482 lines
54 KiB
XML
482 lines
54 KiB
XML
<document id="CF7F68E3588BE8124E9FB7351CEFEADF" ID-CLB-Dataset="29242" ID-DOI="10.11646/zootaxa.4496.1.42" ID-GBIF-Dataset="895f6990-113a-4193-a279-7dfbcd8be850" ID-ISSN="1175-5326" ID-Zenodo-Dep="1447025" ID-ZooBank="F0E7B997-D3E2-45F9-9DB8-1CE60196FEC3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538748284745" checkinUser="plazi" docAuthor="Bartz, Marie L. C., James, Samuel W., Steffen, Gerusa P. K., Antoniolli, Zaida, Steffen, Ricardo B. & Brown, George G." docDate="2018" docId="072D87D2FFCBFF8941DFFED2E0F0FD6F" docLanguage="en" docName="zootaxa.4496.1.42.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glossoscolex GlossosColex, n. sp." docType="treatment" docVersion="7" lastPageNumber="552" masterDocId="FB14FFAAFFC9FF8D4148FFEDE35AFFEA" masterDocTitle="New species-group taxa of Glossoscolex (Clitellata: Glossoscolecidae) from Rio Grande do Sul, Brazil" masterLastPageNumber="561" masterPageNumber="548" pageNumber="550" updateTime="1698566898224" updateUser="ExternalLinkService">
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<mods:titleInfo id="6044D47B9A9F4DFF79EC781C1914C4A5">
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<mods:title id="BC076D26EB560AE18AF1D34161D03D0C">New species-group taxa of Glossoscolex (Clitellata: Glossoscolecidae) from Rio Grande do Sul, Brazil</mods:title>
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<mods:name id="C9F3A2BA0C73C7243B8BAAEE05620A0D" type="personal">
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<mods:namePart id="D3EA4A4188682CE22E148599AC2ADC1B">Bartz, Marie L. C.</mods:namePart>
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<mods:namePart id="6B8D85F76CEF07C9603B5F417B9FE167">James, Samuel W.</mods:namePart>
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<mods:namePart id="FF3235B77A98659EE82872662B965231">Steffen, Gerusa P. K.</mods:namePart>
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<mods:namePart id="372156AB5007D18E27BBE9534D74CD4B">Antoniolli, Zaida</mods:namePart>
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<mods:namePart id="EC0FEAD43C51DEDBBCF693B519B80F2F">Steffen, Ricardo B.</mods:namePart>
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<mods:namePart id="A88C4F327773D0CC700F3CDA7616E70D">Brown, George G.</mods:namePart>
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<mods:title id="276A4AE41FD023A5EE734AF7108D1CDF">Zootaxa</mods:title>
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<mods:part id="EB8B05F14FA3561CFDC04DD0BE063610">
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<mods:date id="A38F33BF65930016FC5837A770EECC15">2018</mods:date>
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<mods:number id="4F2557487CA57691D2D427680F83C87A">2018-10-05</mods:number>
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<mods:number id="BCB10E542CAB8E5E6B66FA99215DADF7">4496</mods:number>
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<mods:identifier id="3BA3ABC2A1FE1256CC4C7E591BA679AD" type="DOI">10.11646/zootaxa.4496.1.42</mods:identifier>
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<treatment id="072D87D2FFCBFF8941DFFED2E0F0FD6F" ID-DOI="http://doi.org/10.5281/zenodo.6488171" ID-GBIF-Taxon="148640196" ID-Zenodo-Dep="6488171" LSID="urn:lsid:plazi:treatment:072D87D2FFCBFF8941DFFED2E0F0FD6F" httpUri="http://treatment.plazi.org/id/072D87D2FFCBFF8941DFFED2E0F0FD6F" lastPageId="4" lastPageNumber="552" pageId="2" pageNumber="550">
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<subSubSection id="C79E654FFFCBFF8F41DFFED2E2D0FE91" pageId="2" pageNumber="550" type="nomenclature">
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<paragraph id="8F3B36C4FFCBFF8F41DFFED2E703FEB0" blockId="2.[151,1113,319,379]" box="[151,1113,319,346]" pageId="2" pageNumber="550">
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<emphasis id="BDF0EAD6FFCBFF8F41DFFED2E703FEB0" bold="true" box="[151,1113,319,346]" pageId="2" pageNumber="550">
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<taxonomicName id="48844D47FFCBFF8F41DFFED2E2B0FEB3" authority="(GlossosColex)" authorityName="GlossosColex" box="[151,490,319,345]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="genus">
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<emphasis id="BDF0EAD6FFCBFF8F41DFFED2E2B0FEB3" bold="true" box="[151,490,319,345]" italics="true" pageId="2" pageNumber="550">GlossosColex (GlossosColex)</emphasis>
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</taxonomicName>
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<taxonomicName id="48844D47FFCBFF8F40BAFED2E753FEB0" authority="Bartz & James" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" box="[498,1033,319,346]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="subSpecies" species="riograndensis" status="sp. nov." subSpecies="riograndensis">
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<emphasis id="BDF0EAD6FFCBFF8F40BAFED2E01FFEB3" bold="true" box="[498,837,319,345]" italics="true" pageId="2" pageNumber="550">riograndensis riograndensis</emphasis>
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Bartz & James
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</taxonomicName>
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,
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<taxonomicNameLabel id="A6C357ADFFCBFF8F455EFEADE703FEB0" box="[1046,1113,320,346]" pageId="2" pageNumber="550" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8F3B36C4FFCBFF8F41DFFE8FE2D0FE91" blockId="2.[151,1113,319,379]" box="[151,394,354,379]" pageId="2" pageNumber="550">
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(
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<figureCitation id="17BF2A41FFCBFF8F41D6FE8FE263FE91" box="[158,313,354,379]" captionStart="FIGURE 1" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[307,1259,233,1868]" captionTargetId="figure@3.[190,1406,193,1912]" captionTargetPageId="3" captionText="FIGURE 1. Glossoscolex (Glossoscolex) riograndensis n. sp.. A, B. Glossoscolex (Glossoscolex) riograndensis riograndensis. C, D. Glossoscolex (Glossoscolex) riograndensis nativus n. ssp. E, F. Glossoscolex (Glossoscolex) riograndensis pollulus n. ssp. FM = female pOres; MP = male pOres; NF = nephridial pOres." httpUri="https://zenodo.org/record/1447027/files/figure.png" pageId="2" pageNumber="550">Figures 1A,B</figureCitation>
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,
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<figureCitation id="17BF2A41FFCBFF8F400DFE8EE2D8FE91" box="[325,386,355,379]" captionStart="FIGURE 2" captionStartId="6.[151,250,1324,1346]" captionTargetBox="[151,1436,339,1302]" captionTargetId="figure@6.[151,1436,339,1302]" captionTargetPageId="6" captionText="FIGURE 2. Sperm-filled nephridia Of Glossoscolex (Glossoscolex) riograndensis n. sp., histOlOgical sectiOns. A, B. Glossoscolex (G.) riograndensis riograndensis n. sp., segment xii (A) and xiii (B). C, D. Glossoscolex (G.) riograndensis pollulus n. ssp., segments Vi (C) and Vii (D)." httpUri="https://zenodo.org/record/1447029/files/figure.png" pageId="2" pageNumber="550">2A,B</figureCitation>
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)
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<materialsCitation id="3FEC3C99FFCBFF8F41DFFE47E7E3FE0D" ID-GBIF-Occurrence="1914265852" collectingDate="2011-11-09" collectionCode="COFM" collectorName="G. P. K. Steffen & R. B. Steffen" country="Brazil" elevation="105" latitude="-30.035471" location="Rosario do Sul" longLatPrecision="1" longitude="-54.581055" pageId="2" pageNumber="550" specimenCode="BRRS0038" specimenCount="1" stateProvince="Rio Grande do Sul" typeStatus="holotype">
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.
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</emphasis>
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<collectionCode id="E995AE01FFCBFF8F405DFE46E23EFE29" box="[277,356,427,451]" pageId="2" pageNumber="550">COFM</collectionCode>
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<specimenCode id="DF229EBFFFCBFF8F4027FE46E2B4FE29" box="[367,494,427,451]" collectionCode="BRRS" pageId="2" pageNumber="550">BRRS0038</specimenCode>
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one adult, fragment of natiVe forest,
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<location id="8A5B601FFFCBFF8F42EEFE47E70FFE29" LSID="urn:lsid:plazi:treatment:072D87D2FFCBFF8941DFFED2E0F0FD6F:8A5B601FFFCBFF8F42EEFE47E70FFE29" box="[934,1109,426,451]" country="Brazil" latitude="-30.035471" longLatPrecision="1" longitude="-54.581055" name="Rosario do Sul" pageId="2" pageNumber="550" stateProvince="Rio Grande do Sul">Rosário do Sul</location>
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,
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<collectingRegion id="4D40F826FFCBFF8F452CFE47E618FE29" box="[1124,1346,426,451]" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="550">
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Rio
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<collectingRegion id="4D40F826FFCBFF8F45DFFE47E618FE29" box="[1175,1346,426,451]" inferredBySuffix="true" pageId="2" pageNumber="550">Grande do Sul</collectingRegion>
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,
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<collectingCountry id="F7937654FFCBFF8F4419FE47E6C2FE29" box="[1361,1432,426,451]" name="Brazil" pageId="2" pageNumber="550">Brazil</collectingCountry>
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,
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<geoCoordinate id="EAB05003FFCBFF8F4066FE23E29CFE0D" box="[302,454,462,487]" direction="west" orientation="longitude" pageId="2" pageNumber="550" precision="1" value="-54.581055">54°34'51,8''O</geoCoordinate>
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,
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<elevation id="04A9D1F7FFCBFF8F4099FE23E160FE0D" box="[465,570,462,487]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="550" unit="m" value="105.0">
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<quantity id="487C9B21FFCBFF8F4099FE23E160FE0D" box="[465,570,462,487]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="550" unit="m" value="105.0">105 masl</quantity>
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</elevation>
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. 0
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<collectingDate id="EB7EE9ECFFCBFF8F431BFE22E045FE0D" box="[595,799,462,487]" pageId="2" pageNumber="550" value="2011-11-09">
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<date id="FB3A1004FFCBFF8F431BFE22E045FE0D" box="[595,799,462,487]" pageId="2" pageNumber="550" value="2011-11-09">9 NoVember 2011</date>
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</collectingDate>
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,
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<collectorName id="22715312FFCBFF8F4262FE22E092FE0D" box="[810,968,462,487]" pageId="2" pageNumber="550">G.P.K. Steffen</collectorName>
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&
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<collectorName id="22715312FFCBFF8F42A3FE22E72CFE0D" box="[1003,1142,462,487]" pageId="2" pageNumber="550">R.B. Steffen</collectorName>
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colls.
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<typeStatus id="503F8866FFCBFF8F418FFE1FE26CFDE1" box="[199,310,498,523]" pageId="2" pageNumber="550" type="paratype">Paratype</typeStatus>
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.
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</emphasis>
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<collectionCode id="E995AE01FFCBFF8F400AFE1EE285FDE1" box="[322,479,499,523]" pageId="2" pageNumber="550">COFM BRRS</collectionCode>
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0039 one adult. Same locality and collecting data as
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<typeStatus id="503F8866FFCBFF8F4565FE1FE7C9FDE1" box="[1069,1171,498,523]" pageId="2" pageNumber="550" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="8F3B36C4FFCBFF8F418FFDFBE257FCED" blockId="2.[151,1437,426,2035]" pageId="2" pageNumber="550">
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<materialsCitation id="3FEC3C99FFCBFF8F418FFDFBE395FD9D" ID-GBIF-Occurrence="1914265857" collectingDate="2011-11-09" collectionCode="COFM" collectorName="G. P. K. Steffen & R. B. Steffen" country="Brazil" elevation="105" latitude="-30.035471" location="Other" longLatPrecision="1" longitude="-54.581055" municipality="Rosario do Sul" pageId="2" pageNumber="550" specimenCode="BRRS0040" specimenCount="2" specimenCount-subadult="2" stateProvince="Rio Grande do Sul">
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<location id="8A5B601FFFCBFF8F418FFDFBE255FDC5" LSID="urn:lsid:plazi:treatment:072D87D2FFCBFF8941DFFED2E0F0FD6F:8A5B601FFFCBFF8F418FFDFBE255FDC5" box="[199,271,534,559]" country="Brazil" latitude="-30.035471" longLatPrecision="1" longitude="-54.581055" municipality="Rosario do Sul" name="Other" pageId="2" pageNumber="550" stateProvince="Rio Grande do Sul">Other</location>
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material.
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</emphasis>
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<specimenCode id="DF229EBFFFCBFF8F40A9FDFAE13AFDC5" box="[481,608,535,559]" collectionCode="BRRS" pageId="2" pageNumber="550">BRRS0040</specimenCode>
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one adult and
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, fragment of natiVe forest,
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<collectingMunicipality id="6F5FACBEFFCBFF8F45F5FDFBE63DFDC5" box="[1213,1383,534,559]" pageId="2" pageNumber="550">Rosário do Sul</collectingMunicipality>
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,
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<collectingRegion id="4D40F826FFCBFF8F443BFDFBE21AFDB9" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="550">
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Rio
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<collectingRegion id="4D40F826FFCBFF8F41DFFDD7E21AFDB9" box="[151,320,570,595]" inferredBySuffix="true" pageId="2" pageNumber="550">Grande do Sul</collectingRegion>
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</collectingRegion>
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,
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<collectingCountry id="F7937654FFCBFF8F4006FDD7E2CFFDB9" box="[334,405,570,595]" name="Brazil" pageId="2" pageNumber="550">Brazil</collectingCountry>
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,
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,
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|
||
<quantity id="487C9B21FFCBFF8F43AAFDD7E017FDB9" box="[738,845,570,595]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="550" unit="m" value="105.0">105 masl</quantity>
|
||
</elevation>
|
||
. 0
|
||
<collectingDate id="EB7EE9ECFFCBFF8F4221FDD6E760FDB9" box="[873,1082,570,595]" pageId="2" pageNumber="550" value="2011-11-09">
|
||
<date id="FB3A1004FFCBFF8F4221FDD6E760FDB9" box="[873,1082,570,595]" pageId="2" pageNumber="550" value="2011-11-09">9 NoVember 2011</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="22715312FFCBFF8F450FFDD6E7BDFDB9" box="[1095,1255,570,595]" pageId="2" pageNumber="550">G.P.K. Steffen</collectorName>
|
||
&
|
||
<collectorName id="22715312FFCBFF8F4458FDD6E6C6FDB9" box="[1296,1436,570,595]" pageId="2" pageNumber="550">R.B. Steffen</collectorName>
|
||
colls
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3FEC3C99FFCBFF8F4194FDB2E61BFD71" ID-GBIF-Occurrence="1914265848" collectingDate="2009-11-09" collectionCode="COFM" collectorName="G. P. K. Steffen & R. B. Steffen" country="Brazil" elevation="174" latitude="-30.035471" location="Rosario do Sul" longLatPrecision="1" longitude="-54.581055" pageId="2" pageNumber="550" specimenCode="BRRS0041" specimenCount="1" stateProvince="Rio Grande do Sul">
|
||
<collectionCode id="E995AE01FFCBFF8F4194FDB2E271FD9D" box="[220,299,607,631]" pageId="2" pageNumber="550">COFM</collectionCode>
|
||
<specimenCode id="DF229EBFFFCBFF8F407CFDB2E2EEFD9D" box="[308,436,607,631]" collectionCode="BRRS" pageId="2" pageNumber="550">BRRS0041</specimenCode>
|
||
fiVe adults and one subadult, fragment of natiVe forest,
|
||
<location id="8A5B601FFFCBFF8F4571FDB3E7BCFD9D" LSID="urn:lsid:plazi:treatment:072D87D2FFCBFF8941DFFED2E0F0FD6F:8A5B601FFFCBFF8F4571FDB3E7BCFD9D" box="[1081,1254,606,631]" country="Brazil" latitude="-30.035471" longLatPrecision="1" longitude="-54.581055" name="Rosario do Sul" pageId="2" pageNumber="550" stateProvince="Rio Grande do Sul">Rosário do Sul</location>
|
||
,
|
||
<collectingRegion id="4D40F826FFCBFF8F45BBFDB3E3E5FD71" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="550">
|
||
Rio
|
||
<collectingRegion id="4D40F826FFCBFF8F446DFDB3E3E5FD71" inferredBySuffix="true" pageId="2" pageNumber="550">Grande do Sul</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="F7937654FFCBFF8F4183FD6FE248FD71" box="[203,274,642,667]" name="Brazil" pageId="2" pageNumber="550">Brazil</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EAB05003FFCBFF8F4055FD6EE2F4FD71" box="[285,430,643,667]" direction="south" orientation="latitude" pageId="2" pageNumber="550" precision="1" value="-30.035471">30°02'07,7''S</geoCoordinate>
|
||
<geoCoordinate id="EAB05003FFCBFF8F40FDFD6FE117FD71" box="[437,589,642,667]" direction="west" orientation="longitude" pageId="2" pageNumber="550" precision="1" value="-54.581055">54°34'51,8''O</geoCoordinate>
|
||
,
|
||
<elevation id="04A9D1F7FFCBFF8F4311FD6EE198FD71" box="[601,706,642,667]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="2" pageNumber="550" unit="m" value="174.0">
|
||
<quantity id="487C9B21FFCBFF8F4311FD6EE198FD71" box="[601,706,642,667]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="2" pageNumber="550" unit="m" value="174.0">174 masl</quantity>
|
||
</elevation>
|
||
. 0
|
||
<collectingDate id="EB7EE9ECFFCBFF8F4394FD6EE0F0FD71" box="[732,938,642,667]" pageId="2" pageNumber="550" value="2009-11-09">
|
||
<date id="FB3A1004FFCBFF8F4394FD6EE0F0FD71" box="[732,938,642,667]" pageId="2" pageNumber="550" value="2009-11-09">9 NoVember 2009</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="22715312FFCBFF8F42FDFD6EE709FD71" box="[949,1107,642,667]" pageId="2" pageNumber="550">G.P.K. Steffen</collectorName>
|
||
&
|
||
<collectorName id="22715312FFCBFF8F453FFD6EE658FD71" box="[1143,1282,642,667]" pageId="2" pageNumber="550">R.B. Steffen</collectorName>
|
||
colls
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3FEC3C99FFCBFF8F4405FD6EE257FCED" ID-GBIF-Occurrence="1914265854" collectingDate="2009-11-09" collectionCode="COFM" collectorName="G. P. K. Steffen & R. B. Steffen" country="Brazil" elevation="151" latitude="-30.097418" location="Rosario do Sul" longLatPrecision="1" longitude="-54.631496" pageId="2" pageNumber="550" specimenCode="BRRS0042" specimenCount="17" stateProvince="Rio Grande do Sul">
|
||
<collectionCode id="E995AE01FFCBFF8F4405FD6EE6C6FD71" box="[1357,1436,643,667]" pageId="2" pageNumber="550">COFM</collectionCode>
|
||
<specimenCode id="DF229EBFFFCBFF8F41DFFD4AE24DFD55" box="[151,279,679,703]" collectionCode="BRRS" pageId="2" pageNumber="550">BRRS0042</specimenCode>
|
||
fiVe adults and one subadult, black acacia plantation,
|
||
<location id="8A5B601FFFCBFF8F42D5FD4BE715FD55" LSID="urn:lsid:plazi:treatment:072D87D2FFCBFF8941DFFED2E0F0FD6F:8A5B601FFFCBFF8F42D5FD4BE715FD55" box="[925,1103,678,703]" country="Brazil" latitude="-30.097418" longLatPrecision="1" longitude="-54.631496" name="Rosario do Sul" pageId="2" pageNumber="550" stateProvince="Rio Grande do Sul">Rosário do Sul</location>
|
||
,
|
||
<collectingRegion id="4D40F826FFCBFF8F4517FD4BE61BFD55" box="[1119,1345,678,703]" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="550">
|
||
Rio
|
||
<collectingRegion id="4D40F826FFCBFF8F45DBFD4BE61BFD55" box="[1171,1345,678,703]" inferredBySuffix="true" pageId="2" pageNumber="550">Grande do Sul</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="F7937654FFCBFF8F4419FD4BE6C2FD55" box="[1361,1432,678,703]" name="Brazil" pageId="2" pageNumber="550">Brazil</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EAB05003FFCBFF8F41DFFD27E27DFD09" box="[151,295,714,739]" direction="south" orientation="latitude" pageId="2" pageNumber="550" precision="1" value="-30.097418">30°05'50,7''S</geoCoordinate>
|
||
<geoCoordinate id="EAB05003FFCBFF8F407BFD27E290FD09" box="[307,458,714,739]" direction="west" orientation="longitude" pageId="2" pageNumber="550" precision="1" value="-54.631496">54°37'53,4''O</geoCoordinate>
|
||
,
|
||
<elevation id="04A9D1F7FFCBFF8F4093FD27E113FD09" box="[475,585,714,739]" metricMagnitude="2" metricUnit="m" metricValue="1.51" pageId="2" pageNumber="550" unit="m" value="151.0">
|
||
<quantity id="487C9B21FFCBFF8F4093FD27E113FD09" box="[475,585,714,739]" metricMagnitude="2" metricUnit="m" metricValue="1.51" pageId="2" pageNumber="550" unit="m" value="151.0">151 masl</quantity>
|
||
</elevation>
|
||
. 0
|
||
<collectingDate id="EB7EE9ECFFCBFF8F432FFD26E067FD09" box="[615,829,714,739]" pageId="2" pageNumber="550" value="2009-11-09">
|
||
<date id="FB3A1004FFCBFF8F432FFD26E067FD09" box="[615,829,714,739]" pageId="2" pageNumber="550" value="2009-11-09">9 NoVember 2009</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="22715312FFCBFF8F4205FD26E0AAFD09" box="[845,1008,714,739]" pageId="2" pageNumber="550">G.P.K. Steffen</collectorName>
|
||
&
|
||
<collectorName id="22715312FFCBFF8F4555FD26E7F6FD09" box="[1053,1196,714,739]" pageId="2" pageNumber="550">R.B. Steffen</collectorName>
|
||
colls.
|
||
<specimenCount id="9982FD4DFFCBFF8F4448FD26E6C7FD09" box="[1280,1437,714,739]" pageId="2" pageNumber="550" type="generic">17 specimens</specimenCount>
|
||
altogether.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C79E654FFFCBFF8F418FFCFFE7BAFCC1" box="[199,1248,786,811]" pageId="2" pageNumber="550" type="etymology">
|
||
<paragraph id="8F3B36C4FFCBFF8F418FFCFFE7BAFCC1" blockId="2.[151,1437,426,2035]" box="[199,1248,786,811]" pageId="2" pageNumber="550">
|
||
<emphasis id="BDF0EAD6FFCBFF8F418FFCFFE214FCC1" bold="true" box="[199,334,786,811]" pageId="2" pageNumber="550">Etymology.</emphasis>
|
||
The species is named due it occurrence in the State of
|
||
<collectingRegion id="4D40F826FFCBFF8F42FDFCFFE7D2FCC1" box="[949,1160,786,811]" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="550">Rio Grande do Sul</collectingRegion>
|
||
,
|
||
<collectingCountry id="F7937654FFCBFF8F45DCFCFFE786FCC1" box="[1172,1244,786,811]" name="Brazil" pageId="2" pageNumber="550">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C79E654FFFCBFF8F418FFCDBE139F945" pageId="2" pageNumber="550" type="description">
|
||
<paragraph id="8F3B36C4FFCBFF8F418FFCDBE1BCFBCD" blockId="2.[151,1437,426,2035]" pageId="2" pageNumber="550">
|
||
<emphasis id="BDF0EAD6FFCBFF8F418FFCDBE203FCA5" bold="true" box="[199,345,822,847]" pageId="2" pageNumber="550">Description.</emphasis>
|
||
Dimensions:
|
||
<typeStatus id="503F8866FFCBFF8F40BCFCDBE106FCA5" box="[500,604,822,847]" pageId="2" pageNumber="550" type="holotype">Holotype</typeStatus>
|
||
<quantity id="487C9B21FFCBFF8F432CFCDBE1E9FCA4" box="[612,691,822,847]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="550" unit="mm" value="54.0">54 mm</quantity>
|
||
by
|
||
<quantity id="487C9B21FFCBFF8F4396FCDAE06EFCA4" box="[734,820,823,847]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="550" unit="mm" value="3.7">3.7 mm</quantity>
|
||
at x, 4.0 mm at clitellum,
|
||
<quantity id="487C9B21FFCBFF8F4515FCDBE7EEFCA4" box="[1117,1204,822,847]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="550" unit="mm" value="3.5">3.5 mm</quantity>
|
||
at xl, 195 segments;
|
||
<typeStatus id="503F8866FFCBFF8F41DFFCB6E3ADFC99" box="[151,247,859,883]" pageId="2" pageNumber="550" type="paratype">paratype</typeStatus>
|
||
<quantity id="487C9B21FFCBFF8F41B6FCB7E216FC98" box="[254,332,858,883]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="550" unit="mm" value="53.0">53 mm</quantity>
|
||
by
|
||
<quantity id="487C9B21FFCBFF8F403DFCB6E290FC98" box="[373,458,859,883]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="550" unit="mm" value="4.1">4.1 mm</quantity>
|
||
at x,
|
||
<quantity id="487C9B21FFCBFF8F434FFCB6E106FC98" box="[519,604,859,883]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="550" unit="mm" value="4.2">4.2 mm</quantity>
|
||
at clitellum,
|
||
<quantity id="487C9B21FFCBFF8F43A7FCB6E01EFC98" box="[751,836,859,883]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="550" unit="mm" value="3.7">3.7 mm</quantity>
|
||
at xl, 125 segments. Other material ranges from
|
||
<quantity id="487C9B21FFCBFF8F442CFCB6E3B2FC7C" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="7.2" metricValueMin="4.1" pageId="2" pageNumber="550" unit="mm" value="56.5" valueMax="72.0" valueMin="41.0">41 to 72 mm</quantity>
|
||
in length, With 132–188 segments. Body cylindrical. Setae closely paired throughout starting in iV; setal formula aa:ab:bc:cd:dd = 7.8:0.3:1.0:0.2:11.2. Setae not Visible before clitellum. Prostomium prolobous. Unpigmented/Whitish. OVipores in a on xiV. Male pores
|
||
<quantity id="487C9B21FFCBFF8F4242FC2AE03AFC34" box="[778,864,967,991]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="550" unit="mm" value="2.8">2.8 mm</quantity>
|
||
apart on xVii Within paired round eleVations (like a mound); clitellum saddle, xV–xxiii and Ventrally until to xxii, in other material (
|
||
<specimenCount id="9982FD4DFFCBFF8F4550FC07E7F9FBE9" box="[1048,1187,1002,1027]" pageId="2" pageNumber="550" type="generic">5 specimens</specimenCount>
|
||
) the clitellum extends dorsally to 3/4 xxii (
|
||
<figureCitation id="17BF2A41FFCBFF8F4033FBE3E2B7FBCD" box="[379,493,1038,1063]" captionStart="FIGURE 1" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[307,1259,233,1868]" captionTargetId="figure@3.[190,1406,193,1912]" captionTargetPageId="3" captionText="FIGURE 1. Glossoscolex (Glossoscolex) riograndensis n. sp.. A, B. Glossoscolex (Glossoscolex) riograndensis riograndensis. C, D. Glossoscolex (Glossoscolex) riograndensis nativus n. ssp. E, F. Glossoscolex (Glossoscolex) riograndensis pollulus n. ssp. FM = female pOres; MP = male pOres; NF = nephridial pOres." httpUri="https://zenodo.org/record/1447027/files/figure.png" pageId="2" pageNumber="550">Fig. 1A,B</figureCitation>
|
||
). Nephropores near b.
|
||
</paragraph>
|
||
<paragraph id="8F3B36C4FFCBFF8F418FFBDEE0F3FAC9" blockId="2.[151,1437,426,2035]" pageId="2" pageNumber="550">
|
||
Septa
|
||
<date id="FB3A1004FFCBFF8F4047FBDFE2DAFBA1" box="[271,384,1074,1099]" pageId="2" pageNumber="550" value="2011-07-06" valueMax="2011-10-06" valueMin="2011-07-06">6/7–10/11</date>
|
||
equally thick and muscular. Alimentary canal With barrel-shaped gizzard in Vi; esophagus With high cheVron-patterned lamellae Vii–xi, ValVular in xii; intestinal origin xiii; typhlosole origin xiV, in xiV–xxV zig-zag With Ventral edge bent oVer to form pockets, xxVi–xlii straight lamina in Z form occupying half of the diameter of the intestine, after xliii gradually becoming simple S form lamina. Calciferous glands paired xii, heartshaped, pedunculated, composite-tubular
|
||
<typeStatus id="503F8866FFCBFF8F433EFB2EE1F0FB31" box="[630,682,1219,1243]" pageId="2" pageNumber="550">type</typeStatus>
|
||
; blood Vessels to gland include large branch of dorsal Vessel to approximate center of each gland. Holonephric, Vesiculate; ducts to body Wall near leVel of b; nephridia in xii, xiii full of sperm (iridescent material) in both specimens
|
||
<typeStatus id="503F8866FFCBFF8F43A1FAE6E043FAC9" box="[745,793,1291,1315]" pageId="2" pageNumber="550">type</typeStatus>
|
||
(
|
||
<figureCitation id="17BF2A41FFCBFF8F4260FAE7E0C7FAC9" box="[808,925,1290,1315]" captionStart="FIGURE 2" captionStartId="6.[151,250,1324,1346]" captionTargetBox="[151,1436,339,1302]" captionTargetId="figure@6.[151,1436,339,1302]" captionTargetPageId="6" captionText="FIGURE 2. Sperm-filled nephridia Of Glossoscolex (Glossoscolex) riograndensis n. sp., histOlOgical sectiOns. A, B. Glossoscolex (G.) riograndensis riograndensis n. sp., segment xii (A) and xiii (B). C, D. Glossoscolex (G.) riograndensis pollulus n. ssp., segments Vi (C) and Vii (D)." httpUri="https://zenodo.org/record/1447029/files/figure.png" pageId="2" pageNumber="550">Fig. 2A,B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8F3B36C4FFCBFF8F418FFAC3E071FA65" blockId="2.[151,1437,426,2035]" pageId="2" pageNumber="550">Vascular system With Ventral trunk, single dorsal trunk, lateral Vessels in Vii–ix, esophageal hearts in x–xi, last pair (xi) free. Extra-esophageal Vessel from pharyngeal glands, along Ventral–lateral face of gizzard, esophagus back to calciferous glands; supraesophageal Vessel in x–xi.</paragraph>
|
||
<paragraph id="8F3B36C4FFCBFF8F418FFA77E139F945" blockId="2.[151,1437,426,2035]" pageId="2" pageNumber="550">
|
||
OVaries, oVarian funnels free in xiii. Male sexual system metandric, testes and funnels in U shaped sac in xi, occupying part of xii; seminal Vesicles tubular, pass through xii–xiii connected lateral-Ventrally to the testis sacs, after xiV seminal Vesicles dorsally and tubular With lobulated edges ranging posteriorly along intestine farther than lxii; Vasa deferentia long, looped from testis sacs in xi, form dense zig-zag on body Wall on route to middle-anterior face of muscular round to oVal copulatory bulbs (
|
||
<quantity id="487C9B21FFCBFF8F4383F9C6E00EF9A8" box="[715,852,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="2" pageNumber="550" unit="mm" value="1.5" valueMax="1.7" valueMin="1.3">1.3–1.7 mm</quantity>
|
||
diameter); bulbs extend oVer xVi–xViii. Copulatory bulbs With thin muscular outer layer, dense, delicate corrugated glandular inner surface With small lumen leading to male pore at approximate center of bulb connection to body Wall; no transVerse muscle bands crossing oVer bulbs, under face of bulb attached to body Wall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C79E654FFFCBFF89418FF957E0F0FD6F" lastPageId="4" lastPageNumber="552" pageId="2" pageNumber="550" type="discussion">
|
||
<paragraph id="8F3B36C4FFCBFF8F418FF957E2BFF841" blockId="2.[151,1437,426,2035]" pageId="2" pageNumber="550">
|
||
<emphasis id="BDF0EAD6FFCBFF8F418FF957E261F939" bold="true" box="[199,315,1722,1747]" pageId="2" pageNumber="550">Remarks.</emphasis>
|
||
<taxonomicName id="48844D47FFCBFF8F400AF956E091F939" authority="riograndensis" box="[322,971,1723,1747]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="riograndensis">
|
||
<emphasis id="BDF0EAD6FFCBFF8F400AF956E091F939" box="[322,971,1723,1747]" italics="true" pageId="2" pageNumber="550">Glossoscolex (Glossoscolex) riograndensis riograndensis</emphasis>
|
||
</taxonomicName>
|
||
is close to
|
||
<taxonomicName id="48844D47FFCBFF8F4504F956E217F91D" authority="Zicsi & Czusdi, 1999" authorityName="Zicsi & Czusdi" authorityYear="1999" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="species" species="minor" subGenus="Glossoscolex">
|
||
<emphasis id="BDF0EAD6FFCBFF8F4504F956E606F938" box="[1100,1372,1723,1747]" italics="true" pageId="2" pageNumber="550">Glossoscolex (G.) minor</emphasis>
|
||
<bibRefCitation id="EB154B35FFCBFF8F442CF957E217F91D" pageId="2" pageNumber="550" refId="ref9066" refString="Zicsi, A. & CsuZdi, C. (1999) Neue und bekannte RegenWurmer aus Verschiedenen Teilen Sudamerikas. Senckenbergiana Biologica, 78 (1 / 2), 123 - 134." type="journal article">Zicsi & Czusdi, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
considering the small body size—all other described species of this subgenus are considerably larger (comp.
|
||
<tableCitation id="C206037FFFCBFF8F407DF8EFE2D2F8F1" box="[309,392,1794,1819]" captionStart="TABLE 1" captionStartId="10.[147,228,591,614]" captionText="TABLE 1. Comparison of characters of the new species and subspecies of Glossoscolex (Glossoscolex) with the type species of the subgenus and with further similar species that have been compared in the remarks sections of the new taxa described." pageId="2" pageNumber="550">Table 1</tableCitation>
|
||
). The differences betWeen
|
||
<taxonomicName id="48844D47FFCBFF8F43F5F8EEE09BF8F1" box="[701,961,1795,1819]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="riograndensis">
|
||
<emphasis id="BDF0EAD6FFCBFF8F43F5F8EEE09BF8F1" box="[701,961,1795,1819]" italics="true" pageId="2" pageNumber="550">G. (G.) r. riograndensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="48844D47FFCBFF8F42B0F8EEE7D7F8F0" authorityName="Zicsi & Czusdi" authorityYear="1999" box="[1016,1165,1795,1818]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="species" species="minor" subGenus="Glossoscolex">
|
||
<emphasis id="BDF0EAD6FFCBFF8F42B0F8EEE7D7F8F0" box="[1016,1165,1795,1818]" italics="true" pageId="2" pageNumber="550">G. (G.) minor</emphasis>
|
||
</taxonomicName>
|
||
are as folloWs, With the characteristics of the latter in parentheses: length
|
||
<quantity id="487C9B21FFCBFF8F439FF8CAE009F8D4" box="[727,851,1831,1855]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="7.2" metricValueMin="4.1" pageId="2" pageNumber="550" unit="mm" value="56.5" valueMax="72.0" valueMin="41.0">41–72 mm</quantity>
|
||
(
|
||
<quantity id="487C9B21FFCBFF8F422CF8CBE0BCF8D5" box="[868,998,1830,1855]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="4.5" metricValueMin="2.4" pageId="2" pageNumber="550" unit="mm" value="34.5" valueMax="45.0" valueMin="24.0">24–45 mm</quantity>
|
||
), number of segments 125–195 (198– 231), clitellum saddle (annular), clitellum extension xV–xxiii (xiV–xxiii), testis sacs U shaped sac in xi–xii (unpaired in xi), seminal Vesicles long tubular With lobulated edges until lxii (band until l), copulatory bulbs round to oVal (elongated). (
|
||
<tableCitation id="C206037FFFCBFF8F40C8F87FE289F841" box="[384,467,1938,1963]" captionStart="TABLE 1" captionStartId="10.[147,228,591,614]" captionText="TABLE 1. Comparison of characters of the new species and subspecies of Glossoscolex (Glossoscolex) with the type species of the subgenus and with further similar species that have been compared in the remarks sections of the new taxa described." pageId="2" pageNumber="550">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8F3B36C4FFCBFF8F418FF85AE3BBF819" blockId="2.[151,1437,426,2035]" pageId="2" pageNumber="550">
|
||
For comparison of
|
||
<taxonomicName id="48844D47FFCBFF8F40D3F85AE1C1F824" authority="Bartz & James & Steffen & Antoniolli & Steffen & Brown, 2018" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" box="[411,667,1974,1998]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="subSpecies" species="riograndensis" status="subsp. nov." subGenus="Glossoscolex" subSpecies="riograndensis">
|
||
<emphasis id="BDF0EAD6FFCBFF8F40D3F85AE1C1F824" box="[411,667,1974,1998]" italics="true" pageId="2" pageNumber="550">G. (G.) r. riograndensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6C357ADFFCBFF8F43E9F85BE1B0F825" box="[673,746,1974,1999]" pageId="2" pageNumber="550" rank="subSpecies">
|
||
<emphasis id="BDF0EAD6FFCBFF8F43E9F85BE1B0F825" bold="true" box="[673,746,1974,1999]" pageId="2" pageNumber="550">n. ssp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
With
|
||
<taxonomicName id="48844D47FFCBFF8F4261F85AE0B3F824" authority="Bartz & James & Steffen & Antoniolli & Steffen & Brown, 2018" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" box="[809,1001,1974,1999]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="subSpecies" species="riograndensis" status="subsp. nov." subGenus="Glossoscolex" subSpecies="pollulus">
|
||
<emphasis id="BDF0EAD6FFCBFF8F4261F85AE060F824" box="[809,826,1975,1998]" italics="true" pageId="2" pageNumber="550">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCBFF8F4219F85AE038F824" box="[849,866,1975,1998]" italics="true" pageId="2" pageNumber="550">G</emphasis>
|
||
.)
|
||
<emphasis id="BDF0EAD6FFCBFF8F4230F85AE0D8F824" box="[888,898,1975,1998]" italics="true" pageId="2" pageNumber="550">r</emphasis>
|
||
.
|
||
<emphasis id="BDF0EAD6FFCBFF8F42C7F85BE0B3F824" box="[911,1001,1974,1998]" italics="true" pageId="2" pageNumber="550">pollulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6C357ADFFCBFF8F42B8F85BE763F825" box="[1008,1081,1974,1999]" pageId="2" pageNumber="550" rank="subSpecies">
|
||
<emphasis id="BDF0EAD6FFCBFF8F42B8F85BE75DF825" bold="true" box="[1008,1031,1974,1999]" pageId="2" pageNumber="550">n.</emphasis>
|
||
<taxonomicName id="48844D47FFCBFF8F4545F85BE763F825" box="[1037,1081,1974,1999]" pageId="2" pageNumber="550" rank="subSpecies" subSpecies="and">
|
||
<emphasis id="BDF0EAD6FFCBFF8F4545F85BE763F825" bold="true" box="[1037,1081,1974,1999]" pageId="2" pageNumber="550">ssp.</emphasis>
|
||
</taxonomicName>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="48844D47FFCBFF8F4527F85AE67CF824" authority="Bartz & James & Steffen & Antoniolli & Steffen & Brown, 2018" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" box="[1135,1318,1975,1999]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="550" phylum="Annelida" rank="subSpecies" species="riograndensis" status="sp. nov." subGenus="Glossoscolex" subSpecies="nativus">
|
||
<emphasis id="BDF0EAD6FFCBFF8F4527F85AE7DAF824" box="[1135,1152,1975,1998]" italics="true" pageId="2" pageNumber="550">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCBFF8F45DEF85AE7FDF824" box="[1174,1191,1975,1998]" italics="true" pageId="2" pageNumber="550">G</emphasis>
|
||
.)
|
||
<emphasis id="BDF0EAD6FFCBFF8F45F6F85AE79DF824" box="[1214,1223,1975,1998]" italics="true" pageId="2" pageNumber="550">r</emphasis>
|
||
.
|
||
<emphasis id="BDF0EAD6FFCBFF8F459DF85AE67CF824" box="[1237,1318,1975,1998]" italics="true" pageId="2" pageNumber="550">nativus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6C357ADFFCBFF8F4464F85BE630F825" box="[1324,1386,1974,1999]" pageId="2" pageNumber="550" rank="species">
|
||
<emphasis id="BDF0EAD6FFCBFF8F4464F85BE630F825" bold="true" box="[1324,1386,1974,1999]" pageId="2" pageNumber="550">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, see beloW.
|
||
</paragraph>
|
||
<caption id="DBFB664CFFCAFF8E41DFF87DE761F809" httpUri="https://zenodo.org/record/1447027/files/figure.png" pageId="3" pageNumber="551" startId="3.[151,250,1936,1958]" targetBox="[307,1259,233,1868]" targetPageId="3">
|
||
<paragraph id="8F3B36C4FFCAFF8E41DFF87DE761F809" blockId="3.[151,1436,1936,2019]" pageId="3" pageNumber="551">
|
||
<emphasis id="BDF0EAD6FFCAFF8E41DFF87DE27AF84C" bold="true" box="[151,288,1936,1958]" pageId="3" pageNumber="551">FIGURE 1.</emphasis>
|
||
<taxonomicName id="48844D47FFCAFF8E407BF87DE058F84C" authority="Bartz & James & Steffen & Antoniolli & Steffen & Brown, 2018" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" box="[307,770,1936,1958]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="551" phylum="Annelida" rank="species" species="riograndensis" status="sp. nov." subGenus="Glossoscolex">
|
||
<emphasis id="BDF0EAD6FFCAFF8E407BF87DE058F84C" box="[307,770,1936,1958]" italics="true" pageId="3" pageNumber="551">Glossoscolex (Glossoscolex) riograndensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6C357ADFFCAFF8E425EF87DE006F84C" box="[790,860,1936,1958]" pageId="3" pageNumber="551" rank="species">
|
||
<emphasis id="BDF0EAD6FFCAFF8E425EF87DE006F84C" bold="true" box="[790,860,1936,1958]" pageId="3" pageNumber="551">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
.
|
||
<emphasis id="BDF0EAD6FFCAFF8E423EF87DE0D2F84C" bold="true" box="[886,904,1936,1958]" pageId="3" pageNumber="551">A</emphasis>
|
||
,
|
||
<emphasis id="BDF0EAD6FFCAFF8E42EAF87DE0E2F84C" bold="true" box="[930,952,1936,1958]" pageId="3" pageNumber="551">B.</emphasis>
|
||
<taxonomicName id="48844D47FFCAFF8E4284F87DE279F82E" authority="riograndensis" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="551" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="riograndensis">
|
||
<emphasis id="BDF0EAD6FFCAFF8E4284F87DE279F82E" italics="true" pageId="3" pageNumber="551">Glossoscolex (Glossoscolex) riograndensis riograndensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="BDF0EAD6FFCAFF8E4073F843E217F82E" bold="true" box="[315,333,1966,1988]" pageId="3" pageNumber="551">C</emphasis>
|
||
,
|
||
<emphasis id="BDF0EAD6FFCAFF8E402CF843E227F82E" bold="true" box="[356,381,1966,1988]" pageId="3" pageNumber="551">D.</emphasis>
|
||
<taxonomicName id="48844D47FFCAFF8E40C6F843E0E8F82F" authority="Bartz & James & Steffen & Antoniolli & Steffen & Brown, 2018" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" box="[398,946,1966,1989]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="551" phylum="Annelida" rank="subSpecies" species="riograndensis" status="subsp. nov." subGenus="Glossoscolex" subSpecies="nativus">
|
||
<emphasis id="BDF0EAD6FFCAFF8E40C6F843E0E8F82F" box="[398,946,1966,1989]" italics="true" pageId="3" pageNumber="551">Glossoscolex (Glossoscolex) riograndensis nativus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BDF0EAD6FFCAFF8E428CF843E769F82E" bold="true" box="[964,1075,1966,1988]" pageId="3" pageNumber="551">
|
||
<taxonomicNameLabel id="A6C357ADFFCAFF8E428CF843E74BF82E" box="[964,1041,1966,1988]" pageId="3" pageNumber="551" rank="subSpecies">n. ssp.</taxonomicNameLabel>
|
||
E
|
||
</emphasis>
|
||
,
|
||
<emphasis id="BDF0EAD6FFCAFF8E4503F843E706F82E" bold="true" box="[1099,1116,1966,1988]" pageId="3" pageNumber="551">F.</emphasis>
|
||
<taxonomicName id="48844D47FFCAFF8E4527F843E226F809" authority="Bartz & James & Steffen & Antoniolli & Steffen & Brown, 2018" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="551" phylum="Annelida" rank="subSpecies" species="riograndensis" status="subsp. nov." subGenus="Glossoscolex" subSpecies="pollulus">
|
||
<emphasis id="BDF0EAD6FFCAFF8E4527F843E226F809" italics="true" pageId="3" pageNumber="551">Glossoscolex (Glossoscolex) riograndensis pollulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6C357ADFFCAFF8E40CAF820E29FF809" box="[386,453,1997,2019]" pageId="3" pageNumber="551" rank="subSpecies">
|
||
<emphasis id="BDF0EAD6FFCAFF8E40CAF820E29FF809" bold="true" box="[386,453,1997,2019]" pageId="3" pageNumber="551">n. ssp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
FM = female pOres; MP = male pOres; NF = nephridial pOres.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8F3B36C4FFCDFF89418FFF7AE0F0FD6F" blockId="4.[151,1436,151,645]" pageId="4" pageNumber="552">
|
||
The presence of sperm cells in some post- and pre-clitellar nephridia of this species is highly unusual in knoWn
|
||
<taxonomicName id="48844D47FFCDFF8941DFFF51E271FF3E" box="[151,299,188,212]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="genus">
|
||
<emphasis id="BDF0EAD6FFCDFF8941DFFF51E271FF3E" box="[151,299,188,212]" italics="true" pageId="4" pageNumber="552">Glossoscolex</emphasis>
|
||
</taxonomicName>
|
||
and the closely related
|
||
<taxonomicName id="48844D47FFCDFF89430CFF51E19FFF3E" authorityName="Michaelsen" authorityYear="1900" box="[580,709,188,212]" class="Clitellata" family="Glossoscolecidae" genus="Fimoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="genus">
|
||
<emphasis id="BDF0EAD6FFCDFF89430CFF51E19FFF3E" box="[580,709,188,212]" italics="true" pageId="4" pageNumber="552">Fimoscolex</emphasis>
|
||
</taxonomicName>
|
||
. Both genera are athecal, i.e. Without spermatophores, and yet nearly all species shoW tremendous inVestment in sperm production and storage, With extremely long and WelldeVeloped seminal Vesicles. All haVe large muscular ejaculatory structures; the copulatory bursae empty Via the male pores. Among the hundreds of specimens that We haVe examined none shoWs signs of parthenogenetic reduction of the male gonads. Only
|
||
<emphasis id="BDF0EAD6FFCDFF89437DFEA1E117FE89" box="[565,589,332,355]" italics="true" pageId="4" pageNumber="552">G.</emphasis>
|
||
sp.1 (see beloW) is aVesiculate but eVen in that species the male anatomy looks otherWise normal. GiVen the large inVestment in the male reproductiVe apparatus one should expect bisexual reproduction Via mutual sperm exchange during copulation, but in fact We can only speculate What is going on in the many
|
||
<taxonomicName id="48844D47FFCDFF894044FE55E2FAFE3A" box="[268,416,440,464]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="genus">
|
||
<emphasis id="BDF0EAD6FFCDFF894044FE55E2FAFE3A" box="[268,416,440,464]" italics="true" pageId="4" pageNumber="552">Glossoscolex</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="48844D47FFCDFF894095FE55E104FE3A" authorityName="Michaelsen" authorityYear="1900" box="[477,606,440,464]" class="Clitellata" family="Glossoscolecidae" genus="Fimoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="genus">
|
||
<emphasis id="BDF0EAD6FFCDFF894095FE55E104FE3A" box="[477,606,440,464]" italics="true" pageId="4" pageNumber="552">Fimoscolex</emphasis>
|
||
</taxonomicName>
|
||
species Where We haVe not Validated the presence of sperm in unusual locations. Prior to this study We haVe reported on strong deVelopment of unusually White spots and tiny pockets on enlarged septa in the region of segments xiii–xiV in
|
||
<taxonomicName id="48844D47FFCDFF894254FDEDE73BFDFD" authorityName="Bartz & James" authorityYear="2012" box="[796,1121,512,536]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="itaguajensis" subGenus="Praedrilus">
|
||
<emphasis id="BDF0EAD6FFCDFF894254FDEDE077FDFD" box="[796,813,512,535]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCDFF894203FDEDE099FDF2" box="[843,963,512,536]" italics="true" pageId="4" pageNumber="552">Praedrilus</emphasis>
|
||
)
|
||
<emphasis id="BDF0EAD6FFCDFF894292FDEDE73BFDFD" box="[986,1121,512,535]" italics="true" pageId="4" pageNumber="552">itaguajensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="48844D47FFCDFF89453EFDEDE662FDF2" box="[1142,1336,512,536]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="lutoculus" subGenus="Praedrilus">
|
||
<emphasis id="BDF0EAD6FFCDFF89453EFDEDE7DDFDFD" box="[1142,1159,512,535]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCDFF8945EEFDECE7EEFDF2" box="[1190,1204,513,536]" italics="true" pageId="4" pageNumber="552">P</emphasis>
|
||
.)
|
||
<emphasis id="BDF0EAD6FFCDFF89459AFDEDE662FDF2" box="[1234,1336,512,536]" italics="true" pageId="4" pageNumber="552">lutoculus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="48844D47FFCDFF894406FDEDE250FDD6" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="uliginosus" subGenus="Praedrilus">
|
||
<emphasis id="BDF0EAD6FFCDFF894406FDEDE605FDFD" box="[1358,1375,512,535]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCDFF894435FDECE6D1FDF2" box="[1405,1419,513,536]" italics="true" pageId="4" pageNumber="552">P</emphasis>
|
||
.)
|
||
<emphasis id="BDF0EAD6FFCDFF8941DFFDC9E250FDD6" box="[151,266,548,572]" italics="true" pageId="4" pageNumber="552">uliginosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="48844D47FFCDFF89405FFDC9E28DFDD6" authorityName="Bartz & James" authorityYear="2012" box="[279,471,548,572]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="mariarum" subGenus="Glossoscolex">
|
||
<emphasis id="BDF0EAD6FFCDFF89405FFDC9E272FDD1" box="[279,296,548,571]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCDFF894077FDC9E20AFDD1" box="[319,336,548,571]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
.)
|
||
<emphasis id="BDF0EAD6FFCDFF89402FFDC8E28DFDD6" box="[359,471,549,572]" italics="true" pageId="4" pageNumber="552">mariarum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="48844D47FFCDFF8940ADFDC9E1F5FDD6" authorityName="Bartz & James" authorityYear="2012" box="[485,687,548,572]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="primaensis" subGenus="Glossoscolex">
|
||
<emphasis id="BDF0EAD6FFCDFF8940ADFDC9E2ACFDD1" box="[485,502,548,571]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCDFF894345FDC9E144FDD1" box="[525,542,548,571]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
.)
|
||
<emphasis id="BDF0EAD6FFCDFF89437CFDC8E1F5FDD6" box="[564,687,549,572]" italics="true" pageId="4" pageNumber="552">primaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="48844D47FFCDFF8943ADFDC9E731FDD6" authority="(Bartz 2012)" authorityName="Bartz" authorityYear="2012" baseAuthorityName="Bartz" baseAuthorityYear="2012" box="[741,1131,548,572]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="sanpredoensis" subGenus="Glossoscolex">
|
||
<emphasis id="BDF0EAD6FFCDFF8943ADFDC9E1ACFDD1" box="[741,758,548,571]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
. (
|
||
<emphasis id="BDF0EAD6FFCDFF894245FDC9E044FDD1" box="[781,798,548,571]" italics="true" pageId="4" pageNumber="552">G</emphasis>
|
||
.)
|
||
<emphasis id="BDF0EAD6FFCDFF89427DFDC9E08EFDD6" box="[821,980,548,572]" italics="true" pageId="4" pageNumber="552">sanpredoensis</emphasis>
|
||
(Bartz 2012)
|
||
</taxonomicName>
|
||
, and We suspected possible sperm storage in those cases. In the case of
|
||
<taxonomicName id="48844D47FFCDFF8943CEFDA5E018FD8A" authorityName="Bartz & James & Steffen & Antoniolli & Steffen & Brown" authorityYear="2018" box="[646,834,584,608]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="riograndensis">
|
||
<emphasis id="BDF0EAD6FFCDFF8943CEFDA5E018FD8A" box="[646,834,584,608]" italics="true" pageId="4" pageNumber="552">G. riograndensis</emphasis>
|
||
</taxonomicName>
|
||
it appears that sperm receiVed in copulation, perhaps by high-pressure injection into nephropores, is stored in the nephridia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |