (CSIRO 7723 - 01, female 295 mm DW) from Honiara, Solomon Islands: (A) dorsal view; (B) ventral view." httpUri="https://zenodo.org/record/1051780/files/figure.png" pageId="2" pageNumber="535">4</figureCitation>
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,
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<figureCitation id="4F942A39B361FF92FEEDCF24FEDCE85E" box="[306,335,1304,1329]" captionStart="FIGURE 5" captionStartId="7.[151,250,1093,1115]" captionTargetBox="[151,1436,198,1064]" captionTargetId="figure@7.[151,1436,193,1071]" captionTargetPageId="7" captionText="FIGURE 5. Oronasal region of: (A) Neotrygon kuhlii (CSIRO 7723 - 01, female 295 mm DW); (B) Neotrygon australiae sp. nov. (CSIRO CA 3247, adult male 379 mm DW); (C) Neotrygon caeruleopunctata sp. nov. (CSIRO H 6202 - 03, adult male 311 mm DW); (D) Neotrygon orientale sp. nov. (CSIRO H 6130 - 01, female 254 mm DW)." httpUri="https://zenodo.org/record/1051782/files/figure.png" pageId="2" pageNumber="535">5a</figureCitation>
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,
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<figureCitation id="4F942A39B361FF92FE84CF24FEEBE85E" box="[347,376,1304,1329]" captionStart="FIGURE 6" captionStartId="12.[151,250,1080,1102]" captionTargetBox="[151,1429,368,1006]" captionTargetId="figure@12.[151,1436,339,1058]" captionTargetPageId="12" captionText="FIGURE 6. Lateral view of the post-caudal sting tail of: (A) Neotrygon kuhlii (CSIRO 7723 - 01, female 295 mm DW); (B) Neotrygon australiae sp. nov. (CSIRO CA 4307, female 306 mm DW); (C) Neotrygon caeruleopunctata sp. nov. (CSIRO H 6202 - 04, female 324 mm DW); (D) Neotrygon orientale sp. nov. (CSIRO H 7848 - 01, female 343 mm DW)." httpUri="https://zenodo.org/record/1051784/files/figure.png" pageId="2" pageNumber="535">6a</figureCitation>
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,
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<figureCitation id="4F942A39B361FF92FE5CCF25FE33E85F" box="[387,416,1305,1328]" captionStart="FIGURE 7" captionStartId="12.[151,250,1773,1795]" captionTargetBox="[151,1436,1264,1721]" captionTargetId="figure@12.[151,1436,1213,1751]" captionTargetPageId="12" captionText="FIGURE 7. Lateral view of the mid-tail showing respective heights of the dorsal and ventral tail folds: (A) Neotrygon kuhlii (CSIRO 7723 - 01, female 295 mm DW); (B) Neotrygon australiae sp. nov. (CSIRO CA 4307, female 306 mm DW); (C) Neotrygon caeruleopunctata sp. nov. (CSIRO H 6202 - 04, female 324 mm DW); (D) Neotrygon orientale sp. nov. (CSIRO H 7848 - 01, female 343 mm DW)." httpUri="https://zenodo.org/record/1051786/files/figure.png" pageId="2" pageNumber="535">7a</figureCitation>
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||
;
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||
<tableCitation id="9A2D0307B361FF92FE72CF24FE6EE85E" box="[429,509,1304,1329]" captionStart="TABLE 1" captionStartId="7.[151,239,1220,1243]" captionText="TABLE 1. Morphometric data for the lectotype (MNHN 2440, 1 of 2), paralectotype (MNHN 2440, 1 of 2), and one other specimen (CSIRO H 7723 - 01) of Neotrygon kuhlii, and for the holotype of Neotrygon australiae sp. nov. (CSIRO H 7016 - 01), and ranges and means for the measured paratypes. Measurements expressed as a percentage of disc width." pageId="2" pageNumber="535">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="9FB56537B361FF92FF48CF5EFE26E88F" pageId="2" pageNumber="535" type="reference_group">
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<paragraph id="D71036BCB361FF92FF48CF5EFC59E819" blockId="2.[151,970,1376,1399]" box="[151,970,1376,1399]" pageId="2" pageNumber="535">
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<taxonomicName id="10AF4D3FB361FF92FF48CF5EFD9AE819" ID-CoL="7CXZF" authority="Muller & Henle, 1841" authorityName="in Muller & Henle" authorityYear="1841" box="[151,521,1376,1399]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="2" pageNumber="535" phylum="Chordata" rank="species" species="kuhlii">
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||
<emphasis id="E5DBEAAEB361FF92FF48CF5EFE8CE819" box="[151,287,1376,1399]" italics="true" pageId="2" pageNumber="535">Trygon kuhlii</emphasis>
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<bibRefCitation id="B33E4B4DB361FF92FEFACF5CFD9AE819" author="Muller" box="[293,521,1376,1398]" pageId="2" pageNumber="535" refString="Muller, J., & Henle, F. G. J. (1841) Systematische Beschreibung der Plagiostomen. Berlin, pp. 103 - 200." type="book chapter" year="1841">Müller & Henle, 1841</bibRefCitation>
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</taxonomicName>
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||
: p. 164, pl. 50 (Vanikoro,
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||
<collectingCountry id="AFB8762CB361FF92FCCECF5CFC2DE819" box="[785,958,1376,1398]" name="Solomon Islands" pageId="2" pageNumber="535">Solomon Islands</collectingCountry>
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||
).
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||
</paragraph>
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<paragraph id="D71036BCB361FF92FF48CF9FFE26E88F" blockId="2.[151,1437,1443,2008]" pageId="2" pageNumber="535">
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<materialsCitation id="67C73CE1B361FF92FF48CF9FFE26E88F" ID-GBIF-Occurrence="1700783815" collectionCode="MNHN" country="Solomon Islands" location="Vanikoro" municipality="Santa Cruz Group" pageId="2" pageNumber="535" specimenCode="MNHN 2440" specimenCount="2" typeStatus="lectotype">
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<emphasis id="E5DBEAAEB361FF92FF48CF9FFE87E8D4" bold="true" box="[151,276,1443,1467]" pageId="2" pageNumber="535">
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<typeStatus id="0814881EB361FF92FF48CF9FFE9CE8D4" box="[151,271,1443,1467]" pageId="2" pageNumber="535" type="lectotype">Lectotype</typeStatus>
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.
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||
</emphasis>
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||
<specimenCode id="87099EC7B361FF92FEC3CF98FE23E8D3" box="[284,432,1444,1468]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="535">MNHN 2440</specimenCode>
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(1 of
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<specimenCount id="C1A9FD35B361FF92FE2ACF98FD17E8D3" box="[501,644,1443,1468]" pageId="2" pageNumber="535" type="generic">2 specimens</specimenCount>
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), female
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DW,
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,
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<collectingMunicipality id="3774ACC6B361FF92FBDFCF98FB5DE8D3" box="[1024,1230,1444,1468]" pageId="2" pageNumber="535">Santa Cruz Group</collectingMunicipality>
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,
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<collectingCountry id="AFB8762CB361FF92FB06CF9FFA0BE8D3" box="[1241,1432,1443,1468]" name="Solomon Islands" pageId="2" pageNumber="535">Solomon Islands</collectingCountry>
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, 1829 (herein designated).
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</materialsCitation>
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</paragraph>
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<subSubSection id="9FB56537B361FF92FF18CFD7FE66EB47" pageId="2" pageNumber="535" type="description">
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<paragraph id="D71036BCB361FF92FF18CFD7FE66EB47" blockId="2.[151,1437,1443,2008]" pageId="2" pageNumber="535">
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<materialsCitation id="67C73CE1B361FF92FF18CFD7FE66EB47" ID-GBIF-Occurrence="1700783798" collectionCode="MNHN" country="Solomon Islands" location="Vanikoro" municipality="Santa Cruz Group" pageId="2" pageNumber="535" specimenCode="MNHN 2440" specimenCount="2" typeStatus="paralectotype">
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<typeStatus id="0814881EB361FF92FF18CFD7FEFDEB6C" box="[199,366,1515,1539]" pageId="2" pageNumber="535" type="paralectotype">Paralectotype</typeStatus>
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.
|
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</emphasis>
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<specimenCode id="87099EC7B361FF92FEA2CFD0FD86EB6B" box="[381,533,1516,1540]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="535">MNHN 2440</specimenCode>
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(2 of
|
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<specimenCount id="C1A9FD35B361FF92FDBBCFD0FD66EB6B" box="[612,757,1515,1540]" pageId="2" pageNumber="535" type="generic">2 specimens</specimenCount>
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; tail missing), female
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<quantity id="10579B59B361FF92FC24CFD7FBCFEB6C" box="[1019,1116,1515,1540]" metricMagnitude="-1" metricUnit="m" metricValue="1.59" pageId="2" pageNumber="535" unit="mm" value="159.0">159 mm</quantity>
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DW,
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<location id="D2706067B361FF92FB7BCFD7FA9DEB6B" LSID="urn:lsid:plazi:treatment:5F0687AAB361FF98FF48CEC9FA1DEC2E:D2706067B361FF92FB7BCFD7FA9DEB6B" box="[1188,1294,1515,1540]" country="Solomon Islands" municipality="Santa Cruz Group" name="Vanikoro" pageId="2" pageNumber="535">Vanikoro</location>
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,
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<collectingMunicipality id="3774ACC6B361FF92FAC3CFD0FF71EB47" pageId="2" pageNumber="535">Santa Cruz Group</collectingMunicipality>
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,
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<collectingCountry id="AFB8762CB361FF92FF33CC33FE39EB47" box="[236,426,1551,1576]" name="Solomon Islands" pageId="2" pageNumber="535">Solomon Islands</collectingCountry>
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, 1829.
|
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9FB56537B361FF92FF18CC0EFF45EB1F" pageId="2" pageNumber="535" type="materials_examined">
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<paragraph id="D71036BCB361FF92FF18CC0EFF45EB1F" blockId="2.[151,1437,1443,2008]" pageId="2" pageNumber="535">
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<materialsCitation id="67C73CE1B361FF92FF18CC0EFF45EB1F" ID-GBIF-Occurrence="1700783859" collectingDate="2015-05-07" collectionCode="CSIRO" country="Solomon Islands" location="Plaza" pageId="2" pageNumber="535" specimenCount="1" stateProvince="Honiara">
|
||
<emphasis id="E5DBEAAEB361FF92FF18CC0EFE12EB24" bold="true" box="[199,385,1586,1611]" pageId="2" pageNumber="535">Other material.</emphasis>
|
||
<taxonomicName id="10AF4D3FB361FF92FE58CC08FE49EB23" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[391,474,1588,1612]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="559" phylum="Arthropoda" rank="genus">
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||
<collectionCode id="B1BEAE79B361FF92FE58CC08FE49EB23" box="[391,474,1588,1612]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="2" pageNumber="535">CSIRO</collectionCode>
|
||
</taxonomicName>
|
||
H 7723-01, female
|
||
<quantity id="10579B59B361FF92FD66CC0FFC86EB24" box="[697,789,1587,1612]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" pageId="2" pageNumber="535" unit="mm" value="295.0">295 mm</quantity>
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||
DW,
|
||
<location id="D2706067B361FF92FC8ACC0FFC01EB23" LSID="urn:lsid:plazi:treatment:5F0687AAB361FF98FF48CEC9FA1DEC2E:D2706067B361FF92FC8ACC0FFC01EB23" box="[853,914,1587,1612]" country="Solomon Islands" name="Plaza" pageId="2" pageNumber="535" stateProvince="Honiara">Plaza</location>
|
||
fish market,
|
||
<collectingRegion id="156BF85EB361FF92FBFBCC0FFB12EB23" box="[1060,1153,1587,1612]" country="Solomon Islands" name="Honiara" pageId="2" pageNumber="535">Honiara</collectingRegion>
|
||
,
|
||
<collectingCountry id="AFB8762CB361FF92FB53CC0FFADAEB23" box="[1164,1353,1587,1612]" name="Solomon Islands" pageId="2" pageNumber="535">Solomon Islands</collectingCountry>
|
||
,
|
||
<collectingDate id="B355E994B361FF92FA8BCC08FF40EB1F" pageId="2" pageNumber="535" value="2015-05-07">
|
||
<date id="A311107CB361FF92FA8BCC08FF40EB1F" pageId="2" pageNumber="535" value="2015-05-07">7 May 2015</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FB56537B361FF93FF18CC46FDBDED97" lastPageId="3" lastPageNumber="536" pageId="2" pageNumber="535" type="diagnosis">
|
||
<paragraph id="D71036BCB361FF93FF18CC46FDBDED97" blockId="2.[151,1437,1443,2008]" lastBlockId="3.[151,1437,151,1977]" lastPageId="3" lastPageNumber="536" pageId="2" pageNumber="535">
|
||
<emphasis id="E5DBEAAEB361FF92FF18CC46FED2EBFC" bold="true" box="[199,321,1658,1683]" pageId="2" pageNumber="535">Diagnosis.</emphasis>
|
||
A medium-sized
|
||
<taxonomicName id="10AF4D3FB361FF92FDD7CC40FDEDEBFC" authorityName="Castelnau" authorityYear="1873" box="[520,638,1660,1683]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="2" pageNumber="535" phylum="Chordata" rank="genus">
|
||
<emphasis id="E5DBEAAEB361FF92FDD7CC40FDEDEBFC" box="[520,638,1660,1683]" italics="true" pageId="2" pageNumber="535">Neotrygon</emphasis>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="10AF4D3FB361FF92FD13CC47FC9FEBFB" authorityName="in Muller & Henle" authorityYear="1841" box="[716,780,1659,1684]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="2" pageNumber="535" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB361FF92FD13CC47FC9FEBFB" box="[716,780,1659,1684]" italics="true" pageId="2" pageNumber="535">kuhlii</emphasis>
|
||
</taxonomicName>
|
||
-complex (reaching at least
|
||
<quantity id="10579B59B361FF92FB9CCC40FB1BEBFC" box="[1091,1160,1660,1684]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="535" unit="cm" value="30.0">30 cm</quantity>
|
||
DW) with the following combination of characters: disc broader than long, width ~1.2 times length; pectoral apices narrowly rounded; snout rather fleshy, broadly angular, angle ~107°, length 1.7–2.1 times interorbital width; maximum width relatively well back on disc, length from snout tip to pectoral-fin insertion 1.8–1.9 times and disc width 2.5–2.6 times horizontal distance from snout tip to maximum disc width; preoral length 2.4–2.8 times mouth width; internasal distance
|
||
<quantity id="10579B59B361FF92FEAFCD13FE71EA27" box="[368,482,1839,1864]" metricMagnitude="-2" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="2" pageNumber="535" unit="in" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 in</quantity>
|
||
prenasal length; interspiracular distance 13–15% DW; nostril length 2.8–4.1% DW; nasal curtain width 8–8.3% DW; small mouth, width 6.4–6.8% DW; horizontal distance from cloaca to caudal sting base ~55% of disc length; thornlets present in nuchal region, absent from tail in all sizes; dermal denticles absent; pectoral-fin radials 113 (based on new specimen); total vertebral centra (including synarcual) 133, trunk centra (including synarcual) 39; blue spots very small and sparse, largest spot on disc 0.3–0.5 times eye width; 0–6 (mean 3.0) blue spots on medial belt, largest ~2.1% DW; mask-like marking pronounced, not covered with dark peppery spots; ventral surface of disc and pelvic fins with broad dark greyish submarginal bands; ventral tail fold and adjacent tail bluish grey when fresh.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FB56537B360FF93FF18CB3FFBCAEC2F" pageId="3" pageNumber="536" type="distribution">
|
||
<paragraph id="D71036BCB360FF93FF18CB3FFBCAEC2F" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="536">
|
||
<emphasis id="E5DBEAAEB360FF93FF18CB3FFEF3EC73" bold="true" box="[199,352,259,284]" pageId="3" pageNumber="536">Distribution.</emphasis>
|
||
Tropical South-West Pacific, off the
|
||
<collectingCountry id="AFB8762CB360FF93FCC3CB38FC4CEC72" box="[796,991,260,285]" name="Solomon Islands" pageId="3" pageNumber="536">Solomon Islands</collectingCountry>
|
||
; Santa Cruz Islands and
|
||
<collectingRegion id="156BF85EB360FF93FAD6CB38FA0AEC72" box="[1289,1433,260,285]" country="Solomon Islands" name="Guadalcanal" pageId="3" pageNumber="536">Guadalcanal</collectingRegion>
|
||
. Probably more widespread in Oceania but limits of its distribution need to be defined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FB56537B360FF98FF18CB70FA1DEC2E" lastPageId="8" lastPageNumber="541" pageId="3" pageNumber="536" type="discussion">
|
||
<paragraph id="D71036BCB360FF93FF18CB70FC8CEFC7" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="536">
|
||
<emphasis id="E5DBEAAEB360FF93FF18CB70FEA8EC0B" bold="true" box="[199,315,332,356]" pageId="3" pageNumber="536">Remarks.</emphasis>
|
||
The two former
|
||
<typeStatus id="0814881EB360FF93FE22CB71FDCCEC0A" box="[509,607,333,357]" pageId="3" pageNumber="536" type="syntype">syntypes</typeStatus>
|
||
(MNHN 2440, 156 and
|
||
<quantity id="10579B59B360FF93FCA6CB70FC44EC0B" box="[889,983,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.59" pageId="3" pageNumber="536" unit="mm" value="159.0">159 mm</quantity>
|
||
DW) from the eastern
|
||
<collectingCountry id="AFB8762CB360FF93FB3FCB70FA0FEC0A" box="[1248,1436,332,357]" name="Solomon Islands" pageId="3" pageNumber="536">Solomon Islands</collectingCountry>
|
||
(Vanikoro) are both females, presumably immature. The designated
|
||
<typeStatus id="0814881EB360FF93FC79CB53FB9EECE7" box="[934,1037,367,392]" pageId="3" pageNumber="536" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<quantity id="10579B59B360FF93FBC0CB53FBECECE8" box="[1055,1151,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="1.56" pageId="3" pageNumber="536" unit="mm" value="156.0">156 mm</quantity>
|
||
DW) has strongly faded skin, almost pale brown with evidence of very small, white ocelli on both (confined to two spots on right side), with the tail intact; white central core of ocellus
|
||
<quantity id="10579B59B360FF93FD6ECB84FCABECA0" box="[689,824,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="3" pageNumber="536" unit="mm" value="1.45" valueMax="1.7" valueMin="1.2">1.2–1.7 mm</quantity>
|
||
diameter, surrounded by thin darker ring ~
|
||
<quantity id="10579B59B360FF93FAC9CB8BFAFCECA0" box="[1302,1391,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="536" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; the tail is mainly pale with two dark saddles confined to the dorsal part after caudal sting; entire tail tip is brownish on both surfaces. The designated
|
||
<typeStatus id="0814881EB360FF93FE2ECBC3FD1BEF77" box="[497,648,511,536]" pageId="3" pageNumber="536" type="paralectotype">paralectotype</typeStatus>
|
||
(
|
||
<quantity id="10579B59B360FF93FD47CBC3FD6BEF78" box="[664,760,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="1.59" pageId="3" pageNumber="536" unit="mm" value="159.0">159 mm</quantity>
|
||
DW) has ~29 similar-sized ocelli (some possibly missing), still evident beside orbits and irregularly dispersed over disc (diameter white core
|
||
<quantity id="10579B59B360FF93FB81C818FB4BEF53" box="[1118,1240,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.6" metricValueMin="1.0" pageId="3" pageNumber="536" unit="mm" value="1.8" valueMax="2.6" valueMin="1.0">1–2.6 mm</quantity>
|
||
); additional dark speckles are present on the orbital membrane, interorbit and over the central disc (<
|
||
<quantity id="10579B59B360FF93FBB6C87BFB50EF30" box="[1129,1219,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="536" unit="mm" value="1.5">1.5 mm</quantity>
|
||
diameter). In both specimens, the posterior margins of ventral disc are probably darker than centrally; outer edge of ventral fold darker than its base (discussed by
|
||
<bibRefCitation id="B33E4B4DB360FF93FDCAC8B3FC81EFC7" author="Muller" box="[533,786,655,680]" pageId="3" pageNumber="536" refString="Muller, J., & Henle, F. G. J. (1841) Systematische Beschreibung der Plagiostomen. Berlin, pp. 103 - 200." type="book chapter" year="1841">Müller & Henle, 1841</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D71036BCB360FF93FF18C888FE57EEEF" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="536">
|
||
A third much smaller
|
||
<typeStatus id="0814881EB360FF93FE64C889FD81EFA2" box="[443,530,693,717]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
(MNHN A7931) collected from New
|
||
<collectingCountry id="AFB8762CB360FF93FC60C888FB83EFA2" box="[959,1040,692,717]" name="Guinea-Bissau" pageId="3" pageNumber="536">Guinea</collectingCountry>
|
||
by Quoy & Gaimard, is a late-term male embryo (~
|
||
<quantity id="10579B59B360FF93FE98C8E4FE35EF80" box="[327,422,728,752]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="536" unit="mm" value="100.0">100 mm</quantity>
|
||
DW) with a rudiment of its umbilical cord still evident. The specimen is heavily bleached, being uniformly pale with no indication of colour (
|
||
<figureCitation id="4F942A39B360FF93FD29C8C0FCDFEE7B" box="[758,844,764,789]" captionStart="FIGURE 3" captionStartId="5.[151,250,1925,1947]" captionTargetBox="[189,1389,218,1887]" captionTargetId="figure@5.[151,1436,171,1910]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of: (A) embryonic syntype of Neotrygon kuhlii from West Papua, New Guinea (MNHN A 7931, embryo 99.7 mm DW; probably equals Neotrygon australiae); (B) dried probable syntype of Neotrygon kuhlii from Java, Indonesia (RMNH 2472, adult male ~ 271 mm DW; probably equals Neotrygon caeruleopunctata)." httpUri="https://zenodo.org/record/1051778/files/figure.png" pageId="3" pageNumber="536">Fig. 3a</figureCitation>
|
||
). Morphometrics of these preserved
|
||
<typeStatus id="0814881EB360FF93FB20C8C1FAA9EE7A" box="[1279,1338,765,789]" pageId="3" pageNumber="536">types</typeStatus>
|
||
may be slightly affected by preservation, as the
|
||
<typeStatus id="0814881EB360FF93FD81C923FD57EE57" box="[606,708,799,824]" pageId="3" pageNumber="536" type="lectotype">lectotype</typeStatus>
|
||
in particular has very flexible skin that may alter some anterior measurements. The fourth
|
||
<typeStatus id="0814881EB360FF93FE1AC979FD8FEE32" box="[453,540,837,861]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
from Leiden is probably RMNH 2472, an adult male (~
|
||
<quantity id="10579B59B360FF93FB49C978FB65EE33" box="[1174,1270,836,860]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="536" unit="mm" value="270.0">270 mm</quantity>
|
||
DW) which is dried and stuffed (
|
||
<figureCitation id="4F942A39B360FF93FEBCC95BFE24EEEF" box="[355,439,871,896]" captionStart="FIGURE 3" captionStartId="5.[151,250,1925,1947]" captionTargetBox="[189,1389,218,1887]" captionTargetId="figure@5.[151,1436,171,1910]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of: (A) embryonic syntype of Neotrygon kuhlii from West Papua, New Guinea (MNHN A 7931, embryo 99.7 mm DW; probably equals Neotrygon australiae); (B) dried probable syntype of Neotrygon kuhlii from Java, Indonesia (RMNH 2472, adult male ~ 271 mm DW; probably equals Neotrygon caeruleopunctata)." httpUri="https://zenodo.org/record/1051778/files/figure.png" pageId="3" pageNumber="536">Fig. 3b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D71036BCB360FF93FF18C9B0FC44EB1A" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="536">
|
||
Identification of the primary
|
||
<typeStatus id="0814881EB360FF93FDCCC9B1FDD0EECA" box="[531,579,909,933]" pageId="3" pageNumber="536">type</typeStatus>
|
||
specimen used in the Müller & Henle’s description and excellent figure to designate a
|
||
<typeStatus id="0814881EB360FF93FEFFC993FE14EEA7" box="[288,391,943,968]" pageId="3" pageNumber="536" type="lectotype">lectotype</typeStatus>
|
||
of the species has proven problematic. The number of blue spots and the presence of two caudal stings in their figure does not coincide with any of the
|
||
<typeStatus id="0814881EB360FF93FC98C9E9FC16EE82" box="[839,901,981,1005]" pageId="3" pageNumber="536">types</typeStatus>
|
||
. Also, the illustration is of a female whereas the probable Leiden
|
||
<typeStatus id="0814881EB360FF93FE85C9C4FE22E97F" box="[346,433,1016,1040]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
is an adult male. Either their illustration is a composite image (most likely) or the dry Leiden adult male is not a
|
||
<typeStatus id="0814881EB360FF93FE1FCE21FD88E95A" box="[448,539,1053,1077]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
. The Leiden
|
||
<typeStatus id="0814881EB360FF93FD73CE21FD4FE95A" box="[684,732,1053,1077]" pageId="3" pageNumber="536">type</typeStatus>
|
||
(RMNH 2472) is listed as from
|
||
<collectingRegion id="156BF85EB360FF93FB96CE21FBEEE95B" box="[1097,1149,1053,1076]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="536">Java</collectingRegion>
|
||
, collected by Kuhl & van Hasselt. Müller & Henle’s description, the
|
||
<typeStatus id="0814881EB360FF93FDA2CE7CFD3EE937" box="[637,685,1088,1112]" pageId="3" pageNumber="536">type</typeStatus>
|
||
location is listed as ‘Indien’ which although often presumed to be
|
||
<collectingCountry id="AFB8762CB360FF93FF48CE58FF40E912" box="[151,211,1124,1149]" name="India" pageId="3" pageNumber="536">India</collectingCountry>
|
||
, was often used for East
|
||
<collectingCountry id="AFB8762CB360FF93FE33CE58FDB5E912" box="[492,550,1124,1149]" name="India" pageId="3" pageNumber="536">India</collectingCountry>
|
||
or East Indies, present day
|
||
<collectingCountry id="AFB8762CB360FF93FC83CE58FC5EE912" box="[860,973,1124,1149]" name="Indonesia" pageId="3" pageNumber="536">Indonesia</collectingCountry>
|
||
. Thus, this provides some support for the Javan specimen as the fourth
|
||
<typeStatus id="0814881EB360FF93FE01CEB5FDA6E9CE" box="[478,565,1161,1185]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="10AF4D3FB360FF93FD86CEB5FD20E9CF" authorityName="in Muller & Henle" authorityYear="1841" box="[601,691,1159,1184]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB360FF93FD86CEB5FD20E9CF" box="[601,691,1159,1184]" italics="true" pageId="3" pageNumber="536">T. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
. But it should be noted that in other species descriptions, Müller & Henle refer to
|
||
<collectingRegion id="156BF85EB360FF93FE9BCE91FEE5E9AB" box="[324,374,1197,1220]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="536">Java</collectingRegion>
|
||
in the locations provided so there is still an element of uncertainty over the exact
|
||
<typeStatus id="0814881EB360FF93FA9ACE91FA0FE9AA" box="[1349,1436,1197,1221]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
specimen. Müller & Henle’s description also refers to a sting (rather than two): ‘der Stachel am Ende des vordem Viertels’. Still, a discrepancy remains as to their sizes; the
|
||
<typeStatus id="0814881EB360FF93FCE9CEC9FC1EE862" box="[822,909,1269,1293]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
measured appears to be about
|
||
<quantity id="10579B59B360FF93FB2DCEC8FAF6E863" box="[1266,1381,1268,1293]" metricMagnitude="-1" metricUnit="m" metricValue="1.395" pageId="3" pageNumber="536" unit="mm" value="139.5">139.5 mm</quantity>
|
||
DW (largest width '5 Zoll, 4 Linie' based on Müller & Henle) which is much smaller than the first size of maturity of any member of the
|
||
<taxonomicName id="10AF4D3FB360FF93FEA3CF00FE2FE83A" authorityName="in Muller & Henle" authorityYear="1841" box="[380,444,1340,1365]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB360FF93FEA3CF00FE2FE83A" box="[380,444,1340,1365]" italics="true" pageId="3" pageNumber="536">kuhlii</emphasis>
|
||
</taxonomicName>
|
||
-complex. The paucity and very small sizes of blue spots appears typical of maskrays from the
|
||
<collectingCountry id="AFB8762CB360FF93FEDFCF63FE53E817" box="[256,448,1375,1400]" name="Solomon Islands" pageId="3" pageNumber="536">Solomon Islands</collectingCountry>
|
||
; blue-spotted maskray populations from
|
||
<collectingCountry id="AFB8762CB360FF93FC50CF63FC5AE817" box="[911,969,1375,1400]" name="India" pageId="3" pageNumber="536">India</collectingCountry>
|
||
(inferentially specified as the location of the fourth
|
||
<typeStatus id="0814881EB360FF93FECDCFB9FEFAE8F2" box="[274,361,1413,1437]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
by Eschmeyer, 2016) and
|
||
<collectingCountry id="AFB8762CB360FF93FD7BCFB8FC82E8F2" box="[676,785,1412,1437]" name="Indonesia" pageId="3" pageNumber="536">Indonesia</collectingCountry>
|
||
(often referred to as ‘Indien’ in old museum labels and incorrectly accorded to
|
||
<collectingCountry id="AFB8762CB360FF93FE7ECF9BFE4FE8AF" box="[417,476,1447,1472]" name="India" pageId="3" pageNumber="536">India</collectingCountry>
|
||
) have medium to large blue spots, usually in much higher densities. The size of spots and their distribution in Müller & Henle’s figure (
|
||
<figureCitation id="4F942A39B360FF93FD16CFF0FC99E88B" box="[713,778,1484,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1167,1189]" captionText="FIGURE 1. Original illustration of Trygon kuhlii in Müller & Henle (1841)." pageId="3" pageNumber="536">Fig. 1</figureCitation>
|
||
) is very similar to that of the two Vanikoro
|
||
<typeStatus id="0814881EB360FF93FB21CFF1FAAAE88A" box="[1278,1337,1485,1509]" pageId="3" pageNumber="536">types</typeStatus>
|
||
(
|
||
<figureCitation id="4F942A39B360FF93FA96CFF0FA1CE88B" box="[1353,1423,1484,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[349,1334,217,1920]" captionTargetId="figure@4.[151,1436,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal view of Neotrygon kuhlii types from Vanikoro, Solomon Islands (preserved): (A) lectotype (MNHN A 2440, 1 of 2, female 154 mm DW); (B) paralectotype (MNHN A 2440, 1 of 2, female 162.7 mm DW)." httpUri="https://zenodo.org/record/1051776/files/figure.png" pageId="3" pageNumber="536">Fig. 2</figureCitation>
|
||
). On that basis, the most complete MNHN specimen, a
|
||
<quantity id="10579B59B360FF93FD22CFD3FCC8EB67" box="[765,859,1519,1544]" metricMagnitude="-1" metricUnit="m" metricValue="1.56" pageId="3" pageNumber="536" unit="mm" value="156.0">156 mm</quantity>
|
||
DW female in good condition with tail and caudal sting intact, is hereby designated as the
|
||
<typeStatus id="0814881EB360FF93FDA4CC28FD75EB42" box="[635,742,1556,1581]" pageId="3" pageNumber="536" type="lectotype">lectotype</typeStatus>
|
||
. The second Vanikoro
|
||
<typeStatus id="0814881EB360FF93FC24CC29FBB8EB42" box="[1019,1067,1557,1581]" pageId="3" pageNumber="536">type</typeStatus>
|
||
becomes a
|
||
<typeStatus id="0814881EB360FF93FB1FCC28FAC8EB42" box="[1216,1371,1556,1581]" pageId="3" pageNumber="536" type="paralectotype">paralectotype</typeStatus>
|
||
. This embryonic
|
||
<typeStatus id="0814881EB360FF93FECACC05FEFFEB3E" box="[277,364,1593,1617]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
(MNHN A7931) probably equals
|
||
<taxonomicName id="10AF4D3FB360FF93FD33CC05FCECEB3F" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[748,895,1591,1616]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="australiae" status="sp. nov.">
|
||
<emphasis id="E5DBEAAEB360FF93FD33CC05FCECEB3F" box="[748,895,1591,1616]" italics="true" pageId="3" pageNumber="536">N. australiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FEE857D5B360FF93FC5ACC05FC4DEB3F" box="[901,990,1593,1616]" pageId="3" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and the dried RMNH
|
||
<typeStatus id="0814881EB360FF93FB04CC05FAA1EB3E" box="[1243,1330,1593,1617]" pageId="3" pageNumber="536" type="syntype">syntype</typeStatus>
|
||
probably equals
|
||
<taxonomicName id="10AF4D3FB360FF93FF3ACC61FE59EB1A" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[229,458,1628,1653]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="caeruleopunctata" status="sp. nov.">
|
||
<emphasis id="E5DBEAAEB360FF93FF3ACC61FE59EB1A" box="[229,458,1628,1653]" italics="true" pageId="3" pageNumber="536">N. caeruleopunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FEE857D5B360FF93FE0ECC61FDBBEB1B" box="[465,552,1629,1652]" pageId="3" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
|
||
based on its size and capture location.
|
||
</paragraph>
|
||
<paragraph id="D71036BCB360FF93FF18CC43FE27EAD7" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="536">
|
||
The meristic details of the
|
||
<taxonomicName id="10AF4D3FB360FF93FE24CCBDFDCEEBF7" authorityName="in Muller & Henle" authorityYear="1841" box="[507,605,1663,1688]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB360FF93FE24CCBDFDCEEBF7" box="[507,605,1663,1688]" italics="true" pageId="3" pageNumber="536">N. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="0814881EB360FF93FDBACCBDFD33EBF6" box="[613,672,1665,1689]" pageId="3" pageNumber="536">types</typeStatus>
|
||
were not obtained so counts are based solely on a newly collected specimen (
|
||
<taxonomicName id="10AF4D3FB360FF93FED0CC98FEF0EBD3" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[271,355,1700,1724]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
H 7723-01): total pectoral-fin radials 113; propterygium 45, mesopterygium 19, metapterygium 49. Pelvic-fin radials: 1 + 25–26. Vertebral centra total (including synarcual) 133; total (excluding synarcual) 128; monospondylous (including synarcual) 44; monospondylous (excluding synarcual) 39; pre-sting diplospondylous 64; post-sting diplospondylous 25. Count of the pectoral-fin radials (i.e. 113) is high for a
|
||
<taxonomicName id="10AF4D3FB360FF93FB78CD33FB74EA47" authorityName="in Muller & Henle" authorityYear="1841" box="[1191,1255,1807,1832]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB360FF93FB78CD33FB74EA47" box="[1191,1255,1807,1832]" italics="true" pageId="3" pageNumber="536">kuhlii</emphasis>
|
||
</taxonomicName>
|
||
-group member. This specimen resembles the Vanikoro
|
||
<typeStatus id="0814881EB360FF93FD92CD08FD27EA22" box="[589,692,1844,1869]" pageId="3" pageNumber="536" type="lectotype">lectotype</typeStatus>
|
||
in shape and colour. Its COI sequence indicates that this species is distinct from all other
|
||
<taxonomicName id="10AF4D3FB360FF93FE43CD6BFE4FEA1F" authorityName="in Muller & Henle" authorityYear="1841" box="[412,476,1879,1904]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB360FF93FE43CD6BFE4FEA1F" box="[412,476,1879,1904]" italics="true" pageId="3" pageNumber="536">kuhlii</emphasis>
|
||
</taxonomicName>
|
||
-group members (Appendix 1). In a similar analysis for the chondrichthyan Tree of Life project using the NADH2 gene, the specimen was basal and separate from all other forms of the
|
||
<taxonomicName id="10AF4D3FB360FF93FACECD40FAC2EAFA" authorityName="in Muller & Henle" authorityYear="1841" box="[1297,1361,1916,1941]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="536" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB360FF93FACECD40FAC2EAFA" box="[1297,1361,1916,1941]" italics="true" pageId="3" pageNumber="536">kuhlii</emphasis>
|
||
</taxonomicName>
|
||
-group (G. Naylor, pers. comm.).
|
||
</paragraph>
|
||
<caption id="83D06634B367FF94FF48CD9DFB34EAB9" httpUri="https://zenodo.org/record/1051776/files/figure.png" pageId="4" pageNumber="537" targetBox="[349,1334,217,1920]" targetPageId="4">
|
||
<paragraph id="D71036BCB367FF94FF48CD9DFB34EAB9" blockId="4.[151,1436,1953,2006]" pageId="4" pageNumber="537">
|
||
<emphasis id="E5DBEAAEB367FF94FF48CD9DFE81EAD9" bold="true" box="[151,274,1953,1975]" pageId="4" pageNumber="537">FIGURE 2.</emphasis>
|
||
Dorsal view of
|
||
<taxonomicName id="10AF4D3FB367FF94FE1DCD9EFDE3EAD8" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[450,624,1953,1975]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="4" pageNumber="537" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB367FF94FE1DCD9EFDE3EAD8" box="[450,624,1953,1975]" italics="true" pageId="4" pageNumber="537">Neotrygon kuhlii</emphasis>
|
||
</taxonomicName>
|
||
types from Vanikoro, Solomon Islands (preserved): (A) lectotype (MNHN A 2440, 1 of 2, female 154 mm DW); (B) paralectotype (MNHN A 2440, 1 of 2, female 162.7 mm DW).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="83D06634B366FF95FF48CDB9FB13EAB6" httpUri="https://zenodo.org/record/1051778/files/figure.png" pageId="5" pageNumber="538" targetBox="[189,1389,218,1887]" targetPageId="5">
|
||
<paragraph id="D71036BCB366FF95FF48CDB9FB13EAB6" blockId="5.[151,1436,1925,2009]" pageId="5" pageNumber="538">
|
||
<emphasis id="E5DBEAAEB366FF95FF48CDB9FE81EAF5" bold="true" box="[151,274,1925,1947]" pageId="5" pageNumber="538">FIGURE 3.</emphasis>
|
||
Dorsal view of: (A) embryonic syntype of
|
||
<taxonomicName id="10AF4D3FB366FF95FD39CDBAFC06EAF4" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[742,917,1925,1947]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="5" pageNumber="538" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB366FF95FD39CDBAFC06EAF4" box="[742,917,1925,1947]" italics="true" pageId="5" pageNumber="538">Neotrygon kuhlii</emphasis>
|
||
</taxonomicName>
|
||
from West Papua, New Guinea (MNHN A7931, embryo 99.7 mm DW; probably equals
|
||
<taxonomicName id="10AF4D3FB366FF95FDE2CD99FC8AEAD5" authorityName="Last & White & Séret" authorityYear="2016" box="[573,793,1956,1978]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="5" pageNumber="538" phylum="Chordata" rank="species" species="australiae">
|
||
<emphasis id="E5DBEAAEB366FF95FDE2CD99FC8AEAD5" box="[573,793,1956,1978]" italics="true" pageId="5" pageNumber="538">Neotrygon australiae</emphasis>
|
||
</taxonomicName>
|
||
); (B) dried probable syntype of
|
||
<taxonomicName id="10AF4D3FB366FF95FBAACD99FAB7EAD5" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[1141,1316,1956,1978]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="5" pageNumber="538" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB366FF95FBAACD99FAB7EAD5" box="[1141,1316,1956,1978]" italics="true" pageId="5" pageNumber="538">Neotrygon kuhlii</emphasis>
|
||
</taxonomicName>
|
||
from Java, Indonesia (RMNH 2472, adult male ~271 mm DW; probably equals
|
||
<taxonomicName id="10AF4D3FB366FF95FC90CDF8FBE2EAB6" authorityName="Last & White & Séret" authorityYear="2016" box="[847,1137,1987,2009]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="5" pageNumber="538" phylum="Chordata" rank="species" species="caeruleopunctata">
|
||
<emphasis id="E5DBEAAEB366FF95FC90CDF8FBE2EAB6" box="[847,1137,1987,2009]" italics="true" pageId="5" pageNumber="538">Neotrygon caeruleopunctata</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="83D06634B365FF96FF48CD9DFDD0EAB9" httpUri="https://zenodo.org/record/1051780/files/figure.png" pageId="6" pageNumber="539" targetBox="[176,1394,212,1911]" targetPageId="6">
|
||
<paragraph id="D71036BCB365FF96FF48CD9DFDD0EAB9" blockId="6.[151,1436,1953,2006]" pageId="6" pageNumber="539">
|
||
<emphasis id="E5DBEAAEB365FF96FF48CD9DFE81EAD9" bold="true" box="[151,274,1953,1975]" pageId="6" pageNumber="539">FIGURE 4.</emphasis>
|
||
Freshly caught specimen of
|
||
<taxonomicName id="10AF4D3FB365FF96FD9BCD9EFD60EAD8" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[580,755,1953,1975]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="6" pageNumber="539" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB365FF96FD9BCD9EFD60EAD8" box="[580,755,1953,1975]" italics="true" pageId="6" pageNumber="539">Neotrygon kuhlii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="10AF4D3FB365FF96FCDDCD9DFCDDEAD8" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[770,846,1953,1975]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
7723-01, female 295 mm DW) from Honiara, Solomon Islands: (A) dorsal view; (B) ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="83D06634B364FF97FF48CE79FB95E9F7" httpUri="https://zenodo.org/record/1051782/files/figure.png" pageId="7" pageNumber="540" targetBox="[151,1436,198,1064]" targetPageId="7">
|
||
<paragraph id="D71036BCB364FF97FF48CE79FB95E9F7" blockId="7.[151,1436,1092,1176]" pageId="7" pageNumber="540">
|
||
<emphasis id="E5DBEAAEB364FF97FF48CE79FE81E935" bold="true" box="[151,274,1093,1115]" pageId="7" pageNumber="540">FIGURE 5.</emphasis>
|
||
Oronasal region of: (A)
|
||
<taxonomicName id="10AF4D3FB364FF97FDCBCE7AFD51E935" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[532,706,1092,1115]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="E5DBEAAEB364FF97FDCBCE7AFD51E935" box="[532,706,1092,1115]" italics="true" pageId="7" pageNumber="540">Neotrygon kuhlii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="10AF4D3FB364FF97FD0FCE78FC8FE935" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[720,796,1092,1114]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
7723-01, female 295 mm DW); (B)
|
||
<taxonomicName id="10AF4D3FB364FF97FB42CE7AFAE4E935" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[1181,1399,1092,1115]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="australiae" status="sp. nov.">
|
||
<emphasis id="E5DBEAAEB364FF97FB42CE7AFAE4E935" box="[1181,1399,1092,1115]" italics="true" pageId="7" pageNumber="540">Neotrygon australiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FEE857D5B364FF97FAA0CE79FF51E916" pageId="7" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<taxonomicName id="10AF4D3FB364FF97FF11CE5FFE89E916" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[206,282,1123,1145]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
CA 3247, adult male 379 mm DW); (C)
|
||
<taxonomicName id="10AF4D3FB364FF97FD66CE58FC48E916" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[697,987,1123,1145]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="caeruleopunctata" status="sp. nov.">
|
||
<emphasis id="E5DBEAAEB364FF97FD66CE58FC48E916" box="[697,987,1123,1145]" italics="true" pageId="7" pageNumber="540">Neotrygon caeruleopunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FEE857D5B364FF97FC3ECE58FBA3E916" box="[993,1072,1124,1145]" pageId="7" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<taxonomicName id="10AF4D3FB364FF97FBE3CE5FFB1BE916" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[1084,1160,1123,1145]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
H 6202-03, adult male 311 mm DW); (D)
|
||
<taxonomicName id="10AF4D3FB364FF97FEF1CEBFFE68E9F7" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[302,507,1154,1176]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="orientale" status="sp. nov.">
|
||
<emphasis id="E5DBEAAEB364FF97FEF1CEBFFE68E9F7" box="[302,507,1154,1176]" italics="true" pageId="7" pageNumber="540">Neotrygon orientale</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FEE857D5B364FF97FDDECEBFFDDCE9F7" box="[513,591,1155,1176]" pageId="7" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<taxonomicName id="10AF4D3FB364FF97FD84CEBEFD34E9F7" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[603,679,1154,1176]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
H 6130-01, female 254 mm DW).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="83D06634B364FF97FF48CEF8FA1FE84D" pageId="7" pageNumber="540">
|
||
<paragraph id="D71036BCB364FF97FF48CEF8FA1FE84D" blockId="7.[151,1436,1220,1314]" pageId="7" pageNumber="540">
|
||
<emphasis id="E5DBEAAEB364FF97FF48CEF8FE99E9B4" bold="true" box="[151,266,1220,1243]" pageId="7" pageNumber="540">TABLE 1.</emphasis>
|
||
Morphometric data for the lectotype (MNHN 2440, 1 of 2), paralectotype (MNHN 2440, 1 of 2), and one other specimen (
|
||
<taxonomicName id="10AF4D3FB364FF97FE97CEDBFE0BE991" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[328,408,1255,1278]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
H 7723-01) of
|
||
<emphasis id="E5DBEAAEB364FF97FD9FCEDBFD68E991" box="[576,763,1254,1278]" italics="true" pageId="7" pageNumber="540">
|
||
<taxonomicName id="10AF4D3FB364FF97FD9FCEDBFD64E991" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[576,759,1254,1278]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="kuhlii">Neotrygon kuhlii</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and for the holotype of
|
||
<taxonomicName id="10AF4D3FB364FF97FBDFCEDBFB77E991" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[1024,1252,1254,1278]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="australiae" status="sp. nov.">
|
||
<emphasis id="E5DBEAAEB364FF97FBDFCEDBFB77E991" box="[1024,1252,1254,1278]" italics="true" pageId="7" pageNumber="540">Neotrygon australiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FEE857D5B364FF97FB34CEDBFAADE991" box="[1259,1342,1255,1278]" pageId="7" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<taxonomicName id="10AF4D3FB364FF97FA93CEDBFA0FE991" authorityName="Medvedev & Lawrence" authorityYear="1984" box="[1356,1436,1255,1278]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="559" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
|
||
H 7016-01), and ranges and means for the measured paratypes. Measurements expressed as a percentage of disc width.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D71036BCB364FF97FD09CF78FAB0E836" blockId="7.[165,1422,1347,2030]" box="[726,1315,1347,1370]" pageId="7" pageNumber="540">
|
||
<emphasis id="E5DBEAAEB364FF97FD09CF78FAB0E836" bold="true" box="[726,1315,1347,1370]" pageId="7" pageNumber="540">
|
||
<emphasis id="E5DBEAAEB364FF97FD09CF78FB59E835" bold="true" box="[726,1226,1347,1370]" italics="true" pageId="7" pageNumber="540">
|
||
<taxonomicName id="10AF4D3FB364FF97FD09CF78FCA0E835" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[726,819,1347,1370]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="kuhlii">N. kuhlii</taxonomicName>
|
||
<taxonomicName id="10AF4D3FB364FF97FB9CCF78FB59E835" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[1091,1226,1347,1370]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="540" phylum="Chordata" rank="species" species="australiae" status="sp. nov.">N. australiae</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="FEE857D5B364FF97FB0FCF78FAB0E836" box="[1232,1315,1348,1369]" pageId="7" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D71036BCB364FF97FD91CF4CFAB5E8EA" blockId="7.[165,1422,1347,2030]" box="[590,1318,1391,1413]" pageId="7" pageNumber="540">
|
||
<typeStatus id="0814881EB364FF97FD91CF4CFD20E8EA" box="[590,691,1392,1413]" pageId="7" pageNumber="540" type="lectotype">Lectotype</typeStatus>
|
||
Paralect. Other
|
||
<typeStatus id="0814881EB364FF97FC07CF53FBA5E8EA" box="[984,1078,1391,1413]" pageId="7" pageNumber="540" type="holotype">Holotype</typeStatus>
|
||
<typeStatus id="0814881EB364FF97FB1CCF4CFAB5E8EA" box="[1219,1318,1392,1413]" pageId="7" pageNumber="540" type="paratype">Paratypes</typeStatus>
|
||
</paragraph>
|
||
<paragraph id="D71036BCB364FF97FCB6CFA0FA1DE8DD" blockId="7.[165,1422,1347,2030]" box="[873,1422,1436,1458]" pageId="7" pageNumber="540">material Min. Max. Mean</paragraph>
|
||
<paragraph id="D71036BCB364FF97FF7ACFFBFABCE8B2" blockId="7.[165,1422,1347,2030]" box="[165,1327,1479,1501]" pageId="7" pageNumber="540">Disc width (mm) 156 159 295 213 171 343</paragraph>
|
||
<paragraph id="D71036BCB364FF98FF7ACFC8FA1DEC2E" blockId="7.[165,1422,1347,2030]" lastBlockId="8.[165,1422,165,321]" lastPageId="8" lastPageNumber="541" pageId="7" pageNumber="540">
|
||
Total length 189.2 – 211.2 224.6 159.1 223.5 199.5 Disc length 82.2 81.3 81.7 82.6 79.8 85.1 82.4 Snout to pectoral-fin insertion 71.4 70.0 71.9 72.0 68.1 74.9 71.7 Disc thickness 12.0 12.3 11.1 12.6 11.8 14.3 12.8 Snout (preorbital) length 17.2 16.4 16.1 14.1 13.3 14.6 13.8 Snout (preorbital) horizontal length 14.4 13.6 13.6 12.0 10.5 11.3 11.0 Pelvic-fin (embedded) length 17.3 17.0 19.9 19.3 18.4 22.3 21.0 Width across pelvic-fin base 14.5 13.8 16.4 15.4 15.6 18.4 16.7 Greatest width across pelvic fins 31.7 34.0 39.7 41.2 29.6 40.1 35.1 Cloaca origin to tail tip 120.2 – 141.1 154.1 90.2 153.7 129.9 Tail width at axil of pelvic fins 7.2 7.4 7.6 8.3 7.3 9.0 8.1
|
||
<emphasis id="E5DBEAAEB364FF98FB88CDE4FB59ED86" italics="true" lastPageId="8" lastPageNumber="541" pageId="7" pageNumber="540">
|
||
......continued on the next page
|
||
<emphasis id="E5DBEAAEB36BFF98FD09CAEFFB59ED86" bold="true" box="[726,1226,210,233]" pageId="8" pageNumber="541">
|
||
<taxonomicName id="10AF4D3FB36BFF98FD09CAEFFCA0ED86" baseAuthorityName="Muller & Henle" baseAuthorityYear="1841" box="[726,819,210,233]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="541" phylum="Chordata" rank="species" species="kuhlii">N. kuhlii</taxonomicName>
|
||
<taxonomicName id="10AF4D3FB36BFF98FB9CCAEFFB59ED86" authority="Last & White & Séret, 2016" authorityName="Last & White & Séret" authorityYear="2016" box="[1091,1226,210,233]" class="Elasmobranchii" family="Dasyatidae" genus="Neotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="541" phylum="Chordata" rank="species" species="australiae" status="sp. nov.">N. australiae</taxonomicName>
|
||
</emphasis>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="FEE857D5B36BFF98FB0FCAEFFAB0ED87" box="[1232,1315,211,232]" pageId="8" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<typeStatus id="0814881EB36BFF98FD91CB3CFD20EC7A" box="[590,691,256,277]" pageId="8" pageNumber="541" type="lectotype">Lectotype</typeStatus>
|
||
Paralect. Other
|
||
<typeStatus id="0814881EB36BFF98FC07CAC3FBA5EC7A" box="[984,1078,255,277]" pageId="8" pageNumber="541" type="holotype">Holotype</typeStatus>
|
||
<typeStatus id="0814881EB36BFF98FB1CCB3CFAB5EC7A" box="[1219,1318,256,277]" pageId="8" pageNumber="541" type="paratype">Paratypes</typeStatus>
|
||
material Min. Max. Mean
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |