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<document id="F792E37BF88B81AF23F493A6E77333DB" ID-DOI="10.4314/met.v34i1.8" ID-ISSN="2307-5031" ID-Zenodo-Dep="14119506" ID-ZooBank="BAAA5966-5EC8-4D47-A025-E64391B62E45" checkinTime="1731462245380" checkinUser="felipe" docAuthor="Zilli, Alberto, László, Gyula M., Kingston, Anthony J. &amp; Larsen, Knud" docDate="2023" docId="077AF01EFFE2C909FF53A62EB72E1240" docLanguage="en" docName="Metamorphosis.34.97-104.pdf" docOrigin="Metamorphosis 34 (1)" docSource="http://dx.doi.org/10.4314/met.v34i1.8" docStyle="DocumentStyle:73CBAE06DE6A380ACC54A94377F5FCBA.1:Metamorphosis.2021-.journal_article" docStyleId="73CBAE06DE6A380ACC54A94377F5FCBA" docStyleName="Metamorphosis.2021-.journal_article" docStyleVersion="1" docTitle="Nyodes paludicola Zilli &amp; László &amp; Kingston &amp; Larsen 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="102" masterDocId="FB438866FFE3C90CFFD7A34FB3731577" masterDocTitle="A peculiar new species of Nyodes Laporte, 1970 (Lepidoptera: Noctuidae)" masterLastPageNumber="104" masterPageNumber="97" pageNumber="98" updateTime="1731462740784" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="F0C66379A0A29A8947AB6E98EC80D07D">A peculiar new species of Nyodes Laporte, 1970 (Lepidoptera: Noctuidae)</mods:title>
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<mods:namePart id="664B837130973F58C89CEC565A0C7975">Zilli, Alberto</mods:namePart>
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<mods:namePart id="9DA7E7306900B79F8A043BB1E51E45EC">László, Gyula M.</mods:namePart>
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<mods:namePart id="3E1A704289E522C7F32EBE8B2EFBB5DB">Kingston, Anthony J.</mods:namePart>
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<mods:namePart id="71C54389AB39F1AAA3C034A5E675031F">Larsen, Knud</mods:namePart>
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<mods:date id="6E5EA4355788EFA347E002814082EC47">2023</mods:date>
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<taxonomicName id="48D33A8BFFE2C90DFF53A62EB238100E" authority="Zilli &amp; László &amp; Kingston &amp; Larsen, 2023" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[132,331,1377,1401]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="paludicola" status="sp. nov.">
<emphasis id="BDA79D1AFFE2C90DFF53A62EB238100E" box="[132,331,1377,1401]" italics="true" pageId="1" pageNumber="98">Nyodes paludicola</emphasis>
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<taxonomicNameLabel id="A6942061FFE2C90DFE85A62DB2DA100D" box="[338,425,1378,1402]" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="17E85D8DFFE2C90DFE60A62EB173100E" box="[439,512,1377,1401]" captionStart="Figures 1" captionStartId="2.[132,209,1869,1890]" captionTargetBox="[136,1448,158,1836]" captionTargetId="figure-232@2.[136,1448,158,1836]" captionTargetPageId="2" captionText="Figures 1 to 9 Adults. 1 Nyodes paludicola sp. nov., HT, ♂, Republic of Congo (ANHRT). 2 Idem, PT, ♂, Liberia (ANHRT). 3 Idem, PT, ♂, Cameroon (RMCA). 4 Idem, PT, ♂, Zambia (ANHRT). 5 Idem, PT, ♂, Kenya (RCAK). 6 Idem, PT, ♂, Uganda, gen. slide No. NHMUK 010318218 (prep. by A. Zilli) (NHMUK). 7 Idem, PT, ♀, DRC (RMCA). 8 Idem, PT, ♀, Sierra Leone (ANHRT). 9 Idem, PT, ♀, Zambia (ANHRT)." figureDoi="http://doi.org/10.5281/zenodo.14119508" httpUri="https://zenodo.org/record/14119508/files/figure.png" pageId="1" pageNumber="98">Figs. 1</figureCitation>
9, 1217, 2022)
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urn:lsid:zoobank.org:act:
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:
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, “
<collectingCountry id="F7C40198FFE2C90DFEF2A6FEB14210BE" box="[293,561,1457,1481]" name="Republic of the Congo" pageId="1" pageNumber="98">REPUBLIC OF CONGO</collectingCountry>
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/
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, /
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bai /
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,
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,
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/ Bakala N., M., Dérozier, V., / Kirk-Spriggs, A.,
<collectorName id="222624DEFFE2C90DFEA0A542B2951352" box="[375,486,1549,1573]" pageId="1" pageNumber="98">László, G.</collectorName>
leg. / ANHRT:2023.3” // unique id.: ANHRTUK 00301104 // gen. slide
<collectorName id="222624DEFFE2C90DFD40A563B1B11334" box="[663,706,1580,1603]" pageId="1" pageNumber="98">No.</collectorName>
: LG 6048 (
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).
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:
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<materialsCitation id="3FBB4B55FFE2C90DFF53A5E9B386122F" collectingDate="2020-01-01" collectingDateMax="2020-01-09" collectingDateMin="2020-01-01" collectionCode="ANHRT" collectorName="Sinyaev, V. &amp; Kalnoi, G." country="Sierra Leone" county="Bongema" elevation="370" latitude="8.488611" location="Kono Province" longLatPrecision="21" longitude="-10.773889" municipality="Gori Hills" pageId="1" pageNumber="98" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="F7C40198FFE2C90DFF53A5E9B26A13C9" box="[132,281,1702,1726]" name="Sierra Leone" pageId="1" pageNumber="98">Sierra Leone</collectingCountry>
:
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,
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,
<collectingMunicipality id="6F08DB72FFE2C90DFDE2A5E9B1DD13C9" box="[565,686,1702,1726]" pageId="1" pageNumber="98">Gori Hills</collectingMunicipality>
, near
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,
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,
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, leg.
<collectorName id="222624DEFFE2C90DFF62A5ABB245138C" box="[181,310,1764,1788]" pageId="1" pageNumber="98">Sinyaev, V.</collectorName>
&amp;
<collectorName id="222624DEFFE2C90DFE8AA5ABB2A1138B" box="[349,466,1764,1788]" pageId="1" pageNumber="98">Kalnoi, G.</collectorName>
, ANHRT:2020.14, unique and DNA sample id.: ANHRTUK 00290107, BOLD process id.: ANLMO4267-23, gen. slide No.: LG 6055 (
<collectionCode id="E9C2D9CDFFE2C90DFF5BA40FB39D122F" box="[140,238,1856,1880]" pageId="1" pageNumber="98">ANHRT</collectionCode>
)
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.
</paragraph>
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<materialsCitation id="3FBB4B55FFE2C90DFF53A410B2291D67" collectingDate="2017-03-16" collectingDateMax="2017-03-17" collectingDateMin="2017-03-16" collectorName="Safian, Sz. &amp; Simonics, G." country="Liberia" elevation="701" latitude="7.5583334" location="Mount Gangra" longLatPrecision="21" longitude="-8.637777" municipality="Mountains" pageId="1" pageNumber="98" specimenCount="1" specimenCount-male="1" stateProvince="Nimba">
<collectingCountry id="F7C40198FFE2C90DFF53A410B3A21200" box="[132,209,1887,1911]" name="Liberia" pageId="1" pageNumber="98">Liberia</collectingCountry>
:
<specimenCount id="99D58A81FFE2C90DFF32A410B2751201" box="[229,262,1887,1911]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
,
<collectingRegion id="4D178FEAFFE2C90DFECEA410B2121200" box="[281,353,1887,1911]" country="Liberia" name="Nimba" pageId="1" pageNumber="98">Nimba</collectingRegion>
<collectingMunicipality id="6F08DB72FFE2C90DFEB9A410B2901200" box="[366,483,1887,1911]" pageId="1" pageNumber="98">Mountains</collectingMunicipality>
,
<location id="8A0C17D3FFE2C90DFE24A410B1E71200" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFE24A410B1E71200" box="[499,660,1887,1911]" country="Liberia" latitude="7.5583334" longLatPrecision="21" longitude="-8.637777" municipality="Mountains" name="Mount Gangra" pageId="1" pageNumber="98" stateProvince="Nimba">Mount Gangra</location>
western slope,
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,
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,
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<date id="FB6D67C8FFE2C90DFD85A432B18312E2" box="[594,752,1917,1941]" pageId="1" pageNumber="98" value="2017-03-16" valueMax="2017-03-17" valueMin="2017-03-16">
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</date>
,
<collectingMethod id="5692391FFFE2C90DFF53A4D3B38B12C4" box="[132,248,1948,1972]" pageId="1" pageNumber="98">Light Trap</collectingMethod>
(250W blended bulb) &amp;
<collectingMethod id="5692391FFFE2C90DFDD2A4D3B27812A5" pageId="1" pageNumber="98">cold cathode UV light bucket trap</collectingMethod>
(8W),
<collectorName id="222624DEFFE2C90DFEA1A4F5B28812A5" box="[374,507,1978,2002]" pageId="1" pageNumber="98">Sáfián, Sz.</collectorName>
,
<collectorName id="222624DEFFE2C90DFDC1A4F5B1DC12A5" box="[534,687,1978,2002]" pageId="1" pageNumber="98">Simonics, G.</collectorName>
leg., ANHRT:2017.36, unique id.: ANHRTUK 00056183, gen. slide No.: LG 6054
</materialsCitation>
;
<materialsCitation id="3FBB4B55FFE2C90DFEB1A4B7B793147D" collectingDate="2017-03-17" collectingDateMax="2017-03-25" collectingDateMin="2017-03-17" collectionCode="ANHRT" collectorName="Safian, Sz. &amp; Simonics, G." country="Liberia" elevation="904" latitude="7.546111" location="Mt Gangra" longLatPrecision="15" longitude="-8.635933" municipality="Mts" pageId="1" pageNumber="98" specimenCount="1" specimenCount-male="1" stateProvince="Nimba">
<specimenCount id="99D58A81FFE2C90DFEB1A4B7B2FB1D67" box="[358,392,2040,2064]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
,
<collectingRegion id="4D178FEAFFE2C90DFE40A4B7B2AC1D67" box="[407,479,2040,2064]" country="Liberia" name="Nimba" pageId="1" pageNumber="98">Nimba</collectingRegion>
<collectingMunicipality id="6F08DB72FFE2C90DFE30A4B6B1601D67" box="[487,531,2041,2064]" pageId="1" pageNumber="98">Mts</collectingMunicipality>
,
<location id="8A0C17D3FFE2C90DFDF7A4B6B1E61D67" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFDF7A4B6B1E61D67" box="[544,661,2040,2064]" country="Liberia" latitude="7.546111" longLatPrecision="15" longitude="-8.635933" municipality="Mts" name="Mt Gangra" pageId="1" pageNumber="98" stateProvince="Nimba">Mt Gangra</location>
summit,
<quantity id="482BECEDFFE2C90DFF53AB59B3B01D59" box="[132,195,2070,2094]" metricMagnitude="2" metricUnit="m" metricValue="9.04" pageId="1" pageNumber="98" unit="m" value="904.0">
<elevation id="04FEA63BFFE2C90DFF53AB59B3B01D59" box="[132,195,2070,2094]" metricMagnitude="2" metricUnit="m" metricValue="9.04" pageId="1" pageNumber="98" unit="m" value="904.0">904m</elevation>
</quantity>
,
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,
<geoCoordinate id="EAE727CFFFE2C90DFEB6AB59B17A1D59" box="[353,521,2070,2094]" degrees="08" direction="west" minutes="38" orientation="longitude" pageId="1" pageNumber="98" precision="1" seconds="09.36" value="-8.635933">08°38'09.36&quot;W</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFDC0AB59B1C61D59" box="[535,693,2070,2094]" pageId="1" pageNumber="98" value="2017-03-17" valueMax="2017-03-25" valueMin="2017-03-17">
<collectingDate id="EB299E20FFE2C90DFDC0AB59B1C61D59" box="[535,693,2070,2094]" pageId="1" pageNumber="98" value="2017-03-17" valueMax="2017-03-25" valueMin="2017-03-17">1725.iii.2017</collectingDate>
</date>
,
<collectingMethod id="5692391FFFE2C90DFD14AB59B3DD1D3A" pageId="1" pageNumber="98">light trap</collectingMethod>
(250W blended bulb)
<collectingMethod id="5692391FFFE2C90DFE6AAB7AB0B415D9" box="[445,967,151,2125]" pageId="1" pageNumber="98">and cold cathode UV light bucket trap</collectingMethod>
(8W),
<collectorName id="222624DEFFE2C90DFBE5A3D8B7C515D8" box="[1074,1206,151,175]" pageId="1" pageNumber="98">Sáfián, Sz.</collectorName>
,
<collectorName id="222624DEFFE2C90DFB06A3D8B61815D8" box="[1233,1387,151,175]" pageId="1" pageNumber="98">Simonics, G.</collectorName>
leg., ANHRT:2017.36, unique and DNA sample id.: ANHRTUK 00214308, BOLD process id.: ANLMO2501- 23, gen. slide No.: LG 5750 (
<collectionCode id="E9C2D9CDFFE2C90DFBA0A3BDB7AA147D" box="[1143,1241,242,266]" pageId="1" pageNumber="98">ANHRT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A25EB6E214F2" blockId="1.[832,1456,151,1923]" pageId="1" pageNumber="98">
<materialsCitation id="3FBB4B55FFE2C90DFC97A25EB70D143F" collectingDate="1992-10-23" collectorName="K. Maes &amp; Nos" country="Cameroon" location="Cameroon" pageId="1" pageNumber="98" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F7C40198FFE2C90DFC97A25EB0DC145E" box="[832,943,273,297]" name="Cameroon" pageId="1" pageNumber="98">Cameroon</collectingCountry>
:
<specimenCount id="99D58A81FFE2C90DFC69A25EB080145E" box="[958,1011,273,297]" count="2" pageId="1" pageNumber="98" type="male">2♂♂</specimenCount>
, Dschang,
<date id="FB6D67C8FFE2C90DFBB9A25EB7AF145E" box="[1134,1244,273,297]" pageId="1" pageNumber="98" value="1992-10-23">
<collectingDate id="EB299E20FFE2C90DFBB9A25EB7AF145E" box="[1134,1244,273,297]" pageId="1" pageNumber="98" value="1992-10-23">23.x.1992</collectingDate>
</date>
,
<collectorName id="222624DEFFE2C90DFB3FA25DB637145E" box="[1256,1348,274,297]" pageId="1" pageNumber="98">K. Maes</collectorName>
leg., gen. slide
<collectorName id="222624DEFFE2C90DFCAFA27EB0D4143F" box="[888,935,305,328]" pageId="1" pageNumber="98">Nos</collectorName>
: LG 5898, LG 5907
</materialsCitation>
;
<specimenCount id="99D58A81FFE2C90DFB5EA27FB7D9143F" box="[1161,1194,304,328]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
, same site and collector,
<date id="FB6D67C8FFE2C90DFC97A200B0DF1410" box="[832,940,335,359]" pageId="1" pageNumber="98" value="1992-10-29">29.x.1992</date>
, gen. slide No.: LG 5899;
<materialsCitation id="3FBB4B55FFE2C90DFB34A200B6FE14F2" collectingDate="1992-10-26" collectionCode="RMCA" collectorName="No." country="Cameroon" pageId="1" pageNumber="98" specimenCount="1" specimenCount-male="1">
<specimenCount id="99D58A81FFE2C90DFB34A200B6761411" box="[1251,1285,335,359]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
, same site and collector,
<date id="FB6D67C8FFE2C90DFC7DA222B76414F2" box="[938,1047,365,389]" pageId="1" pageNumber="98" value="1992-10-26">
<collectingDate id="EB299E20FFE2C90DFC7DA222B76414F2" box="[938,1047,365,389]" pageId="1" pageNumber="98" value="1992-10-26">26.x.1992</collectingDate>
</date>
, gen. slide
<collectorName id="222624DEFFE2C90DFB58A221B7C914F2" box="[1167,1210,366,389]" pageId="1" pageNumber="98">No.</collectorName>
: LG 5900 (
<collectionCode id="E9C2D9CDFFE2C90DFAE5A222B6F514F2" box="[1330,1414,365,389]" country="Belgium" name="Royal Museum for Central Africa" pageId="1" pageNumber="98" type="Museum">RMCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A2C3B0C616BB" blockId="1.[832,1456,151,1923]" pageId="1" pageNumber="98">
<materialsCitation id="3FBB4B55FFE2C90DFC97A2C3B6091768" collectingDate="2023-02-14" collectingDateMax="2023-02-20" collectingDateMin="2023-02-14" collectorName="Bakala N. &amp; Derozier, V. &amp; Kirk-Spriggs, A. &amp; Laszlo, G. &amp; No." country="Republic of the Congo" elevation="372" latitude="2.24" location="Nouabale-Ndoki National Park" longLatPrecision="21" longitude="16.397778" municipality="Mbeli" pageId="1" pageNumber="98" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F7C40198FFE2C90DFC97A2C3B77814D3" box="[832,1035,396,420]" name="Republic of the Congo" pageId="1" pageNumber="98">Republic of Congo</collectingCountry>
:
<specimenCount id="99D58A81FFE2C90DFBCEA2C3B73C14D3" box="[1049,1103,396,420]" count="2" pageId="1" pageNumber="98" type="male">2♂♂</specimenCount>
,
<location id="8A0C17D3FFE2C90DFB8BA2C3B6DF14D3" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFB8BA2C3B6DF14D3" box="[1116,1452,396,420]" country="Republic of the Congo" latitude="2.24" longLatPrecision="21" longitude="16.397778" municipality="Mbeli" name="Nouabale-Ndoki National Park" pageId="1" pageNumber="98">Nouabale-Ndoki National Park</location>
,
<quantity id="482BECEDFFE2C90DFC97A2E5B00C14B5" box="[832,895,426,450]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="1" pageNumber="98" unit="m" value="372.0">
<elevation id="04FEA63BFFE2C90DFC97A2E5B00C14B5" box="[832,895,426,450]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="1" pageNumber="98" unit="m" value="372.0">372m</elevation>
</quantity>
,
<collectingMunicipality id="6F08DB72FFE2C90DFC4BA2E5B0AF14B5" box="[924,988,426,450]" pageId="1" pageNumber="98">Mbeli</collectingMunicipality>
camp,
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,
<geoCoordinate id="EAE727CFFFE2C90DFB3AA2E5B61914B5" box="[1261,1386,426,450]" degrees="16" direction="east" minutes="23" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="52" value="16.397778">16°23'52&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFA5FA2E5B0DC1496" pageId="1" pageNumber="98" value="2023-02-14" valueMax="2023-02-20" valueMin="2023-02-14">
<collectingDate id="EB299E20FFE2C90DFA5FA2E5B0DC1496" pageId="1" pageNumber="98" value="2023-02-14" valueMax="2023-02-20" valueMin="2023-02-14">14 20.ii.2023</collectingDate>
</date>
,
<collectingMethod id="5692391FFFE2C90DFC6CA286B71D1496" box="[955,1134,457,481]" pageId="1" pageNumber="98">actinic light trap</collectingMethod>
,
<collectorName id="222624DEFFE2C90DFBADA286B79B1496" box="[1146,1256,457,481]" pageId="1" pageNumber="98">Bakala N.</collectorName>
, M.,
<collectorName id="222624DEFFE2C90DFAF5A286B6D91496" box="[1314,1450,457,481]" pageId="1" pageNumber="98">Dérozier, V.</collectorName>
,
<collectorName id="222624DEFFE2C90DFC97A2A7B0871777" box="[832,1012,488,512]" pageId="1" pageNumber="98">Kirk-Spriggs, A.</collectorName>
,
<collectorName id="222624DEFFE2C90DFBD7A2A7B7031777" box="[1024,1136,488,512]" pageId="1" pageNumber="98">László, G.</collectorName>
leg., ANHRT:2023.3, unique id.: ANHRTUK 00301089, gen. slide
<collectorName id="222624DEFFE2C90DFB09A148B6791769" box="[1246,1290,519,542]" pageId="1" pageNumber="98">No.</collectorName>
: LG 6056
</materialsCitation>
;
<materialsCitation id="3FBB4B55FFE2C90DFA50A148B72617CF" collectingDate="2023-02-07" collectingDateMax="2023-02-14" collectingDateMin="2023-02-07" collectorName="Bakala N. &amp; Derozier, V. &amp; Kirk-Spriggs, A. &amp; Laszlo, G. &amp; No." country="Republic of Congo" elevation="365" latitude="2.3641665" location="Nouabale-Ndoki National Park" longLatPrecision="21" longitude="16.27389" municipality="Mondika" pageId="1" pageNumber="98" specimenCount="1" specimenCount-female="1">
<specimenCount id="99D58A81FFE2C90DFA50A148B6DA1769" box="[1415,1449,519,543]" count="1" pageId="1" pageNumber="98" type="female">1♀</specimenCount>
,
<location id="8A0C17D3FFE2C90DFC97A16AB7EE174A" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFC97A16AB7EE174A" box="[832,1181,549,573]" country="Republic of Congo" latitude="2.3641665" longLatPrecision="21" longitude="16.27389" municipality="Mondika" name="Nouabale-Ndoki National Park" pageId="1" pageNumber="98">Nouabale-Ndoki National Park</location>
,
<quantity id="482BECEDFFE2C90DFB78A16AB79D174A" box="[1199,1262,549,573]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="1" pageNumber="98" unit="m" value="365.0">
<elevation id="04FEA63BFFE2C90DFB78A16AB79D174A" box="[1199,1262,549,573]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="1" pageNumber="98" unit="m" value="365.0">365m</elevation>
</quantity>
,
<collectingMunicipality id="6F08DB72FFE2C90DFAD7A16AB613174A" box="[1280,1376,549,573]" pageId="1" pageNumber="98">Mondika</collectingMunicipality>
camp,
<geoCoordinate id="EAE727CFFFE2C90DFC97A10BB0B3172C" box="[832,960,580,604]" degrees="02" direction="north" minutes="21" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="51" value="2.3641665">02°21'51&quot;N</geoCoordinate>
,
<geoCoordinate id="EAE727CFFFE2C90DFC1BA10BB73A172C" box="[972,1097,580,604]" degrees="16" direction="east" minutes="16" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="26" value="16.27389">16°16'26&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFB82A10BB798172B" box="[1109,1259,580,604]" pageId="1" pageNumber="98" value="2023-02-07" valueMax="2023-02-14" valueMin="2023-02-07">
<collectingDate id="EB299E20FFE2C90DFB82A10BB798172B" box="[1109,1259,580,604]" pageId="1" pageNumber="98" value="2023-02-07" valueMax="2023-02-14" valueMin="2023-02-07">0714.ii.2023</collectingDate>
</date>
,
<collectingMethod id="5692391FFFE2C90DFB2FA10BB6D8172C" box="[1272,1451,580,604]" pageId="1" pageNumber="98">actinic light trap</collectingMethod>
,
<collectorName id="222624DEFFE2C90DFC97A12DB0DD170D" box="[832,942,610,634]" pageId="1" pageNumber="98">Bakala N.</collectorName>
, M.,
<collectorName id="222624DEFFE2C90DFC3FA12DB703170D" box="[1000,1136,610,634]" pageId="1" pageNumber="98">Dérozier, V.</collectorName>
,
<collectorName id="222624DEFFE2C90DFBABA12DB640170D" box="[1148,1331,610,634]" pageId="1" pageNumber="98">Kirk-Spriggs, A.</collectorName>
,
<collectorName id="222624DEFFE2C90DFAE8A12DB6DC170D" box="[1343,1455,610,634]" pageId="1" pageNumber="98">László, G.</collectorName>
leg., ANHRT:2023.3, unique id.: ANHRTUK 00301057, gen. slide
<collectorName id="222624DEFFE2C90DFC62A1EEB09317CF" box="[949,992,673,696]" pageId="1" pageNumber="98">No.</collectorName>
: LG 6049
</materialsCitation>
;
<materialsCitation id="3FBB4B55FFE2C90DFBB3A1EFB7CF1644" collectingDate="2022-09-29" collectingDateMax="2022-10-01" collectingDateMin="2022-09-29" collectorName="Derozier, V. &amp; Fouka, B. &amp; Kirk-Spriggs, A. &amp; Takano, H." country="Republic of Congo" elevation="352" latitude="2.2133334" location="Nouabale-Ndoki National Park" longLatPrecision="21" longitude="16.39611" municipality="Ndoki Formation" pageId="1" pageNumber="98" specimenCount="1" specimenCount-male="1">
<specimenCount id="99D58A81FFE2C90DFBB3A1EFB7F517CF" box="[1124,1158,672,696]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
,
<location id="8A0C17D3FFE2C90DFB40A1EFB00717A0" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFB40A1EFB00717A0" country="Republic of Congo" latitude="2.2133334" longLatPrecision="21" longitude="16.39611" municipality="Ndoki Formation" name="Nouabale-Ndoki National Park" pageId="1" pageNumber="98">Nouabale-Ndoki National Park</location>
,
<collectingMunicipality id="6F08DB72FFE2C90DFC55A1F0B73217A0" box="[898,1089,703,727]" pageId="1" pageNumber="98">Ndoki Formation</collectingMunicipality>
,
<quantity id="482BECEDFFE2C90DFB87A1F0B7FC17A0" box="[1104,1167,703,727]" metricMagnitude="2" metricUnit="m" metricValue="3.52" pageId="1" pageNumber="98" unit="m" value="352.0">
<elevation id="04FEA63BFFE2C90DFB87A1F0B7FC17A0" box="[1104,1167,703,727]" metricMagnitude="2" metricUnit="m" metricValue="3.52" pageId="1" pageNumber="98" unit="m" value="352.0">352m</elevation>
</quantity>
,
<geoCoordinate id="EAE727CFFFE2C90DFB49A1F0B65317A1" box="[1182,1312,703,727]" degrees="02" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="48" value="2.2133334">02°12'48&quot;N</geoCoordinate>
,
<geoCoordinate id="EAE727CFFFE2C90DFAF9A1F0B6D817A1" box="[1326,1451,703,727]" degrees="16" direction="east" minutes="23" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="46" value="16.39611">16°23'46&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFC97A192B09A1782" box="[832,1001,733,757]" pageId="1" pageNumber="98" value="2022-09-29" valueMax="2022-10-01" valueMin="2022-09-29">
<collectingDate id="EB299E20FFE2C90DFC97A192B09A1782" box="[832,1001,733,757]" pageId="1" pageNumber="98" value="2022-09-29" valueMax="2022-10-01" valueMin="2022-09-29">29.ix.1.x.2022</collectingDate>
</date>
,
<collectingMethod id="5692391FFFE2C90DFC2FA191B7E71782" box="[1016,1172,733,757]" pageId="1" pageNumber="98">MV light trap</collectingMethod>
,
<collectorName id="222624DEFFE2C90DFB74A192B65E1782" box="[1187,1325,733,757]" pageId="1" pageNumber="98">Dérozier, V.</collectorName>
,
<collectorName id="222624DEFFE2C90DFAEBA192B6DA1782" box="[1340,1449,733,757]" pageId="1" pageNumber="98">Fouka, B.</collectorName>
,
<collectorName id="222624DEFFE2C90DFC97A1B3B7731663" box="[832,1024,764,788]" pageId="1" pageNumber="98">Kirk-Spriggs, A.</collectorName>
,
<collectorName id="222624DEFFE2C90DFBC0A1B3B7E81663" box="[1047,1179,764,788]" pageId="1" pageNumber="98">Takano, H.</collectorName>
leg., ANHRT:2022.14, unique id.: ANHRTUK 00253601
</materialsCitation>
;
<materialsCitation id="3FBB4B55FFE2C90DFB1CA054B0C316BB" collectingDate="2023-05-10" collectingDateMax="2023-05-14" collectingDateMin="2023-05-10" collectionCode="ANHRT" collectorName="Derozier, V. &amp; Kirk-Spriggs, A. &amp; Laszlo, G. &amp; Mvouende, S." country="Republic of Congo" elevation="338" latitude="2.2325" location="Nouabale-Ndoki National Park" longLatPrecision="21" longitude="16.20389" municipality="Wali Forest" pageId="1" pageNumber="98" specimenCount="1" specimenCount-male="1">
<specimenCount id="99D58A81FFE2C90DFB1CA054B79F1645" box="[1227,1260,795,819]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
,
<location id="8A0C17D3FFE2C90DFB29A054B09E1626" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFB29A054B09E1626" country="Republic of Congo" latitude="2.2325" longLatPrecision="21" longitude="16.20389" municipality="Wali Forest" name="Nouabale-Ndoki National Park" pageId="1" pageNumber="98">Nouabale-Ndoki National Park</location>
,
<collectingMunicipality id="6F08DB72FFE2C90DFBD8A076B7D41626" box="[1039,1191,825,849]" pageId="1" pageNumber="98">Wali Forest</collectingMunicipality>
,
<quantity id="482BECEDFFE2C90DFB1EA076B67B1626" box="[1225,1288,825,849]" metricMagnitude="2" metricUnit="m" metricValue="3.38" pageId="1" pageNumber="98" unit="m" value="338.0">
<elevation id="04FEA63BFFE2C90DFB1EA076B67B1626" box="[1225,1288,825,849]" metricMagnitude="2" metricUnit="m" metricValue="3.38" pageId="1" pageNumber="98" unit="m" value="338.0">338m</elevation>
</quantity>
,
<geoCoordinate id="EAE727CFFFE2C90DFAFDA076B6D81626" box="[1322,1451,825,849]" degrees="02" direction="north" minutes="13" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="57" value="2.2325">02°13'57&quot;N</geoCoordinate>
,
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,
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,
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,
<collectorName id="222624DEFFE2C90DFAF3A017B6D91607" box="[1316,1450,856,880]" pageId="1" pageNumber="98">Dérozier, V.</collectorName>
,
<collectorName id="222624DEFFE2C90DFC97A038B77516F8" box="[832,1030,887,911]" pageId="1" pageNumber="98">Kirk-Spriggs, A.</collectorName>
,
<collectorName id="222624DEFFE2C90DFBF4A038B7D616F8" box="[1059,1189,887,911]" pageId="1" pageNumber="98">László, G.</collectorName>
,
<collectorName id="222624DEFFE2C90DFB15A038B61B16F8" box="[1218,1384,887,911]" pageId="1" pageNumber="98">Mvouende, S.</collectorName>
leg., ANHRT:2023.6, unique id.: ANHRTUK 00329637 (
<collectionCode id="E9C2D9CDFFE2C90DFC90A0FBB0DA16BB" box="[839,937,948,972]" pageId="1" pageNumber="98">ANHRT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A09CB77E115F" blockId="1.[832,1456,151,1923]" pageId="1" pageNumber="98">
<materialsCitation id="3FBB4B55FFE2C90DFC97A09CB77A115F" collectingDate="1949-05-30" collectionCode="RMCA" collectorName="Dr. Fontaine &amp; No." country="Democratic Republic of the Congo" location="Democratic Republic of the Congo" pageId="1" pageNumber="98" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F7C40198FFE2C90DFC97A09CB7AB169C" box="[832,1240,979,1003]" name="Democratic Republic of the Congo" pageId="1" pageNumber="98">Democratic Republic of the Congo</collectingCountry>
:
<specimenCount id="99D58A81FFE2C90DFB38A09CB662169D" box="[1263,1297,979,1003]" count="1" pageId="1" pageNumber="98" type="female">1♀</specimenCount>
, Leopoldville [Kinshasa],
<date id="FB6D67C8FFE2C90DFC13A0BEB742117E" box="[964,1073,1009,1033]" pageId="1" pageNumber="98" value="1949-05-30">
<collectingDate id="EB299E20FFE2C90DFC13A0BEB742117E" box="[964,1073,1009,1033]" pageId="1" pageNumber="98" value="1949-05-30">30.v.1949</collectingDate>
</date>
,
<collectorName id="222624DEFFE2C90DFBE8A0BDB7B9117E" box="[1087,1226,1009,1033]" pageId="1" pageNumber="98">Dr. Fontaine</collectorName>
leg., gen. slide
<collectorName id="222624DEFFE2C90DFA56A0BDB6DF117E" box="[1409,1452,1010,1033]" pageId="1" pageNumber="98">No.</collectorName>
: LG 5908 (
<collectionCode id="E9C2D9CDFFE2C90DFC7AA75FB772115F" box="[941,1025,1040,1064]" country="Belgium" name="Royal Museum for Central Africa" pageId="1" pageNumber="98" type="Museum">RMCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A760B0CE11F3" blockId="1.[832,1456,151,1923]" pageId="1" pageNumber="98">
<materialsCitation id="3FBB4B55FFE2C90DFC97A760B7551112" collectingDate="1951-07" collectionCode="NHMUK" collectorName="T. H. E Jackson" country="Uganda" location="Uganda" pageId="1" pageNumber="98" specimenCode="NHMUK 015208916" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F7C40198FFE2C90DFC97A760B0E01130" box="[832,915,1071,1095]" name="Uganda" pageId="1" pageNumber="98">Uganda</collectingCountry>
:
<specimenCount id="99D58A81FFE2C90DFC75A760B0B71131" box="[930,964,1071,1095]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
, Entebe [sic],
<date id="FB6D67C8FFE2C90DFBB4A760B7CE1130" box="[1123,1213,1071,1095]" pageId="1" pageNumber="98" value="1951-07">
<collectingDate id="EB299E20FFE2C90DFBB4A760B7CE1130" box="[1123,1213,1071,1095]" pageId="1" pageNumber="98" value="1951-07">vii.1951</collectingDate>
</date>
,
<collectorName id="222624DEFFE2C90DFB1DA760B6141130" box="[1226,1383,1071,1095]" pageId="1" pageNumber="98">T.H.E Jackson</collectorName>
[leg.],
<specimenCode id="DF75E973FFE2C90DFC97A701B7551112" box="[832,1062,1101,1125]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="98" type="Museum">NHMUK 015208916</specimenCode>
</materialsCitation>
,
<materialsCitation id="3FBB4B55FFE2C90DFBE7A701B0CA11F3" collectionCode="NHMUK" country="Uganda" pageId="1" pageNumber="98" specimenCode="NHMUK 010318218" specimenCount="1">
gen. slide No.
<specimenCode id="DF75E973FFE2C90DFB1CA701B6DD1112" box="[1227,1454,1101,1125]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="98" type="Museum">NHMUK 010318218</specimenCode>
(
<collectionCode id="E9C2D9CDFFE2C90DFC90A723B0C111F3" box="[839,946,1132,1156]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="98" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A7C4B0D11188" blockId="1.[832,1456,151,1923]" pageId="1" pageNumber="98">
<materialsCitation id="3FBB4B55FFE2C90DFC97A7C4B0ED1188" collectingDate="2011-11-24" collectionCode="RCAK" collectorName="A. J. Kingston &amp; No." country="Kenya" county="Karamaini Estate" elevation="1550" latitude="-1.0455555" location="Central Province" longLatPrecision="21" longitude="36.99111" municipality="Thika District" pageId="1" pageNumber="98" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F7C40198FFE2C90DFC97A7C4B0F611D4" box="[832,901,1163,1187]" name="Kenya" pageId="1" pageNumber="98">Kenya</collectingCountry>
:
<specimenCount id="99D58A81FFE2C90DFC4FA7C4B0CA11D5" box="[920,953,1163,1187]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
,
<location id="8A0C17D3FFE2C90DFC1CA7C4B7F511D4" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFC1CA7C4B7F511D4" box="[971,1158,1163,1187]" country="Kenya" county="Karamaini Estate" latitude="-1.0455555" longLatPrecision="21" longitude="36.99111" municipality="Thika District" name="Central Province" pageId="1" pageNumber="98">Central Province</location>
,
<collectingMunicipality id="6F08DB72FFE2C90DFB41A7C4B64211D4" box="[1174,1329,1163,1187]" pageId="1" pageNumber="98">Thika District</collectingMunicipality>
,
<locationDeviation id="9EDE582AFFE2C90DFA96A7C4B0F211B6" location="Thika" pageId="1" pageNumber="98">
<quantity id="482BECEDFFE2C90DFA96A7C4B60F11D4" box="[1345,1404,1163,1187]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="98" unit="km" value="8.0">8 km</quantity>
SW Thika
</locationDeviation>
,
<collectingCounty id="660D3984FFE2C90DFC5CA7E6B73511B6" box="[907,1094,1193,1217]" pageId="1" pageNumber="98">Karamaini Estate</collectingCounty>
,
<quantity id="482BECEDFFE2C90DFB87A7E6B7EF11B6" box="[1104,1180,1193,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="98" unit="m" value="1550.0">
<elevation id="04FEA63BFFE2C90DFB87A7E6B7EF11B6" box="[1104,1180,1193,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="98" unit="m" value="1550.0">1550m</elevation>
</quantity>
,
<geoCoordinate id="EAE727CFFFE2C90DFB70A7E6B65111B6" box="[1191,1314,1193,1217]" degrees="01" direction="south" minutes="02" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="44" value="-1.0455555">01°02'44&quot;S</geoCoordinate>
,
<geoCoordinate id="EAE727CFFFE2C90DFAF9A7E6B6D811B6" box="[1326,1451,1193,1217]" degrees="36" direction="east" minutes="59" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="28" value="36.99111">36°59'28&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFC97A787B0C11197" box="[832,946,1224,1248]" pageId="1" pageNumber="98" value="2011-11-24">
<collectingDate id="EB299E20FFE2C90DFC97A787B0C11197" box="[832,946,1224,1248]" pageId="1" pageNumber="98" value="2011-11-24">24.xi.2011</collectingDate>
</date>
,
<collectorName id="222624DEFFE2C90DFC14A787B7291197" box="[963,1114,1224,1248]" pageId="1" pageNumber="98">A.J. Kingston</collectorName>
leg., gen. slide
<collectorName id="222624DEFFE2C90DFAC5A786B64E1197" box="[1298,1341,1225,1248]" pageId="1" pageNumber="98">No.</collectorName>
: LG 5747 (
<collectionCode id="E9C2D9CDFFE2C90DFC90A7A8B0E41188" box="[839,919,1255,1279]" pageId="1" pageNumber="98">RCAK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A64AB0D112F4" blockId="1.[832,1456,151,1923]" pageId="1" pageNumber="98">
<materialsCitation id="3FBB4B55FFE2C90DFC97A64AB6D810EF" collectingDate="2014-05-05" collectingDateMax="2014-05-08" collectingDateMin="2014-05-05" collectorName="Smith &amp; Takano &amp; Chmurova, L. &amp; Smith, L. &amp; No." country="Zambia" elevation="1316" latitude="-11.606112" location="Mwinilunga" longLatPrecision="21" longitude="24.554722" municipality="Nkwaji" pageId="1" pageNumber="98" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F7C40198FFE2C90DFC97A64AB0E2106A" box="[832,913,1285,1309]" name="Zambia" pageId="1" pageNumber="98">Zambia</collectingCountry>
:
<specimenCount id="99D58A81FFE2C90DFC72A64AB0B5106A" box="[933,966,1285,1309]" count="1" pageId="1" pageNumber="98" type="female">1♀</specimenCount>
,
<collectingMunicipality id="6F08DB72FFE2C90DFC0DA64AB758106A" box="[986,1067,1285,1309]" pageId="1" pageNumber="98">Nkwaji</collectingMunicipality>
,
<location id="8A0C17D3FFE2C90DFBEBA64AB7B2106A" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFBEBA64AB7B2106A" box="[1084,1217,1285,1309]" country="Zambia" latitude="-11.606112" longLatPrecision="21" longitude="24.554722" municipality="Nkwaji" name="Mwinilunga" pageId="1" pageNumber="98">Mwinilunga</location>
,
<quantity id="482BECEDFFE2C90DFB06A64AB66D106A" box="[1233,1310,1285,1309]" metricMagnitude="3" metricUnit="m" metricValue="1.316" pageId="1" pageNumber="98" unit="m" value="1316.0">
<elevation id="04FEA63BFFE2C90DFB06A64AB66D106A" box="[1233,1310,1285,1309]" metricMagnitude="3" metricUnit="m" metricValue="1.316" pageId="1" pageNumber="98" unit="m" value="1316.0">1316m</elevation>
</quantity>
,
<geoCoordinate id="EAE727CFFFE2C90DFAF8A64AB6D9106A" box="[1327,1450,1285,1309]" degrees="11" direction="south" minutes="36" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="22" value="-11.606112">11°36'22&quot;S</geoCoordinate>
,
<geoCoordinate id="EAE727CFFFE2C90DFC97A66BB0CE104B" box="[832,957,1316,1340]" degrees="24" direction="east" minutes="33" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="17" value="24.554722">24°33'17&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFC1FA66BB730104B" box="[968,1091,1316,1340]" pageId="1" pageNumber="98" value="2014-05-05" valueMax="2014-05-08" valueMin="2014-05-05">
<collectingDate id="EB299E20FFE2C90DFC1FA66BB730104B" box="[968,1091,1316,1340]" pageId="1" pageNumber="98" value="2014-05-05" valueMax="2014-05-08" valueMin="2014-05-05">58.v.2014</collectingDate>
</date>
,
<collectingMethod id="5692391FFFE2C90DFB99A66BB7C1104B" box="[1102,1202,1316,1340]" pageId="1" pageNumber="98">light trap</collectingMethod>
,
<collectorName id="222624DEFFE2C90DFB6AA66BB78C104B" box="[1213,1279,1316,1340]" pageId="1" pageNumber="98">Smith</collectorName>
, R.,
<collectorName id="222624DEFFE2C90DFAE7A66BB6F0104B" box="[1328,1411,1316,1340]" pageId="1" pageNumber="98">Takano</collectorName>
, H.,
<collectorName id="222624DEFFE2C90DFC97A60CB0A8102D" box="[832,987,1347,1371]" pageId="1" pageNumber="98">Chmurova, L.</collectorName>
,
<collectorName id="222624DEFFE2C90DFC3CA60CB726102D" box="[1003,1109,1347,1371]" pageId="1" pageNumber="98">Smith, L.</collectorName>
leg., ANHRT:2017.11, unique and DNA sample id.: ANHRTUK 00235851, BOLD process id.: ANLMO3001-23, gen. slide
<collectorName id="222624DEFFE2C90DFADDA6CEB64510EF" box="[1290,1334,1409,1432]" pageId="1" pageNumber="98">No.</collectorName>
: LG 6057
</materialsCitation>
;
<materialsCitation id="3FBB4B55FFE2C90DFC97A6D0B0EF13FA" collectingDate="2018-11-09" collectingDateMax="2018-11-15" collectingDateMin="2018-11-09" collectorName="MV Light Trap &amp; Aristophanous, M. &amp; Derozier, V. &amp; Laszlo, G. &amp; Oram, D. &amp; Nos" country="Zambia" county="Lukwakwa" elevation="1147" latitude="-12.661111" location="Crytosepalum forest" longLatPrecision="21" longitude="24.436943" municipality="West Lunga" pageId="1" pageNumber="98" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="99D58A81FFE2C90DFC97A6D0B01210C1" box="[832,865,1439,1463]" count="1" pageId="1" pageNumber="98" type="male">1♂</specimenCount>
,
<specimenCount id="99D58A81FFE2C90DFCA0A6D0B0EA10C1" box="[887,921,1439,1463]" count="1" pageId="1" pageNumber="98" type="female">1♀</specimenCount>
,
<collectingCounty id="660D3984FFE2C90DFC67A6D0B75A10C0" box="[944,1065,1439,1463]" pageId="1" pageNumber="98">Lukwakwa</collectingCounty>
,
<collectingMunicipality id="6F08DB72FFE2C90DFBEAA6D0B7BB10C0" box="[1085,1224,1439,1463]" pageId="1" pageNumber="98">West Lunga</collectingMunicipality>
NP, (
<location id="8A0C17D3FFE2C90DFAC0A6D0B00C10A2" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFAC0A6D0B00C10A2" country="Zambia" county="Lukwakwa" latitude="-12.661111" longLatPrecision="21" longitude="24.436943" municipality="West Lunga" name="Crytosepalum forest" pageId="1" pageNumber="98">Crytosepalum forest</location>
/
<location id="8A0C17D3FFE2C90DFC50A6F2B0A410A2" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFC50A6F2B0A410A2" box="[903,983,1469,1493]" country="Zambia" county="Lukwakwa" latitude="-12.661111" longLatPrecision="21" longitude="24.436943" municipality="West Lunga" name="Dambo" pageId="1" pageNumber="98">Dambo</location>
),
<quantity id="482BECEDFFE2C90DFC2DA6F2B73510A2" box="[1018,1094,1469,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.147" pageId="1" pageNumber="98" unit="m" value="1147.0">
<elevation id="04FEA63BFFE2C90DFC2DA6F2B73510A2" box="[1018,1094,1469,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.147" pageId="1" pageNumber="98" unit="m" value="1147.0">1147m</elevation>
</quantity>
,
<geoCoordinate id="EAE727CFFFE2C90DFBB5A6F2B7AE10A2" box="[1122,1245,1469,1493]" degrees="12" direction="south" minutes="39" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="40" value="-12.661111">12°39'40&quot;S</geoCoordinate>
,
<geoCoordinate id="EAE727CFFFE2C90DFB2CA6F2B60510A2" box="[1275,1398,1469,1493]" degrees="24" direction="east" minutes="26" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="13" value="24.436943">24°26'13&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFA42A6F2B0C71083" pageId="1" pageNumber="98" value="2018-11-09" valueMax="2018-11-15" valueMin="2018-11-09">
<collectingDate id="EB299E20FFE2C90DFA42A6F2B0C71083" pageId="1" pageNumber="98" value="2018-11-09" valueMax="2018-11-15" valueMin="2018-11-09">9 15.xi.2018</collectingDate>
</date>
,
<collectorName id="222624DEFFE2C90DFC17A692B71B1083" box="[960,1128,1500,1524]" pageId="1" pageNumber="98">MV Light Trap</collectorName>
,
<collectorName id="222624DEFFE2C90DFBA3A693B64D1083" box="[1140,1342,1500,1524]" pageId="1" pageNumber="98">Aristophanous, M.</collectorName>
,
<collectorName id="222624DEFFE2C90DFA9DA693B0281365" pageId="1" pageNumber="98">Dérozier, V.</collectorName>
,
<collectorName id="222624DEFFE2C90DFCBCA6B4B0AC1364" box="[875,991,1531,1555]" pageId="1" pageNumber="98">László, G.</collectorName>
,
<collectorName id="222624DEFFE2C90DFC38A6B4B7241365" box="[1007,1111,1531,1555]" pageId="1" pageNumber="98">Oram, D.</collectorName>
leg., ANHRT:2018.40, unique and DNA sample ids: ANHRTUK 00066254, ANHRTUK 00066842, BOLD process ids: ANLMO557-23, ANLMO586-23, gen. slide
<collectorName id="222624DEFFE2C90DFBB4A518B7E11319" box="[1123,1170,1623,1646]" pageId="1" pageNumber="98">Nos</collectorName>
: LG 5780 (male), LG 5714 (female)
</materialsCitation>
;
<materialsCitation id="3FBB4B55FFE2C90DFC7CA53AB0AA127F" collectingDate="2014-11-21" collectingDateMax="2014-11-24" collectingDateMin="2014-11-21" collectionCode="ANHRT" collectorName="Smith &amp; Takano, H." country="Zambia" latitude="-11.865001" location="Jiwundu Swamp" longLatPrecision="21" longitude="25.555555" pageId="1" pageNumber="98" specimenCount="1" specimenCount-female="1">
<specimenCount id="99D58A81FFE2C90DFC7CA53AB0BE13FA" box="[939,973,1653,1677]" count="1" pageId="1" pageNumber="98" type="female">1♀</specimenCount>
,
<location id="8A0C17D3FFE2C90DFC09A53AB7E513FA" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFC09A53AB7E513FA" box="[990,1174,1653,1677]" country="Zambia" latitude="-11.865001" longLatPrecision="21" longitude="25.555555" name="Jiwundu Swamp" pageId="1" pageNumber="98">Jiwundu Swamp</location>
,
<geoCoordinate id="EAE727CFFFE2C90DFB73A53AB66D13FA" box="[1188,1310,1653,1677]" degrees="11" direction="south" minutes="51" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="54" value="-11.865001">11°51'54&quot;S</geoCoordinate>
,
<geoCoordinate id="EAE727CFFFE2C90DFAF9A53AB6D813FA" box="[1326,1451,1653,1677]" degrees="25" direction="east" minutes="33" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="20" value="25.555555">25°33'20&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFC97A5DBB0AF13DB" box="[832,988,1684,1708]" pageId="1" pageNumber="98" value="2014-11-21" valueMax="2014-11-24" valueMin="2014-11-21">
<collectingDate id="EB299E20FFE2C90DFC97A5DBB0AF13DB" box="[832,988,1684,1708]" pageId="1" pageNumber="98" value="2014-11-21" valueMax="2014-11-24" valueMin="2014-11-21">2124.xi.2014</collectingDate>
</date>
,
<collectingMethod id="5692391FFFE2C90DFC38A5DBB72813DB" box="[1007,1115,1684,1708]" pageId="1" pageNumber="98">light trap</collectingMethod>
,
<collectorName id="222624DEFFE2C90DFBB9A5DBB7C313DB" box="[1134,1200,1684,1708]" pageId="1" pageNumber="98">Smith</collectorName>
, R.,
<collectorName id="222624DEFFE2C90DFB25A5DBB60113DB" box="[1266,1394,1684,1708]" pageId="1" pageNumber="98">Takano, H.</collectorName>
leg., ANHRT:2017.12, unique and DNA sample id.: ANHRTUK 00277792, BOLD process id.: ANLMO3661- 23 (
<collectionCode id="E9C2D9CDFFE2C90DFCA7A5BFB0A1127F" box="[880,978,1776,1800]" pageId="1" pageNumber="98">ANHRT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3FBB4B55FFE2C90DFC3CA5BFB0ED12F4" collectingDate="2018-11-06" collectionCode="RCAK" collectorName="Kingston &amp; P. Schmit &amp; No." country="Zambia" elevation="1261" latitude="-12.3575" location="Kafue River" longLatPrecision="21" longitude="27.529167" municipality="Chimfunshi Wildlife Orphanage" pageId="1" pageNumber="98" specimenCount="1" specimenCount-female="1" stateProvince="Copperbelt">
<specimenCount id="99D58A81FFE2C90DFC3CA5BFB77F127F" box="[1003,1036,1776,1800]" count="1" pageId="1" pageNumber="98" type="female">1♀</specimenCount>
,
<collectingRegion id="4D178FEAFFE2C90DFBF6A5BFB675127F" box="[1057,1286,1776,1800]" country="Zambia" name="Copperbelt" pageId="1" pageNumber="98">Copperbelt Province</collectingRegion>
,
<locationDeviation id="9EDE582AFFE2C90DFACEA5BFB0D71250" location="Chingola" pageId="1" pageNumber="98">
<quantity id="482BECEDFFE2C90DFACEA5BFB617127F" box="[1305,1380,1776,1800]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="98" unit="km" value="40.0">40 km</quantity>
W of Chingola
</locationDeviation>
,
<collectingMunicipality id="6F08DB72FFE2C90DFC64A440B66B1250" box="[947,1304,1807,1831]" pageId="1" pageNumber="98">Chimfunshi Wildlife Orphanage</collectingMunicipality>
,
<location id="8A0C17D3FFE2C90DFAF0A440B6DD1250" LSID="urn:lsid:plazi:treatment:077AF01EFFE2C909FF53A62EB72E1240:8A0C17D3FFE2C90DFAF0A440B6DD1250" box="[1319,1454,1807,1831]" country="Zambia" latitude="-12.3575" longLatPrecision="21" longitude="27.529167" municipality="Chimfunshi Wildlife Orphanage" name="Kafue River" pageId="1" pageNumber="98" stateProvince="Copperbelt">Kafue River</location>
floodplain,
<quantity id="482BECEDFFE2C90DFC6EA462B7761232" box="[953,1029,1837,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.261" pageId="1" pageNumber="98" unit="m" value="1261.0">
<elevation id="04FEA63BFFE2C90DFC6EA462B7761232" box="[953,1029,1837,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.261" pageId="1" pageNumber="98" unit="m" value="1261.0">1261m</elevation>
</quantity>
,
<geoCoordinate id="EAE727CFFFE2C90DFBD9A462B7FA1232" box="[1038,1161,1837,1861]" degrees="12" direction="south" minutes="21" orientation="latitude" pageId="1" pageNumber="98" precision="15" seconds="27" value="-12.3575">12°21'27&quot;S</geoCoordinate>
,
<geoCoordinate id="EAE727CFFFE2C90DFB43A462B6621232" box="[1172,1297,1837,1861]" degrees="27" direction="east" minutes="31" orientation="longitude" pageId="1" pageNumber="98" precision="15" seconds="45" value="27.529167">27°31'45&quot;E</geoCoordinate>
,
<date id="FB6D67C8FFE2C90DFACDA462B6F11232" box="[1306,1410,1837,1861]" pageId="1" pageNumber="98" value="2018-11-06">
<collectingDate id="EB299E20FFE2C90DFACDA462B6F11232" box="[1306,1410,1837,1861]" pageId="1" pageNumber="98" value="2018-11-06">6.xi.2018</collectingDate>
</date>
, AJ.
<collectorName id="222624DEFFE2C90DFC97A403B0D11213" box="[832,930,1868,1892]" pageId="1" pageNumber="98">Kingston</collectorName>
&amp;
<collectorName id="222624DEFFE2C90DFC04A402B7311213" box="[979,1090,1868,1892]" pageId="1" pageNumber="98">P. Schmit</collectorName>
leg., gen. slide
<collectorName id="222624DEFFE2C90DFADDA402B6461213" box="[1290,1333,1869,1892]" pageId="1" pageNumber="98">No.</collectorName>
: LG 5748 (
<collectionCode id="E9C2D9CDFFE2C90DFC90A424B0E412F4" box="[839,919,1899,1923]" pageId="1" pageNumber="98">RCAK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C7C91283FFE2C908FC97A4EDB01E1038" lastPageId="4" lastPageNumber="101" pageId="1" pageNumber="98" type="description">
<paragraph id="8F6C4108FFE2C90DFC97A4EDB0B012CD" blockId="1.[832,1455,1954,2101]" box="[832,963,1954,1978]" pageId="1" pageNumber="98">
<emphasis id="BDA79D1AFFE2C90DFC97A4EDB0B012CD" bold="true" box="[832,963,1954,1978]" pageId="1" pageNumber="98">Description</emphasis>
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A48FB08A12AF" blockId="1.[832,1455,1954,2101]" box="[832,1017,1984,2008]" pageId="1" pageNumber="98">
Male (
<figureCitation id="17E85D8DFFE2C90DFC53A48FB0BD12AF" box="[900,974,1984,2008]" captionStart="Figures 1" captionStartId="2.[132,209,1869,1890]" captionTargetBox="[136,1448,158,1836]" captionTargetId="figure-232@2.[136,1448,158,1836]" captionTargetPageId="2" captionText="Figures 1 to 9 Adults. 1 Nyodes paludicola sp. nov., HT, ♂, Republic of Congo (ANHRT). 2 Idem, PT, ♂, Liberia (ANHRT). 3 Idem, PT, ♂, Cameroon (RMCA). 4 Idem, PT, ♂, Zambia (ANHRT). 5 Idem, PT, ♂, Kenya (RCAK). 6 Idem, PT, ♂, Uganda, gen. slide No. NHMUK 010318218 (prep. by A. Zilli) (NHMUK). 7 Idem, PT, ♀, DRC (RMCA). 8 Idem, PT, ♀, Sierra Leone (ANHRT). 9 Idem, PT, ♀, Zambia (ANHRT)." figureDoi="http://doi.org/10.5281/zenodo.14119508" httpUri="https://zenodo.org/record/14119508/files/figure.png" pageId="1" pageNumber="98">Figs. 1</figureCitation>
7):
</paragraph>
<paragraph id="8F6C4108FFE2C90DFC97A490B7F11D42" blockId="1.[832,1455,1954,2101]" pageId="1" pageNumber="98">
Forewing length
<quantity id="482BECEDFFE2C90DFC2FA490B7031280" box="[1016,1136,2015,2039]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="1" pageNumber="98" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
. Ground colour very variable from greyish to chocolate or rusty brown, often with marked purplish or rosy tinge.
</paragraph>
<caption id="DBAC1180FFE1C90EFF53A402B29012C2" ID-DOI="http://doi.org/10.5281/zenodo.14119508" ID-Zenodo-Dep="14119508" httpUri="https://zenodo.org/record/14119508/files/figure.png" pageId="2" pageNumber="99" startId="2.[132,209,1869,1890]" targetBox="[136,1448,158,1836]" targetPageId="2" targetType="figure">
<paragraph id="8F6C4108FFE1C90EFF53A402B29012C2" blockId="2.[132,1456,1868,1973]" pageId="2" pageNumber="99">
<emphasis id="BDA79D1AFFE1C90EFF53A402B2611215" bold="true" box="[132,274,1869,1890]" pageId="2" pageNumber="99">Figures 1 to 9</emphasis>
Adults.
<emphasis id="BDA79D1AFFE1C90EFEAFA402B2E51215" bold="true" box="[376,406,1869,1890]" pageId="2" pageNumber="99">1 </emphasis>
<taxonomicName id="48D33A8BFFE1C90EFE4AA403B1221215" authority="Zilli &amp; László &amp; Kingston &amp; Larsen, 2023" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[413,593,1868,1890]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="paludicola" status="sp. nov.">
<emphasis id="BDA79D1AFFE1C90EFE4AA403B1221215" box="[413,593,1868,1890]" italics="true" pageId="2" pageNumber="99">Nyodes paludicola</emphasis>
</taxonomicName>
<emphasis id="BDA79D1AFFE1C90EFD8FA402B1D51215" bold="true" box="[600,678,1869,1890]" pageId="2" pageNumber="99">
<taxonomicNameLabel id="A6942061FFE1C90EFD8FA402B1D51215" box="[600,678,1869,1890]" pageId="2" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, HT, ♂, Republic of Congo (ANHRT).
<emphasis id="BDA79D1AFFE1C90EFBFBA402B7381215" bold="true" box="[1068,1099,1869,1890]" pageId="2" pageNumber="99">2 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFB85A403B7F11215" box="[1106,1154,1868,1890]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♂, Liberia (ANHRT).
<emphasis id="BDA79D1AFFE1C90EFA46A402B6C31215" bold="true" box="[1425,1456,1869,1890]" pageId="2" pageNumber="99">3 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFF53A427B3C71209" box="[132,180,1896,1918]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♂, Cameroon (RMCA).
<emphasis id="BDA79D1AFFE1C90EFE02A427B280120A" bold="true" box="[469,499,1896,1917]" pageId="2" pageNumber="99">4 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFE2DA427B1591209" box="[506,554,1896,1918]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♂, Zambia (ANHRT).
<emphasis id="BDA79D1AFFE1C90EFCECA427B029120A" bold="true" box="[827,858,1896,1917]" pageId="2" pageNumber="99">5 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFCB6A427B0E21209" box="[865,913,1896,1918]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♂, Kenya (RCAK).
<emphasis id="BDA79D1AFFE1C90EFB50A427B7D5120A" bold="true" box="[1159,1190,1896,1917]" pageId="2" pageNumber="99">6 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFB7BA427B7AF1209" box="[1196,1244,1896,1918]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♂, Uganda, gen. slide No. NHMUK 010318218 (prep. by A. Zilli) (NHMUK).
<emphasis id="BDA79D1AFFE1C90EFD0AA4CBB18812EE" bold="true" box="[733,763,1924,1945]" pageId="2" pageNumber="99">7 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFCD6A4CCB04212EE" box="[769,817,1923,1945]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♀, DRC (RMCA).
<emphasis id="BDA79D1AFFE1C90EFBC9A4CBB74F12EE" bold="true" box="[1054,1084,1924,1945]" pageId="2" pageNumber="99">8 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFB94A4CCB70012EE" box="[1091,1139,1923,1945]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♀, Sierra Leone (ANHRT).
<emphasis id="BDA79D1AFFE1C90EFF53A4EFB3D112C2" bold="true" box="[132,162,1952,1973]" pageId="2" pageNumber="99">9 </emphasis>
<emphasis id="BDA79D1AFFE1C90EFF7FA4D0B3AB12C2" box="[168,216,1951,1973]" italics="true" pageId="2" pageNumber="99">Idem</emphasis>
, PT, ♀, Zambia (ANHRT).
</paragraph>
</caption>
<caption id="DBAC1180FFE0C90FFF65A1B2B6F11664" ID-DOI="http://doi.org/10.5281/zenodo.14119512" ID-Zenodo-Dep="14119512" box="[178,1410,765,787]" httpUri="https://zenodo.org/record/14119512/files/figure.png" pageId="3" pageNumber="100" startId="3.[178,255,765,786]" targetBox="[240,1348,158,726]" targetPageId="3" targetType="figure">
<paragraph id="8F6C4108FFE0C90FFF65A1B2B6F11664" blockId="3.[178,1410,765,787]" box="[178,1410,765,787]" pageId="3" pageNumber="100">
<emphasis id="BDA79D1AFFE0C90FFF65A1B2B2271665" bold="true" box="[178,340,765,786]" pageId="3" pageNumber="100">Figures 1011 </emphasis>
Adults.
<emphasis id="BDA79D1AFFE0C90FFE72A1B2B2A31665" bold="true" box="[421,464,765,786]" pageId="3" pageNumber="100">10 </emphasis>
<taxonomicName id="48D33A8BFFE0C90FFE01A1B2B1FD1664" authority=", HT" authorityName="HT" box="[470,654,765,787]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE0C90FFE01A1B2B12C1664" box="[470,607,765,787]" italics="true" pageId="3" pageNumber="100">Nyodes steelei</emphasis>
, HT
</taxonomicName>
, ♂, Cameroon (NHMUK).
<emphasis id="BDA79D1AFFE0C90FFC41A1B2B0B21665" bold="true" box="[918,961,765,786]" pageId="3" pageNumber="100">11 </emphasis>
<taxonomicName id="48D33A8BFFE0C90FFC10A1B1B7F81664" authority=", HT" authorityName="HT" box="[967,1163,765,787]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="punctatoides">
<emphasis id="BDA79D1AFFE0C90FFC10A1B1B72E1664" box="[967,1117,765,787]" italics="true" pageId="3" pageNumber="100">N. punctatoides</emphasis>
, HT
</taxonomicName>
, ♂, Tanzania (NHMUK).
</paragraph>
</caption>
<paragraph id="8F6C4108FFE0C90FFF53A07DB23911AF" blockId="3.[132,756,818,2130]" pageId="3" pageNumber="100">Head: Small, positioned quite ventral; frons flat, roughly scaled such as vertex; eye globular, hairy; antenna bipectinate, with finely ciliated rami ending with conspicuous bristle, and as long as width of flagellum in most basal flagellomeres, then gradually lengthened up to 1.8x the width of flagellum in the medial section of antenna, and eventually shortened so as to leave a short filiform apical flagellar section, flagellomeres also with prominent midventral conical process terminated by short seta, and covered with pale greyish brown scales dorsally; proboscis short, likely unfunctional; labial palpus semiporrect, short and compactly clothed dorsally and at sides, roughly so ventrally. Collum with thin dark blackish brown vestiture of scales.</paragraph>
<paragraph id="8F6C4108FFE0C90FFF53A790B7EF137C" blockId="3.[132,756,818,2130]" lastBlockId="3.[832,1455,818,2130]" pageId="3" pageNumber="100">Thorax: Dorsally very gibbous, roughly scaled, mostly unicolorous or with some paler terminal lining of patagia and tegulae; forewing short and compact, distally squared off with nearly vertical termen, and straight costal and anal margins, apex rounded, tornal angle broadly rounded; basal field generally paler than median one, but in some specimens costal area of median field may be similarly pale; subbasal line thin, dark brown, as two short strokes separated by Cubitus; antemedial line white followed by brown, smoothly convex or more often sigmoid, more or less shallowly concave in discal cell and smoothly convex below this, inwardly curved at anal margin; orbicular stigma most often absent, or alternatively very narrow and sub-horizontal, white circled by dark brown, reniform stigma variable also, as a thin vertical white mark lined externally by dark brown or reduced to only a narrow dark brown vertical stroke, distal half of section of CuA2 crossing median field conspicuously lined by bright white stroke; median field overall only slightly constricted below discal cell; postmedial line dark brown, occasionally indistinct against particularly dark unicolorous median field, lined externally by white, acutely pointed outwardly below costa, then almost straight vertical, and shallowly concave below CuA1, often poorly expressed just below discal cell so as that colouring of median field merges with that of distal field, but highlighted bright white in the section from CuA2 to anal margin; distal field concolorous or slightly darker than median one, with decreasing intensity of dark tinge towards apical area, crossed inferiorly by bright white stroke lining CuA2, such stroke combining with white one of median field and white lining of postmedial line to produce in tornal area conspicuous mark in shape of letter “y”, the triangular space between the strokes of the “v- section” of the y- mark showing admixture of white and brown scales; submarginal line dark brown lined externally by white, smooth or tinily irregularly waved, with shallow inward indentation in correspondence of M2, and indistinct below white subtornal y-mark; adterminal field narrow and of variable colouration, from paler to darker than median field; terminal line little distinct, dark brown; fringe greyish brown to reddish brown. Hindwing rounded, pale beige or purplish tinged with extensive suffusion of grey scales, especially in distal area, pattern elements either indistinct or well expressed, if so consisting of grey discal dot or lunule and postmedial line, the latter rounded, very proximally positioned with respect to discal mark, and faded in anal area; termen and fringe as in forewing. underside of wings unicolorous, paler than upperside with some silky shine, showing grey discal spots and diffuse postmedial lines on both wings discernible. Legs with femora and tibiae densely covered in long, dark to reddish brown hairs, spur formula 0-2-4.</paragraph>
<paragraph id="8F6C4108FFE0C90FFC97A55DB731135D" blockId="3.[832,1455,818,2130]" box="[832,1090,1554,1578]" pageId="3" pageNumber="100">Abdomen: Unicolorous.</paragraph>
<paragraph id="8F6C4108FFE0C908FC97A57FB25F14B5" blockId="3.[832,1455,818,2130]" lastBlockId="4.[132,756,151,450]" lastPageId="4" lastPageNumber="101" pageId="3" pageNumber="100">Male genitalia (Figs. 1217): Tegumen short, domeshaped, posteriorly with moderately dilated, finely scobinated arms; vinculum arms ca. 1.2x longer than tegumen, moderately broad, V-oriented, saccus well prominent, comparatively narrow, ca. 1/3 the length of vinculum arms, apically pointed; valva broadly oval before cucullus, with costal margin slightly concave at base, and ventral one evenly arched; sacculus wide and arched, joined anteriorly with outstanding large and soft, strongly setose conico-triangular lobe, of possible clavar derivation, that occupies most of central area of valva; clasper narrow, rather long and arcuate; ampulla in shape of short erect papilla with apical bundle of long robust setae; cucullus with short and narrow neck, distally trilobate, dilated into ventral triangular plate bearing middorsally a fin-like plate, the former edged by corona of short acute spines; costal prolongations on diaphragma relatively wide, ribbon-like, not co-joined into unique transtilla; juxta very large, broadly cordiform; uncus flat and relatively short, with almost parallel or slightly concave sides, quite variable in width between samples, and with truncated apex often showing feebly concave tip, covered in both dorsal and ventral surfaces with sparse short hairs; tuba analis wide and membranous; phallus shaft elongated and of approximately same width, regularly arched dorsally, with short and rounded coecum; vesica long, slightly rugose, with short basal swelling and gradually tapered main tube, lacking cornuti.</paragraph>
<paragraph id="8F6C4108FFE7C908FC97A3D8B74015BA" blockId="4.[832,1455,151,205]" pageId="4" pageNumber="101">more fuscous hindwing, and less hairy legs. Colouration can be slightly plainer.</paragraph>
<caption id="DBAC1180FFE7C908FC97A186B7AF163A" ID-DOI="http://doi.org/10.5281/zenodo.14119516" ID-Zenodo-Dep="14119516" httpUri="https://zenodo.org/record/14119516/files/figure.png" pageId="4" pageNumber="101" startId="4.[832,909,713,734]" targetBox="[835,1459,234,678]" targetPageId="4" targetType="figure">
<paragraph id="8F6C4108FFE7C908FC97A186B7AF163A" blockId="4.[832,1456,712,845]" pageId="4" pageNumber="101">
<emphasis id="BDA79D1AFFE7C908FC97A186B09517A9" bold="true" box="[832,998,713,734]" pageId="4" pageNumber="101">Figures 2022 </emphasis>
Female genitalia (
<emphasis id="BDA79D1AFFE7C908FB76A186B7BD17A9" bold="true" box="[1185,1230,713,734]" pageId="4" pageNumber="101">20 </emphasis>
ventral view,
<emphasis id="BDA79D1AFFE7C908FA88A186B6C317A9" bold="true" box="[1375,1456,713,734]" pageId="4" pageNumber="101">2122 </emphasis>
dorsal view).
<emphasis id="BDA79D1AFFE7C908FC11A1ABB081178E" bold="true" box="[966,1010,740,761]" pageId="4" pageNumber="101">20 </emphasis>
<taxonomicName id="48D33A8BFFE7C908FC2DA1ABB7DD178D" authority="Zilli &amp; László &amp; Kingston &amp; Larsen, 2023" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[1018,1198,740,762]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="paludicola" status="sp. nov.">
<emphasis id="BDA79D1AFFE7C908FC2DA1ABB7DD178D" box="[1018,1198,740,762]" italics="true" pageId="4" pageNumber="101">Nyodes paludicola</emphasis>
</taxonomicName>
<emphasis id="BDA79D1AFFE7C908FB61A1ABB676178E" bold="true" box="[1206,1285,740,761]" pageId="4" pageNumber="101">
<taxonomicNameLabel id="A6942061FFE7C908FB61A1ABB676178E" box="[1206,1285,740,761]" pageId="4" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, PT, Republic of Congo, gen. slide No.: LG 6049 (ANHRT).
<emphasis id="BDA79D1AFFE7C908FB33A04FB67D1662" bold="true" box="[1252,1294,768,789]" pageId="4" pageNumber="101">21 </emphasis>
<emphasis id="BDA79D1AFFE7C908FAC4A1B0B6301662" box="[1299,1347,767,789]" italics="true" pageId="4" pageNumber="101">Idem</emphasis>
, PT, Sierra Leone, gen. slide No.: LG 6055 (ANHRT).
<emphasis id="BDA79D1AFFE7C908FAD8A054B6331647" bold="true" box="[1295,1344,795,816]" pageId="4" pageNumber="101">22 </emphasis>
<emphasis id="BDA79D1AFFE7C908FA9BA054B60F1646" box="[1356,1404,795,817]" italics="true" pageId="4" pageNumber="101">Idem</emphasis>
, PT, Zambia, gen. slide No.: LG 5748 (RCAK).
</paragraph>
</caption>
<caption id="DBAC1180FFE7C908FF53A660B1E41371" ID-DOI="http://doi.org/10.5281/zenodo.14119518" ID-Zenodo-Dep="14119518" httpUri="https://zenodo.org/record/14119518/files/figure.png" pageId="4" pageNumber="101" startId="4.[132,209,1327,1348]" targetBox="[136,760,479,1287]" targetPageId="4" targetType="figure">
<paragraph id="8F6C4108FFE7C908FF53A660B1E41371" blockId="4.[132,757,1326,1542]" pageId="4" pageNumber="101">
<emphasis id="BDA79D1AFFE7C908FF53A660B2571033" bold="true" box="[132,292,1327,1348]" pageId="4" pageNumber="101">Figures 1217 </emphasis>
Male genitalia, a: clasping apparatus, b: phallus.
<emphasis id="BDA79D1AFFE7C908FF53A605B3C21028" bold="true" box="[132,177,1354,1375]" pageId="4" pageNumber="101">12 </emphasis>
<taxonomicName id="48D33A8BFFE7C908FF6EA605B21D1017" authority="Zilli &amp; László &amp; Kingston &amp; Larsen, 2023" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[185,366,1354,1376]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="paludicola" status="sp. nov.">
<emphasis id="BDA79D1AFFE7C908FF6EA605B21D1017" box="[185,366,1354,1376]" italics="true" pageId="4" pageNumber="101">Nyodes paludicola</emphasis>
</taxonomicName>
<emphasis id="BDA79D1AFFE7C908FEA1A605B2B51028" bold="true" box="[374,454,1354,1375]" pageId="4" pageNumber="101">
<taxonomicNameLabel id="A6942061FFE7C908FEA1A605B2B51028" box="[374,454,1354,1375]" pageId="4" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, HT, Republic of Congo, gen. slide No.: LG 6048 (ANHRT).
<emphasis id="BDA79D1AFFE7C908FE6AA629B29A100C" bold="true" box="[445,489,1382,1403]" pageId="4" pageNumber="101">13 </emphasis>
<emphasis id="BDA79D1AFFE7C908FE26A629B152100B" box="[497,545,1382,1404]" italics="true" pageId="4" pageNumber="101">Idem</emphasis>
, PT, Cameroon, gen. slide No.: LG 5899 (RMCA);
<emphasis id="BDA79D1AFFE7C908FE65A6CDB2AD10E0" bold="true" box="[434,478,1410,1431]" pageId="4" pageNumber="101">14 </emphasis>
PT, Liberia, gen. slide No.: LG 6054 (ANHRT).
<emphasis id="BDA79D1AFFE7C908FE86A6D2B20E10C5" bold="true" box="[337,381,1437,1458]" pageId="4" pageNumber="101">15 </emphasis>
<emphasis id="BDA79D1AFFE7C908FE53A6D2B2C710C4" box="[388,436,1437,1459]" italics="true" pageId="4" pageNumber="101">Idem</emphasis>
, PT, Zambia, gen. slide No.: LG 5780 (ANHRT).
<emphasis id="BDA79D1AFFE7C908FEFEA6F6B22710B9" bold="true" box="[297,340,1465,1486]" pageId="4" pageNumber="101">16 </emphasis>
<emphasis id="BDA79D1AFFE7C908FE8CA6F7B2F810B9" box="[347,395,1464,1486]" italics="true" pageId="4" pageNumber="101">Idem</emphasis>
, PT, Kenya, gen. slide No.: LG 5747 (RCAK);
<emphasis id="BDA79D1AFFE7C908FF21A69BB247109E" bold="true" box="[246,308,1492,1513]" pageId="4" pageNumber="101">17 </emphasis>
<emphasis id="BDA79D1AFFE7C908FE9AA69BB20E109D" box="[333,381,1492,1514]" italics="true" pageId="4" pageNumber="101">Idem</emphasis>
, PT, Uganda, gen. slide No.: NHMUK010318218 (prepared by A. Zilli) (NHMUK).
</paragraph>
</caption>
<caption id="DBAC1180FFE7C908FF53A41BB29C12CA" ID-DOI="http://doi.org/10.5281/zenodo.14119520" ID-Zenodo-Dep="14119520" httpUri="https://zenodo.org/record/14119520/files/figure.png" pageId="4" pageNumber="101" startId="4.[132,209,1876,1897]" targetBox="[136,760,1570,1840]" targetPageId="4" targetType="figure">
<paragraph id="8F6C4108FFE7C908FF53A41BB29C12CA" blockId="4.[132,757,1876,1981]" pageId="4" pageNumber="101">
<emphasis id="BDA79D1AFFE7C908FF53A41BB257121E" bold="true" box="[132,292,1876,1897]" pageId="4" pageNumber="101">Figures 1819 </emphasis>
Male genitalia, a: clasping apparatus, b: phallus.
<emphasis id="BDA79D1AFFE7C908FF53A43FB3C112F2" bold="true" box="[132,178,1904,1925]" pageId="4" pageNumber="101">18 </emphasis>
<taxonomicName id="48D33A8BFFE7C908FF6BA43EB23612F1" authority=", PT" authorityName="PT" box="[188,325,1904,1926]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE7C908FF6BA43EB26B12F1" box="[188,280,1904,1926]" italics="true" pageId="4" pageNumber="101">N. steelei</emphasis>
, PT
</taxonomicName>
, Cameroon, gen. slide No.: BM
<taxonomicName id="48D33A8BFFE7C908FD46A43FB18012F1" box="[657,755,1904,1926]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="family">Noctuidae</taxonomicName>
5803 (NHMUK).
<emphasis id="BDA79D1AFFE7C908FEE5A4C3B22E12D6" bold="true" box="[306,349,1932,1953]" pageId="4" pageNumber="101">19 </emphasis>
<taxonomicName id="48D33A8BFFE7C908FEB4A4C3B15A12D6" authority=", HT" authorityName="HT" box="[355,553,1931,1953]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="punctatoides">
<emphasis id="BDA79D1AFFE7C908FEB4A4C3B28912D6" box="[355,506,1931,1953]" italics="true" pageId="4" pageNumber="101">N. punctatoides</emphasis>
, HT
</taxonomicName>
, Tanzania, gen. slide No.: BM
<taxonomicName id="48D33A8BFFE7C908FF37A4E8B23012CA" box="[224,323,1959,1981]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="family">Noctuidae</taxonomicName>
5805 (NHMUK).
</paragraph>
</caption>
<paragraph id="8F6C4108FFE7C908FF53A493B2261283" blockId="4.[132,756,2012,2128]" box="[132,341,2012,2036]" pageId="4" pageNumber="101">Female (Figs. 79):</paragraph>
<paragraph id="8F6C4108FFE7C908FF53A4B4B01E1038" blockId="4.[132,756,2012,2128]" lastBlockId="4.[832,1455,875,1359]" pageId="4" pageNumber="101">Size, habitus and pattern as in male, with smaller head (ca. 20% less in diameter), labial palpi being particularly smaller, and with filiform, finely ciliated antennae, overall Female genitalia (Figs. 2022): Segment A8 short and broad, narrower dorsally; apophysis anterior short and thin; ostium bursae wide and shallow cup-like, lightly sclerotised; ductus bursae comparatively long and wide, ca. as long as apophysis posterior, narrowest posteriorly and gradually dilated towards corpus bursae, with numerous strongly sclerotised ridges; corpus bursae wholly membranous, posteriorly elongated ovoid in shape, and anteriorly strongly bent to left into large, caudally oriented sack-like fundus; appendix bursae conspicuous with very broad basal section branching before terminal end of fundus bursae; ovipositor short, rather blunt, with short intersegmental membrane A8-A9 and large papillae anales; papilla trapezoid in side view, sparsely setose; apophysis posterior rod-like, ca. 4x longer than anterior one.</paragraph>
</subSubSection>
<subSubSection id="C7C91283FFE7C909FC97A620B7DA16F8" lastPageId="5" lastPageNumber="102" pageId="4" pageNumber="101" type="diagnosis">
<paragraph id="8F6C4108FFE7C908FC97A620B0DE10F0" blockId="4.[832,1456,1391,2120]" box="[832,941,1391,1415]" pageId="4" pageNumber="101">
<emphasis id="BDA79D1AFFE7C908FC97A620B0DE10F0" bold="true" box="[832,941,1391,1415]" pageId="4" pageNumber="101">Diagnosis</emphasis>
</paragraph>
<paragraph id="8F6C4108FFE7C909FC97A6C2B2D816DA" blockId="4.[832,1456,1391,2120]" lastBlockId="5.[132,756,151,941]" lastPageId="5" lastPageNumber="102" pageId="4" pageNumber="101">
The new species is readily distinguished from all other
<taxonomicName id="48D33A8BFFE7C908FC97A6E3B0FD10B3" authorityName="Laporte" authorityYear="1970" box="[832,910,1452,1476]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="BDA79D1AFFE7C908FC97A6E3B0FD10B3" box="[832,910,1452,1476]" italics="true" pageId="4" pageNumber="101">Nyodes</emphasis>
</taxonomicName>
by its considerably less elongate and apically less pointed forewing and the chocolate brown to rusty brown colouration with some purplish tinge lacking any mossy green mottling, which is typical for the majority of
<taxonomicName id="48D33A8BFFE7C908FAB6A547B6DC1357" authorityName="Laporte" authorityYear="1970" box="[1377,1455,1544,1568]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="BDA79D1AFFE7C908FAB6A547B6DC1357" box="[1377,1455,1544,1568]" italics="true" pageId="4" pageNumber="101">Nyodes</emphasis>
</taxonomicName>
taxa. The only taxon which is somewhat externally reminiscent of the new species is
<taxonomicName id="48D33A8BFFE7C908FB7BA509B6DC132A" authority="Laporte, 1971" authorityName="Laporte" authorityYear="1971" box="[1196,1455,1605,1629]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE7C908FB7BA509B662132A" box="[1196,1297,1605,1629]" italics="true" pageId="4" pageNumber="101">N. steelei</emphasis>
Laporte, 1971
</taxonomicName>
(Fig. 10), due to its predominantly dark brown colouration, although it shows some greenish tinge on the pale markings of forewing. A further similarity between the two species consists of a conspicuous pale sub-horizontal streak starting from the median field of forewing that upon crossing the postmedial line bends posteriorly and widens into a triangular mark ending at tornus. However, in the new species this pattern element starts as a short streak only from the middle of the median field and it wholly joins the triangular mark so as to produce a sharply defined, white coloured y, the space in- between the two strokes (“v”) of such y-mark being darker with an admixture of brown and white scales (to be noted that the most posterior of the two strokes coincides with the lower section of the postmedial line). In
<taxonomicName id="48D33A8BFFE7C908FC4EAB5DB0881D5E" authorityName="Laporte" authorityYear="1971" box="[921,1019,2065,2089]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE7C908FC4EAB5DB0881D5E" box="[921,1019,2065,2089]" italics="true" pageId="4" pageNumber="101">N. steelei</emphasis>
</taxonomicName>
, the median streak is a thicker and longer greenish beige stroke that crosses almost wholly the median field, and it is thinly albeit distinctly interrupted by the postmedial line, while the tornal triangular mark appears more as an entirely uniform greenish beige stroke. Notably, the new species also has distinctly bipectinate male antennae contrary to filiform ones of
<taxonomicName id="48D33A8BFFE6C909FD55A25DB19E145E" authorityName="Laporte" authorityYear="1971" box="[642,749,273,297]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE6C909FD55A25DB19E145E" box="[642,749,273,297]" italics="true" pageId="5" pageNumber="102">N. steelei</emphasis>
</taxonomicName>
, forewing more squared off with almost vertical termen, white instead of greenish beige pattern elements, smaller stigmata, the orbicular being usually obliterated and the reniform occasionally reduced to thin dark brown dash only, shallower concavity of the inferior section of the postmedial line, and a considerably paler brown hindwing compared to
<taxonomicName id="48D33A8BFFE6C909FED8A2A6B2021777" authorityName="Laporte" authorityYear="1971" box="[271,369,488,512]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE6C909FED8A2A6B2021777" box="[271,369,488,512]" italics="true" pageId="5" pageNumber="102">N. steelei</emphasis>
</taxonomicName>
. In the male genitalia (Figs. 1218), the new species greatly differs from
<taxonomicName id="48D33A8BFFE6C909FDC8A147B1F51768" authorityName="Laporte" authorityYear="1971" box="[543,646,519,543]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE6C909FDC8A147B1F51768" box="[543,646,519,543]" italics="true" pageId="5" pageNumber="102">N. steelei</emphasis>
</taxonomicName>
, although both of them show an almost parallel-sided and apically truncated uncus. Main differences are to be found in the valval outline, broadly oval in the new species excepting for the cucullus, a massive, soft and sparsely setose lobe of likely clavar origin occupying the middle of valva, the arched clasper without free apical process, a short albeit prominent, papilla-like ampulla with conspicuous bundle of setae, and the narrow-necked albeit apically conspicuously dilated and trilobate cucullus, while this is gradually tapered with sharp apex in
<taxonomicName id="48D33A8BFFE6C909FDC6A075B1061626" authorityName="Laporte" authorityYear="1971" box="[529,629,825,849]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE6C909FDC6A075B1061626" box="[529,629,825,849]" italics="true" pageId="5" pageNumber="102">N. steelei</emphasis>
</taxonomicName>
. In the new species, the phallus is longer and slenderer, without cornuti, whereas the vesica of
<taxonomicName id="48D33A8BFFE6C909FE04A037B14A16F8" authorityName="Laporte" authorityYear="1971" box="[467,569,887,911]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE6C909FE04A037B14A16F8" box="[467,569,887,911]" italics="true" pageId="5" pageNumber="102">N. steelei</emphasis>
</taxonomicName>
bears a group of short, thorn-shaped cornuti.
</paragraph>
<paragraph id="8F6C4108FFE6C909FF53A083B2581275" blockId="5.[132,756,972,1794]" pageId="5" pageNumber="102">
A somewhat externally similar species to
<taxonomicName id="48D33A8BFFE6C909FD8CA082B1B11693" authorityName="Laporte" authorityYear="1971" box="[603,706,972,996]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE6C909FD8CA082B1B11693" box="[603,706,972,996]" italics="true" pageId="5" pageNumber="102">N. steelei</emphasis>
</taxonomicName>
and thence to the new species is
<taxonomicName id="48D33A8BFFE6C909FE66A0A3B1871174" authority="Laporte, 1973" authorityName="Laporte" authorityYear="1973" box="[433,756,1003,1027]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="punctatoides">
<emphasis id="BDA79D1AFFE6C909FE66A0A3B12A1174" box="[433,601,1003,1027]" italics="true" pageId="5" pageNumber="102">N. punctatoides</emphasis>
Laporte, 1973
</taxonomicName>
(Fig. 11), which however has even wider and more intense green markings than
<taxonomicName id="48D33A8BFFE6C909FEB6A765B2B11136" authorityName="Laporte" authorityYear="1971" box="[353,450,1065,1089]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="steelei">
<emphasis id="BDA79D1AFFE6C909FEB6A765B2B11136" box="[353,450,1065,1089]" italics="true" pageId="5" pageNumber="102">N. steelei</emphasis>
</taxonomicName>
itself. Interestingly, together with species like
<taxonomicName id="48D33A8BFFE6C909FE53A707B1831128" authority="Hampson, 1918" authorityName="Hampson" authorityYear="1918" box="[388,752,1095,1119]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="rufifusa">
<emphasis id="BDA79D1AFFE6C909FE53A707B17D1128" box="[388,526,1095,1119]" italics="true" pageId="5" pageNumber="102">N. rufifusa</emphasis>
Hampson, 1918
</taxonomicName>
,
<taxonomicName id="48D33A8BFFE6C909FF53A728B2B51109" authority="Laporte, 1973" authorityName="Laporte" authorityYear="1973" box="[132,454,1126,1150]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="rufifusoides">
<emphasis id="BDA79D1AFFE6C909FF53A728B2521109" box="[132,289,1126,1150]" italics="true" pageId="5" pageNumber="102">N. rufifusoides</emphasis>
Laporte, 1973
</taxonomicName>
,
<taxonomicName id="48D33A8BFFE6C909FE03A728B19C1109" authority="Laporte, 1973" authorityName="Laporte" authorityYear="1973" box="[468,751,1126,1150]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="jucunda">
<emphasis id="BDA79D1AFFE6C909FE03A728B13F1109" box="[468,588,1126,1150]" italics="true" pageId="5" pageNumber="102">N. jucunda</emphasis>
Laporte, 1973
</taxonomicName>
, and
<taxonomicName id="48D33A8BFFE6C909FF61A7CAB29F11EB" authority="Laporte, 1977" authorityName="Laporte" authorityYear="1977" box="[182,492,1156,1180]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="nigrioides">
<emphasis id="BDA79D1AFFE6C909FF61A7CAB23611EB" box="[182,325,1156,1180]" italics="true" pageId="5" pageNumber="102">N. nigrioides</emphasis>
Laporte, 1977
</taxonomicName>
,
<taxonomicName id="48D33A8BFFE6C909FE2BA7CAB1D411EB" authorityName="Laporte" authorityYear="1973" box="[508,679,1156,1180]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="punctatoides">
<emphasis id="BDA79D1AFFE6C909FE2BA7CAB1D411EB" box="[508,679,1156,1180]" italics="true" pageId="5" pageNumber="102">N. punctatoides</emphasis>
</taxonomicName>
shares with the new species the same configuration of the uncus and multi-lobate cucullus, but in all such species the cucullus is shorter-necked, its distal processes are differently oriented, shorter and generally more sharply tipped, often with a bundle of spines, so that the cuculli are overall tapered. Possibly, the closest species to the new one in the configuration of the male genitalia is really
<taxonomicName id="48D33A8BFFE6C909FF53A634B25F10E5" authorityName="Laporte" authorityYear="1973" box="[132,300,1402,1426]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="punctatoides">
<emphasis id="BDA79D1AFFE6C909FF53A634B25F10E5" box="[132,300,1402,1426]" italics="true" pageId="5" pageNumber="102">N. punctatoides</emphasis>
</taxonomicName>
(Fig. 19), as the juxta is similarly broadly cordiform, the ventral plate of the cucullus is distally dilated (albeit lesser), it is edged by a row of spines, and dorsally there is another acutely pointed process. This however well separate from the ventral plate, and the cucullus is also reached by a further hook-like process consisting of the distal free termination of a long straight clasper proceeding from the valvula. Further to this, the cucullus of
<taxonomicName id="48D33A8BFFE6C909FED7A53FB2D413F0" authorityName="Laporte" authorityYear="1973" box="[256,423,1647,1671]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="punctatoides">
<emphasis id="BDA79D1AFFE6C909FED7A53FB2D413F0" box="[256,423,1647,1671]" italics="true" pageId="5" pageNumber="102">N. punctatoides</emphasis>
</taxonomicName>
has no narrow stalk and is thus shorter and more broadly joined to the valvula, there is no voluminous setose lobe on the middle of valva, the uncus is much wider, and the phallus bears a series of small conical cornuti.
</paragraph>
<caption id="DBAC1180FFE6C909FC97A6F7B6D210BA" ID-DOI="http://doi.org/10.5281/zenodo.14119524" ID-Zenodo-Dep="14119524" box="[832,1441,1464,1486]" httpUri="https://zenodo.org/record/14119524/files/figure.png" pageId="5" pageNumber="102" startId="5.[832,900,1464,1485]" targetBox="[835,1459,1117,1429]" targetPageId="5" targetType="figure">
<paragraph id="8F6C4108FFE6C909FC97A6F7B6D210BA" blockId="5.[832,1441,1464,1486]" box="[832,1441,1464,1486]" pageId="5" pageNumber="102">
<emphasis id="BDA79D1AFFE6C909FC97A6F7B0C710BA" bold="true" box="[832,948,1464,1485]" pageId="5" pageNumber="102">Figure 23 </emphasis>
Distribution records of
<taxonomicName id="48D33A8BFFE6C909FB4DA6F7B63D10B9" authority="Zilli &amp; László &amp; Kingston &amp; Larsen, 2023" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[1178,1358,1464,1486]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="paludicola" status="sp. nov.">
<emphasis id="BDA79D1AFFE6C909FB4DA6F7B63D10B9" box="[1178,1358,1464,1486]" italics="true" pageId="5" pageNumber="102">Nyodes paludicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6942061FFE6C909FA83A6F7B6D210BA" box="[1364,1441,1464,1485]" pageId="5" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8F6C4108FFE6C909FF53A46EB22E124E" blockId="5.[132,756,1825,2125]" box="[132,349,1825,1849]" pageId="5" pageNumber="102">
<emphasis id="BDA79D1AFFE6C909FF53A46EB22E124E" bold="true" box="[132,349,1825,1849]" pageId="5" pageNumber="102">Genetic divergence</emphasis>
</paragraph>
<paragraph id="8F6C4108FFE6C909FF53A40FB7DA16F8" blockId="5.[132,756,1825,2125]" lastBlockId="5.[832,1456,151,911]" pageId="5" pageNumber="102">
The intraspecific divergence of the COI mtDNA barcodes of
<taxonomicName id="48D33A8BFFE6C909FF48A410B25C1201" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[159,303,1886,1910]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="BDA79D1AFFE6C909FF48A410B25C1201" box="[159,303,1886,1910]" italics="true" pageId="5" pageNumber="102">N. paludicola</emphasis>
</taxonomicName>
was found in the range of 0.00-2.50%. The genetic divergence within single populations seems to be null, in fact it is 0.00% in the Zambian population (n = 4), but it already increases at low geographic distance, e.g. 0.46% when
<specimenCount id="99D58A81FFE6C909FEDAA495B2ED1286" box="[269,414,2009,2033]" count="1" pageId="5" pageNumber="102" type="generic">one specimen</specimenCount>
from
<collectingCountry id="F7C40198FFE6C909FE0CA496B1111286" box="[475,610,2009,2033]" name="Sierra Leone" pageId="5" pageNumber="102">Sierra Leone</collectingCountry>
and one from
<collectingCountry id="F7C40198FFE6C909FF53A4B7B3A21D67" box="[132,209,2040,2064]" name="Liberia" pageId="5" pageNumber="102">Liberia</collectingCountry>
are compared. The pairwise distance between more separate populations is higher, attaining 2.30-2.50% between the Zambian and West African (n = 2) samples, 1.23% between the Zambian and Kenyan (n = 1) ones, and 1.68-2.00% between those from
<collectingCountry id="F7C40198FFE6C909FB71A3F9B79815B9" box="[1190,1259,182,206]" name="Kenya" pageId="5" pageNumber="102">Kenya</collectingCountry>
and West Africa. Insights into the substitutions occurring in the COI barcodes by these populations however fail to recover full congruence with geographic provenance of the specimens. For instance, two out of the three mutations that separate the Liberian specimen from that of
<collectingCountry id="F7C40198FFE6C909FB07A200B62F1411" box="[1232,1372,335,359]" name="Sierra Leone" pageId="5" pageNumber="102">Sierra Leone</collectingCountry>
do not occur in the southeastern African samples, or six out of the eight nucleotides that make the Zambian samples differ from the Kenyan one are unmodified in the West African specimens, while the remaining two differ in the Kenyan specimen and not in the West African + Zambian lot. This overall situation is suggestive of a normal pattern of genetic divergence by distance (cf.
<bibRefCitation id="EB423CF9FFE6C909FB2BA169B6F0174A" author="MAYR, E." box="[1276,1411,549,574]" pageId="5" pageNumber="102" refId="ref6565" refString="MAYR, E. 1963. Animal Species and Evolution. The Belknap Press of Harvard University Press, Cambridge MS. xiv + 797 pp." type="book" year="1963">Mayr 1963</bibRefCitation>
), a circumstance that should always be thoroughly checked with careful analysis of geographically interposed populations when taxonomic conclusions are drawn from barcode data on allopatric populations, and is likely enhanced by the scattered distribution of the swamp-like environments where
<taxonomicName id="48D33A8BFFE6C909FBF6A191B7C01782" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[1057,1203,733,757]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="BDA79D1AFFE6C909FBF6A191B7C01782" box="[1057,1203,733,757]" italics="true" pageId="5" pageNumber="102">N. paludicola</emphasis>
</taxonomicName>
occurs. Further to this, as no definite morphological substantiation of any geographic groupings could be found, e.g. in the narrowness of uncus, broadness of valva or vesical diverticula, the samples are here treated as belonging to a single, widely distributed species.
</paragraph>
</subSubSection>
<subSubSection id="C7C91283FFE6C909FC97A0E1B0E81136" pageId="5" pageNumber="102" type="etymology">
<paragraph id="8F6C4108FFE6C909FC97A0E1B0C816B1" blockId="5.[832,1455,942,1089]" box="[832,955,942,966]" pageId="5" pageNumber="102">
<emphasis id="BDA79D1AFFE6C909FC97A0E1B0C816B1" bold="true" box="[832,955,942,966]" pageId="5" pageNumber="102">Etymology</emphasis>
</paragraph>
<paragraph id="8F6C4108FFE6C909FC97A083B0E81136" blockId="5.[832,1455,942,1089]" pageId="5" pageNumber="102">
The name
<taxonomicName id="48D33A8BFFE6C909FC78A083B7521693" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[943,1057,972,996]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="BDA79D1AFFE6C909FC78A083B7521693" box="[943,1057,972,996]" italics="true" pageId="5" pageNumber="102">paludicola</emphasis>
</taxonomicName>
(= swamp-dwelling) attributed to the new species is inspired by its preferences for marshes and other wetlands, as features of all known collecting sites indicate.
</paragraph>
</subSubSection>
<subSubSection id="C7C91283FFE6C909FC97A6A2B72E1240" pageId="5" pageNumber="102" type="distribution">
<paragraph id="8F6C4108FFE6C909FC97A6A2B75C1372" blockId="5.[832,1455,1517,1847]" box="[832,1071,1517,1541]" pageId="5" pageNumber="102">
<emphasis id="BDA79D1AFFE6C909FC97A6A2B0B81372" bold="true" box="[832,971,1517,1541]" pageId="5" pageNumber="102">Distribution</emphasis>
(
<figureCitation id="17E85D8DFFE6C909FC0EA6A2B7541372" box="[985,1063,1517,1541]" captionStart="Figure 23" captionStartId="5.[832,900,1464,1485]" captionTargetBox="[835,1459,1117,1429]" captionTargetId="figure-1255@5.[835,1459,1117,1429]" captionTargetPageId="5" captionText="Figure 23 Distribution records of Nyodes paludicola sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14119524" httpUri="https://zenodo.org/record/14119524/files/figure.png" pageId="5" pageNumber="102">Fig. 23</figureCitation>
)
</paragraph>
<paragraph id="8F6C4108FFE6C909FC97A544B72E1240" blockId="5.[832,1455,1517,1847]" pageId="5" pageNumber="102">
<taxonomicName id="48D33A8BFFE6C909FC97A544B7651354" authorityName="Zilli &amp; László &amp; Kingston &amp; Larsen" authorityYear="2023" box="[832,1046,1547,1571]" class="Insecta" family="Noctuidae" genus="Nyodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="BDA79D1AFFE6C909FC97A544B7651354" box="[832,1046,1547,1571]" italics="true" pageId="5" pageNumber="102">Nyodes paludicola</emphasis>
</taxonomicName>
is widely distributed throughout Equatorial Africa with records from
<collectingCountry id="F7C40198FFE6C909FB12A565B6231336" box="[1221,1360,1578,1602]" name="Sierra Leone" pageId="5" pageNumber="102">Sierra Leone</collectingCountry>
,
<collectingCountry id="F7C40198FFE6C909FA8CA565B6D81335" box="[1371,1451,1578,1602]" name="Liberia" pageId="5" pageNumber="102">Liberia</collectingCountry>
,
<collectingCountry id="F7C40198FFE6C909FC97A507B0C21317" box="[832,945,1608,1632]" name="Cameroon" pageId="5" pageNumber="102">Cameroon</collectingCountry>
,
<collectingCountry id="F7C40198FFE6C909FC17A507B7E41317" box="[960,1175,1608,1632]" name="Republic of the Congo" pageId="5" pageNumber="102">Republic of Congo</collectingCountry>
,
<collectingCountry id="F7C40198FFE6C909FB71A507B0C81308" name="Democratic Republic of the Congo" pageId="5" pageNumber="102">Democratic Republic of the Congo</collectingCountry>
,
<collectingCountry id="F7C40198FFE6C909FC07A528B7551308" box="[976,1062,1639,1663]" name="Uganda" pageId="5" pageNumber="102">Uganda</collectingCountry>
,
<collectingCountry id="F7C40198FFE6C909FBECA527B7F213F7" box="[1083,1153,1640,1664]" name="Kenya" pageId="5" pageNumber="102">Kenya</collectingCountry>
and
<collectingCountry id="F7C40198FFE6C909FB1EA528B66D1308" box="[1225,1310,1639,1663]" name="Zambia" pageId="5" pageNumber="102">Zambia</collectingCountry>
. Despite its extensive range, the species appear to be rather rare and local, confined to wetlands (
<figureCitation id="17E85D8DFFE6C909FBA8A5EAB67B13CA" box="[1151,1288,1701,1725]" captionStart-0="Figure 24" captionStart-1="Figure 25" captionStart-2="Figure 26" captionStartId-0="6.[132,200,651,672]" captionStartId-1="6.[132,200,1259,1280]" captionStartId-2="6.[132,200,1867,1888]" captionTargetBox-0="[136,760,148,616]" captionTargetBox-1="[136,760,756,1224]" captionTargetBox-2="[136,758,1364,1832]" captionTargetId-0="figure-718@6.[136,760,148,616]" captionTargetId-1="figure-721@6.[136,760,756,1224]" captionTargetId-2="figure-724@6.[136,758,1364,1832]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="6" captionText-0="Figure 24 Mbeli Bai (swampy forest clearing) in the Nouabale- Ndoki National Park, Republic of Congo, type locality of Nyodes paludicola sp. nov. (photo: Gy. M. László)." captionText-1="Figure 25 Lukwakwa Dambo (seasonal wetland) in the West Lunga National Park, Zambia, habitat of Nyodes paludicola sp. nov. (photo: Gy. M. László)." captionText-2="Figure 26 Karamaini Estate in Thika District, Kenya, collecting site of Nyodes paludicola sp. nov. (photo: A.J. Kingston)." figureDoi-0="http://doi.org/10.5281/zenodo.14119526" figureDoi-1="http://doi.org/10.5281/zenodo.14119530" figureDoi-2="http://doi.org/10.5281/zenodo.14119534" httpUri-0="https://zenodo.org/record/14119526/files/figure.png" httpUri-1="https://zenodo.org/record/14119530/files/figure.png" httpUri-2="https://zenodo.org/record/14119534/files/figure.png" pageId="5" pageNumber="102">Figs. 2426</figureCitation>
). The fact that marshes and swamps have never been considered popular collecting sites by entomologists in the Afrotropics may have led to the low representation of this peculiar species in
<taxonomicName id="48D33A8BFFE6C909FC8CA450B0AF1240" authorityName="Linnaeus" authorityYear="1758" box="[859,988,1823,1847]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="102" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
collections.
</paragraph>
</subSubSection>
</treatment>
</document>