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<mods:title id="8295A0A6AB0FDEE9643C427D369CF2B2">Revision of Eucorydia Hebard, 1929 from China, with notes on the genus and species worldwide (Blattodea, Corydioidea, Corydiidae)</mods:title>
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<mods:namePart id="30304E27FA4D2D8F051AFF163F07050A">Qiu, Lu</mods:namePart>
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<treatment id="34C9447927A80DF58460E8DAEA6B10EC" ID-GBIF-Taxon="135450479" LSID="urn:lsid:plazi:treatment:34C9447927A80DF58460E8DAEA6B10EC" httpUri="http://treatment.plazi.org/id/34C9447927A80DF58460E8DAEA6B10EC" lastPageNumber="46" pageId="29" pageNumber="46">
<subSubSection id="656BCBD0C8FD34C0C92939C24F6E3820" pageId="29" pageNumber="46" type="nomenclature">
<paragraph id="450F14ADDB46419AC50734E85298C89A" pageId="29" pageNumber="46">
<taxonomicName id="8756278072AD10B3C993D276992B04C6" ID-CoL="YPTF" authority="Hanitsch, 1915" authorityName="Hanitsch" authorityYear="1915" class="Insecta" family="Corydiidae" genus="Eucorydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucorydia maxwelli" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="maxwelli">
<pageBreakToken id="61A709C5CD69DC5861A508142A216E62" pageId="29" pageNumber="46" start="start">Eucorydia</pageBreakToken>
maxwelli (Hanitsch, 1915)
</taxonomicName>
<taxonomicNameLabel id="E909DD3C2FAB14EC42AC7F14D956A121" pageId="29" pageNumber="46">stat. rev.</taxonomicNameLabel>
Fig. 13
<normalizedToken id="6975BA4CB4FB4808BAA58C29733E5DAE" originalValue="HK">H-K</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="F330A225D9F9946317E92127C036B319" pageId="29" pageNumber="46" type="reference_group">
<paragraph id="DB28287379B1925CE9A2663EB320B2BC" pageId="29" pageNumber="46">
<taxonomicName id="9C32A0EE88669D1EFB1A95DB92788D91" class="Insecta" family="Corydiidae" genus="Corydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corydia maxwelli" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="maxwelli">Corydia maxwelli</taxonomicName>
Hanitsch, 1915: 126 (2 males respectively from
<normalizedToken id="3A3FFA1BE58B5D8D59C103AFBB674B44" originalValue="Maxwells">Maxwell's</normalizedToken>
Hill, Perak and Lingga Mt., Sarawak; 2 females from Bukit Kutu, Selangor);
<bibRefCitation id="66920ABA47E085164A21208A8EC560C7" author="Hanitsch, R" journalOrPublisher="Journal Straits Branch Royal Asiatic Society" pageId="32" pageNumber="49" pagination="395 - 474" title="Malayan Blattidae. Part II." volume="1" year="1923">Hanitsch 1923</bibRefCitation>
: 466.
</paragraph>
<paragraph id="6E5A6A2A983B249B5717978A27A1BAA2" pageId="29" pageNumber="46">
<taxonomicName id="5592CAC577D6238549C560085C9AE3AE" class="Insecta" family="Corydiidae" genus="Eucorydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucorydia maxwelli" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="maxwelli">Eucorydia maxwelli</taxonomicName>
:
<bibRefCitation id="ADE8CB77F97B0CCFBC8C9DBB3FCC0DE4" author="Hebard, M" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="32" pageNumber="49" pagination="1 - 109" title="Studies in Malayan Blattidae (Orthoptera)." volume="81" year="1929">Hebard 1929</bibRefCitation>
: 14.
</paragraph>
<paragraph id="9D53EB3D9014053475B1CBCFF5F87323" pageId="29" pageNumber="46">
<taxonomicName id="10BE69131708ADCCC2108527998598B0" class="Insecta" family="Corydiidae" genus="Eucorydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucorydia westwoodi" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="westwoodi">Eucorydia westwoodi</taxonomicName>
:
<bibRefCitation id="139BC5D521FD2DD32B6A7C7247C502F2" author="Hanitsch, R" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="32" pageNumber="49" pagination="48 - 92" title="Beccari and Modigliani's collection of Sumatran Blattidae in the museo civico, Genoa." volume="36" year="1932">Hanitsch 1932</bibRefCitation>
: 80 (1 male from Siboga, Sumatra);
<bibRefCitation id="DD721E573C44368BF2B4306F67C3BBC5" author="Bruijning, CFA" journalOrPublisher="Zoologische Mededelingen" pageId="32" pageNumber="49" pagination="1 - 174" title="Studies on Malayan Blattidae." volume="29" year="1948">Bruijning 1948</bibRefCitation>
: 149;
<bibRefCitation id="2C2CD050ED0DF60CFC937159F97D055E" pageId="29" pageNumber="46">Princis 1963</bibRefCitation>
: 83.
</paragraph>
</subSubSection>
<subSubSection id="67B8F337AA50FBF6149D1C0BFD07312A" pageId="29" pageNumber="46" type="type material">
<paragraph id="5AC1CDA36FDDC692F63C028CCEAA3DDC" pageId="29" pageNumber="46">Type material examined.</paragraph>
<paragraph id="199EBCEC27A6AEE0C7B796610AF0C6CA" pageId="29" pageNumber="46">
TYPE of
<taxonomicName id="BAB18D6606CA3A81A12A01EDC9E5BF99" class="Insecta" family="Corydiidae" genus="Corydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corydia maxwelli" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="maxwelli">Corydia maxwelli</taxonomicName>
, female (OUM, TYPE ORTH 251), two identical labels: &quot;April 1915./ SELANGOR, F. M. S., 3457 ft, 35 m N. N.
<taxonomicName id="E5FD62C1A3EFB5CE4EFA9EF25D7A8CF2" genus="E." lsidName="E." pageId="29" pageNumber="46" rank="genus">E.</taxonomicName>
of Kuala Lumpur, Bukit Kutu./ R. Hanitsch.&quot;; &quot;Bukit Kutu/ Selangor/ April 1915/ 3457&quot;; &quot;♀ TYPE/ HANITSCH./
<taxonomicName id="810F60B656EA772172B0763C7E597E2C" class="Insecta" family="Corydiidae" genus="Corydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corydia maxwelli" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="maxwelli">Corydia maxwelli</taxonomicName>
(handwritten)/ J.S.B., R.A.S./ No. 69. p 126, pl.ii, fig.10.1915.&quot;; &quot;Corydia maxwelli, Han: (handwritten)&quot;; &quot;Eucorydia westwoodi Princis, 1963 (handwritten)&quot;; &quot;TYPE ORTH: 251/
<taxonomicName id="DCDE27F3A5DB2AE60F9301B2B2171DFF" class="Insecta" family="Corydiidae" genus="Corydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corydia maxwelli" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="maxwelli">Corydia maxwelli</taxonomicName>
Hanitsch/ HOPE DEPT. OXFORD&quot;.
</paragraph>
</subSubSection>
<subSubSection id="02854F4E9746A3511DD5F62D0DB5A2C8" pageId="29" pageNumber="46" type="diagnosis">
<paragraph id="8CB87383CC0410A0E4725874DB5DB47F" pageId="29" pageNumber="46">Diagnosis.</paragraph>
<paragraph id="7790AFCD30C23F711EE0FB90A7E6AECB" pageId="29" pageNumber="46">This species may show sexual dimorphism. Male overall length about 11.0 mm (including tegmina); head orange, pronotum black; tegmina orange, each tegmen with a round black spot in the center, and with apex blackish; coxae, femora and abdomen orange, tibiae, tarsi and cerci black. Female overall length about 11.0 mm; head, pronotum, legs and cerci black, tegmina orange; lateral of tegmen with a stripe occupying the entire half of basal margin, and curved inward near the median, the curved part slightly enlarged and quadrate, the entire anal areas are encircled by the two stripes, apex of tegmina blackish; abdomen black, lateral margins orange.</paragraph>
</subSubSection>
<subSubSection id="3ECE30DEC36718A29CE1E6ED169F6825" pageId="29" pageNumber="46" type="remarks">
<paragraph id="A4300B0BBCA94179EB06F0EF675CDDF3" pageId="29" pageNumber="46">Remarks.</paragraph>
<paragraph id="271583BE604832C540EBB1964CF81D79" pageId="29" pageNumber="46">
<bibRefCitation id="E28DE886A710A2DAB063A135A003861A" author="Hanitsch, R" journalOrPublisher="Journal Straits Branch Royal Asiatic Society" pageId="32" pageNumber="49" pagination="17 - 178" title="Malayan Blattidae." volume="69" year="1915">Hanitsch (1915)</bibRefCitation>
described
<taxonomicName id="56E280B5977B9F5DBA99903662A861F1" class="Insecta" family="Corydiidae" genus="Corydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corydia maxwelli" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="species" species="maxwelli">Corydia maxwelli</taxonomicName>
from southeast Asia. He realized this species showed sexual dimorphism, and later synonymized it under
<taxonomicName id="3711B7A906C3329CE1F9CD20B5C617F9" lsidName="E. westwoodi" pageId="29" pageNumber="46" rank="species" species="westwoodi">E. westwoodi</taxonomicName>
(Hanitsch, 1932) based on the female tegmina pattern. However, this was in error:
<taxonomicName id="A1CF9A07985B335BBB92B73129ED2FF3" lsidName="E. westwoodi" pageId="29" pageNumber="46" rank="species" species="westwoodi">E. westwoodi</taxonomicName>
indeed is similar to
<taxonomicName id="25E114844BAC46EA15805A245D14D741" lsidName="E. maxwelli" pageId="29" pageNumber="46" rank="species" species="maxwelli">E. maxwelli</taxonomicName>
by the lateral blackish strips and the blackish apex on tegmina; but the former had an additional large oval spot near the anal area, while the later lacks a spot near the anal area. In addition,
<taxonomicName id="95EF9F85AC442DD01B718955C114310E" lsidName="E. westwoodi" pageId="29" pageNumber="46" rank="species" species="westwoodi">E. westwoodi</taxonomicName>
is distributed in North India and Nepal, while
<taxonomicName id="FD12E4A51FFD6191995962302CE100C9" lsidName="E. maxwelli" pageId="29" pageNumber="46" rank="species" species="maxwelli">E. maxwelli</taxonomicName>
is in southeast Asia. Since the two species are geographically disjunct we believe they are unlikely to be the same species, and we suggest restoring the status of
<taxonomicName id="ECAD4FB84FB76121EE01348CC6CDA5DF" lsidName="E. maxwelli" pageId="29" pageNumber="46" rank="species" species="maxwelli">E. maxwelli</taxonomicName>
.
</paragraph>
<paragraph id="DAA29A39D1ACAB0B4465BFF94EEC479D" pageId="29" pageNumber="46">
The sexual dimorphism of this species is unusual in
<taxonomicName id="7CB886AAD8D3E58794D4560ABA606FF3" class="Insecta" family="Corydiidae" genus="Eucorydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucorydia" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="genus">Eucorydia</taxonomicName>
. Other
<taxonomicName id="86D2A0337C50A453EAE5F50A1084504E" class="Insecta" family="Corydiidae" genus="Eucorydia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucorydia" order="Blattodea" pageId="29" pageNumber="46" phylum="Arthropoda" rank="genus">Eucorydia</taxonomicName>
species show weak sexual dimorphism; the female generally resembles the male, but with shorter tegmina and wings.
<bibRefCitation id="D859CE5E6E8A78D923D200420ED4644D" author="Hanitsch, R" journalOrPublisher="Journal Straits Branch Royal Asiatic Society" pageId="32" pageNumber="49" pagination="17 - 178" title="Malayan Blattidae." volume="69" year="1915">Hanitsch (1915)</bibRefCitation>
at first has stated that the female he described may be another species. Later, he correctly treated it as sexual dimorphism and synonymized
<taxonomicName id="9EFA0D9D08CEEB05DF4EA98501949688" lsidName="C. maxwelli" pageId="29" pageNumber="46" rank="species" species="maxwelli">C. maxwelli</taxonomicName>
under
<taxonomicName id="17DEB323EC0CD37BE6B4EE9AD769D3B2" lsidName="E. westwoodi" pageId="29" pageNumber="46" rank="species" species="westwoodi">E. westwoodi</taxonomicName>
(Hanitsch, 1932), but without convincing reasons. According to the collection data in
<bibRefCitation id="F2CB916F8D540F55C820F7F8175446BD" author="Hanitsch, R" journalOrPublisher="Journal Straits Branch Royal Asiatic Society" pageId="32" pageNumber="49" pagination="17 - 178" title="Malayan Blattidae." volume="69" year="1915">Hanitsch (1915)</bibRefCitation>
, the males and females are not from the same place, which could lead to false pairing. We consider the female more likely to be a separate species.
</paragraph>
</subSubSection>
<subSubSection id="73ABA9AE7290AF4C806EF444EF0CCE4C" pageId="29" pageNumber="46" type="distribution">
<paragraph id="4CE6BAEFA9C6A646DC23137A50BF8900" pageId="29" pageNumber="46">Distribution.</paragraph>
<paragraph id="666BE9DA4180BD278115C2EC6A66A20A" pageId="29" pageNumber="46">Malay Peninsula; Sumatra and Borneo.</paragraph>
</subSubSection>
</treatment>
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