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<mods:title id="44BFB7DD6C251F5381880D47381FDC3B">A monographic catalogue on the systematics and phylogeny of the South American iguanian lizard family Liolaemidae (Squamata, Iguania)</mods:title>
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<mods:namePart id="0D3B547E8A34DAA92B04F169355F568C">Pincheira-Donoso, Daniel</mods:namePart>
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<mods:namePart id="5717B789D91C2C8C6B403EBEEA06EEF2">Scolaro, J. Alejandro</mods:namePart>
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<mods:namePart id="873FC90A89A764B3F285E280698CA4B2">Sura, Piotr</mods:namePart>
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<mods:title id="308361CACEB2728185D19AF6CB751089">Zootaxa</mods:title>
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<mods:date id="EF43CF598574D3FE36EB5B8F38B0065F">2008</mods:date>
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<treatment id="341FFA64FFA3610B6CFBEEE0FC658E86" ID-DOI="http://doi.org/10.5281/zenodo.6789361" ID-GBIF-Taxon="196374877" ID-Zenodo-Dep="6789361" LSID="urn:lsid:plazi:treatment:341FFA64FFA3610B6CFBEEE0FC658E86" httpUri="http://treatment.plazi.org/id/341FFA64FFA3610B6CFBEEE0FC658E86" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection id="F4AC18F9FFA3610B6CFBEEE0FDD88F40" box="[151,535,1535,1561]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="BC094B72FFA3610B6CFBEEE0FDD88F40" blockId="19.[151,1436,1535,2015]" box="[151,535,1535,1561]" pageId="19" pageNumber="20">
<heading id="E741FC1EFFA3610B6CFBEEE0FDD88F40" bold="true" box="[151,535,1535,1561]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="7BB630F1FFA3610B6CFBEEE0FDD88F40" authority="(Philippi)" baseAuthorityName="Philippi" box="[151,535,1535,1561]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="bisignatus">
<emphasis id="8EC29760FFA3610B6CFBEEE0FDD88F40" bold="true" box="[151,535,1535,1561]" pageId="19" pageNumber="20">
<emphasis id="8EC29760FFA3610B6CFBEEE0FE568F40" bold="true" box="[151,409,1535,1561]" italics="true" pageId="19" pageNumber="20">Liolaemus bisignatus</emphasis>
(Philippi)
</emphasis>
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</heading>
</paragraph>
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<paragraph id="BC094B72FFA3610B6CFBED37FAEF8F66" blockId="19.[151,1436,1535,2015]" box="[151,1312,1576,1599]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="9CA66C5CFFA3610B6CFBED37FAEF8F66" box="[151,1312,1576,1599]" pageId="19" pageNumber="20">
<treatmentCitation id="3D176D63FFA3610B6CFBED37FD938F66" author="Philippi, R. A." box="[151,604,1576,1599]" page="166" pageId="19" pageNumber="20" year="1860">
<taxonomicName id="7BB630F1FFA3610B6CFBED37FD938F66" ID-CoL="77XX5" authority="Philippi 1860: 166" authorityName="Philippi" authorityPageNumber="166" authorityYear="1860" box="[151,604,1576,1599]" class="Reptilia" family="Liolaemidae" genus="Proctotretus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="bisignatus">
<emphasis id="8EC29760FFA3610B6CFBED37FE418F66" box="[151,398,1576,1599]" italics="true" pageId="19" pageNumber="20">Proctotretus bisignatus</emphasis>
<materialsCitation id="0CDE412FFFA3610B6DF9ED37FD938F66" ID-GBIF-Occurrence="3827715324" box="[405,604,1576,1599]" country="Chile" location="Caldera" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Atacama">
<bibRefCitation id="D8273683FFA3610B6DF9ED37FD938F66" author="Philippi" box="[405,604,1576,1599]" firstAuthor="Philippi" page="166" pageId="19" pageNumber="20" refId="ref69099" refString="Philippi, R. A. (1860) Reise durch die Wuste Atacama, auf Befehl der chilenischen Regierung in Sommer 1853 - 1854, Liberia de Eduard Anton, Halle, 192 pp." type="book" year="1860">Philippi 1860: 166</bibRefCitation>
</materialsCitation>
</taxonomicName>
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(
<typeStatus id="630DF5D0FFA3610B6E06ED37FD568F66" box="[618,665,1576,1599]" pageId="19" pageNumber="20">type</typeStatus>
locality:
<location id="B9691DA9FFA3610B6E92ED37FC9B8F66" LSID="urn:lsid:plazi:treatment:341FFA64FFA3610B6CFBEEE0FC658E86:B9691DA9FFA3610B6E92ED37FC9B8F66" box="[766,852,1576,1599]" country="Chile" name="Caldera" pageId="19" pageNumber="20" stateProvince="Atacama">Caldera</location>
,
<collectingRegion id="7E728590FFA3610B6F32ED37FBC48F66" box="[862,1035,1576,1599]" country="Chile" name="Atacama" pageId="19" pageNumber="20">Atacama Desert</collectingRegion>
,
<collectingRegion id="7E728590FFA3610B687AED37FB038F66" box="[1046,1228,1576,1599]" country="Chile" name="Atacama" pageId="19" pageNumber="20">Atacama Region</collectingRegion>
,
<collectingCountry id="C4A10BE2FFA3610B68BAED37FADB8F66" box="[1238,1300,1576,1599]" name="Chile" pageId="19" pageNumber="20">Chile</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC094B72FFA3610B6CFBED57FE718F26" blockId="19.[151,1436,1535,2015]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="9CA66C5CFFA3610B6CFBED57FE718F26" pageId="19" pageNumber="20">
<treatmentCitation id="3D176D63FFA3610B6CFBED57FC468F06" author="Muller, L. &amp; Hellmich, W." box="[151,905,1608,1631]" page="132" pageId="19" pageNumber="20" year="1933">
<taxonomicName id="7BB630F1FFA3610B6CFBED57FC468F06" authority="Muller &amp; Hellmich 1933 b: 132" authorityName="Muller &amp; Hellmich" authorityPageNumber="132" authorityYear="1933" box="[151,905,1608,1631]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="nigromaculatus" subSpecies="bisignatus">
<emphasis id="8EC29760FFA3610B6CFBED57FDE48F06" box="[151,555,1608,1631]" italics="true" pageId="19" pageNumber="20">Liolaemus nigromaculatus bisignatus</emphasis>
<bibRefCitation id="D8273683FFA3610B6E5FED57FC468F06" author="Muller &amp; Hellmich" box="[563,905,1608,1631]" firstAuthor="Muller" page="132" pageId="19" pageNumber="20" pagination="128 - 142" refId="ref66814" refString="Muller, L. &amp; Hellmich, W. (1933 b) Beitrage zur Kenntnis der Herpetofauna Chiles. VII. Der Rassenkreis des Liolaemus nigromaculatus. Zoologischer Anzeiger, 103, 128 - 142." type="journal article" year="1933" yearSuffix="b">Müller &amp; Hellmich 1933b: 132</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="3D176D63FFA3610B6FFAED57FBAC8F06" author="Hellmich, W." box="[918,1123,1608,1631]" page="53" pageId="19" pageNumber="20" year="1934">
<bibRefCitation id="D8273683FFA3610B6FFAED57FBAC8F06" author="Hellmich" box="[918,1123,1608,1631]" firstAuthor="Hellmich" page="53" pageId="19" pageNumber="20" refId="ref61273" refString="Hellmich, W. (1934) Die Eidechsen Chiles insbesonders die Gattung Liolaemus, Nach den Sammlungen Goetsch-Hellmich, Abhandlungen Bayerische Akademie der Wissenschaften, 24, 140 pp." type="book" year="1934">Hellmich 1934: 53</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="3D176D63FFA3610B681DED57FA578F06" author="Donoso-Barros, R." box="[1137,1432,1608,1631]" page="170" pageId="19" pageNumber="20" year="1966">
<bibRefCitation id="D8273683FFA3610B681DED57FA578F06" author="Donoso-Barros" box="[1137,1432,1608,1631]" firstAuthor="Donoso-Barros" page="170" pageId="19" pageNumber="20" refId="ref57808" refString="Donoso-Barros, R. (1966 a) Reptiles de Chile. Ediciones de la Universidad de Chile, Santiago, 458 pp." type="book" year="1966" yearSuffix="a">Donoso-Barros 1966a: 170</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="3D176D63FFA3610B6CABED77FE8F8F26" author="Donoso-Barros, R." box="[199,320,1640,1663]" page="189" pageId="19" pageNumber="20" year="1970">
<bibRefCitation id="D8273683FFA3610B6CABED77FE8F8F26" author="Donoso-Barros" box="[199,320,1640,1663]" firstAuthor="Donoso-Barros" page="189" pageId="19" pageNumber="20" pagination="170 - 195" refId="ref58010" refString="Donoso-Barros, R. (1970 a) Liolaemus. In: Peters, J. A. &amp; Donoso-Barros, R. (Ed.), Catalogue of the Neotropical Squamata. Part II. Lizards and Amphisbaenians. Bulletin United States National Museum, 297, 170 - 195." type="journal article" year="1970" yearSuffix="a">1970a: 189</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="3D176D63FFA3610B6D27ED77FE758F26" author="Donoso-Barros, R." box="[331,442,1640,1663]" page="96" pageId="19" pageNumber="20" year="1970">
<bibRefCitation id="D8273683FFA3610B6D27ED77FE758F26" author="Donoso-Barros" box="[331,442,1640,1663]" firstAuthor="Donoso-Barros" page="96" pageId="19" pageNumber="20" pagination="49 - 124" refId="ref58063" refString="Donoso-Barros, R. (1970 b) Catalogo herpetologico chileno. Boletin del Museo Nacional de Historia Natural de Chile, 31, 49 - 124." type="journal article" year="1970" yearSuffix="b">1970b: 96</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC094B72FFA3610B6CFBED97FC048FE6" blockId="19.[151,1436,1535,2015]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="9CA66C5CFFA3610B6CFBED97FC048FE6" pageId="19" pageNumber="20">
<treatmentCitation id="3D176D63FFA3610B6CFBED97FC7D8FC6" author="Muller, L. &amp; Hellmich, W." box="[151,946,1672,1695]" page="135" pageId="19" pageNumber="20" year="1933">
<taxonomicName id="7BB630F1FFA3610B6CFBED97FC7D8FC6" authority="Muller &amp; Hellmich 1933 b: 135" authorityName="Muller &amp; Hellmich" authorityPageNumber="135" authorityYear="1933" box="[151,946,1672,1695]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="nigromaculatus" subSpecies="copiapensis">
<emphasis id="8EC29760FFA3610B6CFBED97FD8D8FC6" box="[151,578,1672,1695]" italics="true" pageId="19" pageNumber="20">Liolaemus nigromaculatus copiapensis</emphasis>
<bibRefCitation id="D8273683FFA3610B6E22ED97FC7D8FC6" author="Muller &amp; Hellmich" box="[590,946,1672,1695]" firstAuthor="Muller" page="135" pageId="19" pageNumber="20" pagination="128 - 142" refId="ref66814" refString="Muller, L. &amp; Hellmich, W. (1933 b) Beitrage zur Kenntnis der Herpetofauna Chiles. VII. Der Rassenkreis des Liolaemus nigromaculatus. Zoologischer Anzeiger, 103, 128 - 142." type="journal article" year="1933" yearSuffix="b">Müller &amp; Hellmich 1933b: 135</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(synonym
<emphasis id="8EC29760FFA3610B6858ED97FB938FC6" box="[1076,1116,1672,1695]" italics="true" pageId="19" pageNumber="20">fide</emphasis>
<bibRefCitation id="D8273683FFA3610B6805ED97FECE8FE6" author="Pincheira-Donoso &amp; Nunez" firstAuthor="Pincheira-Donoso" pageId="19" pageNumber="20" refId="ref69923" refString="Pincheira-Donoso, D. &amp; Nunez, H. (2005) Las especies chilenas del genero Liolaemus (Iguania, Tropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion, Publicacion Ocasional del Museo Nacional de Historia Natural de Chile, Santiago, 486 pp." type="book" year="2005">Pincheira-Donoso &amp; Núñez 2005</bibRefCitation>
);
<treatmentCitation id="3D176D63FFA3610B6D7FEDB7FE158FE6" author="Hellmich, W." box="[275,474,1704,1727]" page="55" pageId="19" pageNumber="20" year="1934">
<bibRefCitation id="D8273683FFA3610B6D7FEDB7FE158FE6" author="Hellmich" box="[275,474,1704,1727]" firstAuthor="Hellmich" page="55" pageId="19" pageNumber="20" refId="ref61273" refString="Hellmich, W. (1934) Die Eidechsen Chiles insbesonders die Gattung Liolaemus, Nach den Sammlungen Goetsch-Hellmich, Abhandlungen Bayerische Akademie der Wissenschaften, 24, 140 pp." type="book" year="1934">Hellmich 1934: 55</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="3D176D63FFA3610B6E2AEDB7FCA88FE6" author="Donoso-Barros, R." box="[582,871,1704,1727]" page="189" pageId="19" pageNumber="20" year="1970">
<bibRefCitation id="D8273683FFA3610B6E2AEDB7FCA88FE6" author="Donoso-Barros" box="[582,871,1704,1727]" firstAuthor="Donoso-Barros" page="189" pageId="19" pageNumber="20" pagination="170 - 195" refId="ref58010" refString="Donoso-Barros, R. (1970 a) Liolaemus. In: Peters, J. A. &amp; Donoso-Barros, R. (Ed.), Catalogue of the Neotropical Squamata. Part II. Lizards and Amphisbaenians. Bulletin United States National Museum, 297, 170 - 195." type="journal article" year="1970" yearSuffix="a">Donoso-Barros 1970a: 189</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC094B72FFA3610B6CFBEDD7FA448F86" blockId="19.[151,1436,1535,2015]" box="[151,1419,1736,1759]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="9CA66C5CFFA3610B6CFBEDD7FA448F86" box="[151,1419,1736,1759]" pageId="19" pageNumber="20">
<treatmentCitation id="3D176D63FFA3610B6CFBEDD7FCA38F86" author="Donoso-Barros, R." box="[151,876,1736,1759]" page="255" pageId="19" pageNumber="20" year="1966">
<taxonomicName id="7BB630F1FFA3610B6CFBEDD7FCA38F86" authority="Donoso-Barros 1966 a: 255" authorityName="Donoso-Barros" authorityYear="1966" box="[151,876,1736,1759]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="nigromaculatus" subSpecies="copiapoensis">
<emphasis id="8EC29760FFA3610B6CFBEDD7FD8B8F86" box="[151,580,1736,1759]" italics="true" pageId="19" pageNumber="20">Liolaemus nigromaculatus copiapoensis</emphasis>
<bibRefCitation id="D8273683FFA3610B6E27EDD7FCA38F86" author="Donoso-Barros" box="[587,876,1736,1759]" firstAuthor="Donoso-Barros" page="255" pageId="19" pageNumber="20" refId="ref57808" refString="Donoso-Barros, R. (1966 a) Reptiles de Chile. Ediciones de la Universidad de Chile, Santiago, 458 pp." type="book" year="1966" yearSuffix="a">Donoso-Barros 1966a: 255</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<emphasis id="8EC29760FFA3610B6F10EDD7FB9E8F86" box="[892,1105,1736,1759]" italics="true" pageId="19" pageNumber="20">nomen enmendatum</emphasis>
; in part); 1970b: 96 (in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC094B72FFA3610B6CFBEDF7FDB18E66" blockId="19.[151,1436,1535,2015]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="9CA66C5CFFA3610B6CFBEDF7FDB18E66" pageId="19" pageNumber="20">
<treatmentCitation id="3D176D63FFA3610B6CFBEDF7FDFA8FA6" author="Ortiz, J." box="[151,565,1768,1791]" page="251" pageId="19" pageNumber="20" year="1981">
<taxonomicName id="7BB630F1FFA3610B6CFBEDF7FDFA8FA6" authority="Ortiz 1981: 251" authorityName="Ortiz" authorityPageNumber="251" authorityYear="1981" box="[151,565,1768,1791]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="bisignatus">
<emphasis id="8EC29760FFA3610B6CFBEDF7FEB38FA6" box="[151,380,1768,1791]" italics="true" pageId="19" pageNumber="20">Liolaemus bisignatus</emphasis>
<bibRefCitation id="D8273683FFA3610B6DE9EDF7FDFA8FA6" author="Ortiz" box="[389,565,1768,1791]" firstAuthor="Ortiz" page="251" pageId="19" pageNumber="20" pagination="247 - 265" refId="ref68353" refString="Ortiz, J. (1981) Estudio multivariado de las especies de Liolaemus del grupo nigromaculatus (Squamata, Iguanidae). Anales del Museo de Historia Natural de Valparaiso, 14, 247 - 265." type="journal article" year="1981">Ortiz 1981: 251</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="3D176D63FFA3610B6E2EEDF7FCDC8FA6" author="Vanzolini, P. E." box="[578,787,1768,1791]" page="13" pageId="19" pageNumber="20" year="1986">
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<bibRefCitation id="D8273683FFA3610B6FD0EC17FB168E46" author="Escobar" box="[956,1241,1800,1823]" etAl="et al." firstAuthor="Escobar" page="1679" pageId="19" pageNumber="20" pagination="1677 - 1690" refId="ref58855" refString="Escobar, C. A., Labra, A. &amp; Niemeyer, H. M. (2001) Chemical composition of precloacal secretions of Liolaemus lizards. Journal of Chemical Ecology, 27, 1677 - 1690." type="journal article" year="2001">
Escobar
<emphasis id="8EC29760FFA3610B6877EC17FB9C8E46" box="[1051,1107,1800,1823]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2001: 1679
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<bibRefCitation id="D8273683FFA3610B688AEC17FA578E46" author="Lobo" box="[1254,1432,1800,1823]" firstAuthor="Lobo" page="139" pageId="19" pageNumber="20" pagination="137 - 150" refId="ref64638" refString="Lobo, F. (2001) A phylogenetic analysis of lizards of the Liolaemus chiliensis group (Iguania: Tropiduridae). Herpetological Journal, 11, 137 - 150." type="journal article" year="2001">Lobo 2001: 139</bibRefCitation>
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<bibRefCitation id="D8273683FFA3610B6CABEC37FEA28E66" author="Sura" box="[199,365,1832,1855]" firstAuthor="Sura" page="428" pageId="19" pageNumber="20" refId="ref73002" refString="Sura, P. 2005. Encyclopedia of extant amphibians and reptiles, Wydawnictwo Fundacja, Nowy Sacz, 544 pp." type="book" year="2005">Sura 2005: 428</bibRefCitation>
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<bibRefCitation id="D8273683FFA3610B6D14EC37FDB68E66" author="Diaz &amp; Lobo" box="[376,633,1832,1855]" firstAuthor="Diaz" page="264" pageId="19" pageNumber="20" pagination="261 - 274" refId="ref57471" refString="Diaz, J. M. &amp; Lobo, F. (2006) Historical biogeography of a clade of Liolaemus (Iguania: Liolaemidae) based on ancestral areas and dispersal-vicariance analysis (DIVA). Papeis Avulsos de Zoologia, 46, 261 - 274." type="journal article" year="2006">Díaz &amp; Lobo 2006: 264</bibRefCitation>
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<emphasis id="8EC29760FFA3610B6CFBEC57FE418E06" box="[151,398,1864,1887]" italics="true" pageId="19" pageNumber="20">Liolaemus copiapensis</emphasis>
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<emphasis id="8EC29760FFA3610B6CFBECB7FE588EE6" box="[151,407,1960,1983]" italics="true" pageId="19" pageNumber="20">Liolaemus copiapoensis</emphasis>
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<bibRefCitation id="D8273683FFA3610B6F51ECB7FB848EE6" author="Nunez &amp; Jaksic" box="[829,1099,1960,1983]" firstAuthor="Nunez" page="71" pageId="19" pageNumber="20" pagination="63 - 91" refId="ref67479" refString="Nunez, H. &amp; Jaksic, F. M. (1992) Lista comentada de los reptiles terrestres de Chile continental. Boletin del Museo Nacional de Historia Natural de Chile, 43, 63 - 91." type="journal article" year="1992">Núñez &amp; Jaksic 1992: 71</bibRefCitation>
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<treatmentCitation id="3D176D63FFA3610B6CFBECD7FC6A8E86" author="Pincheira-Donoso, D. &amp; Nunez, H." box="[151,933,1992,2015]" page="394" pageId="19" pageNumber="20" year="2005">
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<emphasis id="8EC29760FFA3610B6CFBECD7FDCE8E86" box="[151,513,1992,2015]" italics="true" pageId="19" pageNumber="20">Liolaemus (Liolaemus) bisignatus</emphasis>
<bibRefCitation id="D8273683FFA3610B6E64ECD7FC6A8E86" author="Pincheira-Donoso &amp; Nunez" box="[520,933,1992,2015]" firstAuthor="Pincheira-Donoso" page="394" pageId="19" pageNumber="20" refId="ref69923" refString="Pincheira-Donoso, D. &amp; Nunez, H. (2005) Las especies chilenas del genero Liolaemus (Iguania, Tropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion, Publicacion Ocasional del Museo Nacional de Historia Natural de Chile, Santiago, 486 pp." type="book" year="2005">Pincheira-Donoso &amp; Núñez 2005: 394</bibRefCitation>
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.
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