254 lines
33 KiB
XML
254 lines
33 KiB
XML
<document id="99D57CCF438ED78C7E9B9361DDA04BD5" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FF9DF751035110DF4EB8C9D8" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada mckenziei Köhler 2010, n.sp." docType="treatment" docUuid="34B98E97-0DFA-413B-8A45-7596855D4940" docUuidSource="ZooBank" docVersion="3" lastPageNumber="230" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="229" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="22FBDCCF54F04688E7E642709EF7400B">
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<mods:title id="A778786CF07F7FC9F4C54B96A1EB8F69">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9D2C1E70336273CD6DF11D5A0A734234" type="personal">
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<mods:roleTerm id="986C4EAA48D513CB4FFCD4BAF5333AC9">Author</mods:roleTerm>
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<mods:namePart id="61CEFD16D2C653793532A0E1F849EBFD">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="4ADA1EB4EBCCE8730B50E06D0B951B74">text</mods:typeOfResource>
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<mods:title id="B4AAC2A9878FE886C368337BA1763288">Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part id="F2A0F209C731755A02D582E1AFA6341B">
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<mods:date id="F59072BB38669C56432E515B7E645A50">2010</mods:date>
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<mods:detail id="B5B6EEE346DFC4D01940AE631E1BAE3C" type="pubDate">
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<mods:number id="96E21D0261D6A76ED9A1D5E5711A523A">2010-11-24</mods:number>
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</mods:detail>
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<mods:detail id="BCDF2E41D31C6E089C74F4B388FABC98" type="volume">
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<mods:number id="C73E61D23336686098EAF7B1D93CACDA">62</mods:number>
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</mods:detail>
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<mods:detail id="5DE9A94428359C35108F0BA3EC67B92E" type="issue">
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<mods:number id="DF6C3FAE760EF7801DE38DB9F6109623">2</mods:number>
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</mods:detail>
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<mods:extent id="10E99412A89E3401FCE04F37B188C6B7" unit="page">
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<mods:start id="E709ED5DC8B18B57CF5538E97A3D3740">217</mods:start>
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<mods:end id="5E6ABCDE9E97334672448D396FE861E1">284</mods:end>
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<mods:location id="E16CAA27FE3639F0B4AF4208C61DFF6F">
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<mods:url id="E843019A79A7C21A23762DBB13F0F40C">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
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</mods:location>
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<mods:classification id="57BC5535A0B8CFD6A9360D09959B1ADD">journal article</mods:classification>
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<mods:identifier id="A228CD45A4F2CE107456B97CD778D08C" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
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<mods:identifier id="D53BDD17D2F7B5E5F0A242C12EB0A091" type="ISSN">2201-4349</mods:identifier>
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<mods:identifier id="8EF3CDE2D35B48CA2525FF2AB969456D" type="Zenodo-Dep">5239086</mods:identifier>
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<mods:identifier id="4AC2D8670E431ED7722F12005D6C404C" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
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<treatment id="03F987C5FF9DF751035110DF4EB8C9D8" ID-DOI="http://doi.org/10.5281/zenodo.5239015" ID-Zenodo-Dep="5239015" LSID="urn:lsid:zoobank.org:act:34B98E97-0DFA-413B-8A45-7596855D4940" httpUri="http://treatment.plazi.org/id/03F987C5FF9DF751035110DF4EB8C9D8" lastPageId="13" lastPageNumber="230" pageId="12" pageNumber="229">
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<subSubSection id="C34A6558FF9DF750035110DF4F89CEC0" box="[292,668,154,180]" pageId="12" pageNumber="229" type="nomenclature">
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<paragraph id="8BEF36D3FF9DF750035110DF4F89CEC0" blockId="12.[292,668,154,180]" box="[292,668,154,180]" pageId="12" pageNumber="229">
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<heading id="D0A781BFFF9DF750035110DF4F89CEC0" box="[292,668,154,180]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="229" reason="2">
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<emphasis id="B924EAC1FF9DF750035110DF4F89CEC0" bold="true" box="[292,668,154,180]" pageId="12" pageNumber="229">
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<taxonomicName id="4C504D50FF9DF750035110DF4F4DCEC0" ID-CoL="D7RJ" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[292,600,154,180]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="229" phylum="Mollusca" rank="species" species="mckenziei" status="sp. nov.">
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<emphasis id="B924EAC1FF9DF750035110DF4F4DCEC0" bold="true" box="[292,600,154,180]" italics="true" pageId="12" pageNumber="229">Amplirhagada mckenziei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21757BAFF9DF750001510DF4F89CEC0" box="[608,668,154,180]" pageId="12" pageNumber="229" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6558FF9DF75002D310924F44CC64" pageId="12" pageNumber="229" type="materials_examined">
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<paragraph id="8BEF36D3FF9DF75002D310924E0FCF32" blockId="12.[166,794,215,326]" pageId="12" pageNumber="229">
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<materialsCitation id="3B383C8EFF9DF75002D310924E03CF32" collectingDate="1988-06-18" collectionCode="V" collectorName="Kessner & A. Longbottom" county="northwestern Kimberley" latitude="-14.094999" location="Admiralty Gulf" longLatPrecision="21" longitude="125.74166" municipality="Bonaparte Archipelago" pageId="12" pageNumber="229" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B924EAC1FF9DF75002D310924C55CE9B" bold="true" box="[166,320,215,239]" pageId="12" pageNumber="229">
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||
<typeStatus id="54EB8871FF9DF75002D310924DF5CE9B" box="[166,224,215,239]" pageId="12" pageNumber="229">Type</typeStatus>
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||
locality
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||
</emphasis>
|
||
(
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<figureCitation id="136B2A56FF9DF750032610924C82CE9A" box="[339,407,215,238]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="12" pageNumber="229">Fig. 1</figureCitation>
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).
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||
<collectingRegion id="4994F831FF9DF75003C710924F69CE9A" box="[434,636,215,238]" country="Australia" name="Western Australia" pageId="12" pageNumber="229">Western Australia</collectingRegion>
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,
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<collectingCounty id="628E4E5FFF9DF75000FF10924C0CCF7F" pageId="12" pageNumber="229">northwestern Kimberley</collectingCounty>
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,
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<collectingMunicipality id="6B8BACA9FF9DF750035610B14F08CF7F" box="[291,541,244,267]" pageId="12" pageNumber="229">Bonaparte Archipelago</collectingMunicipality>
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||
,
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||
<location id="8E8F6008FF9DF750005010B14FD8CF7F" LSID="urn:lsid:plazi:treatment:03F987C5FF9DF751035110DF4EB8C9D8:8E8F6008FF9DF750005010B14FD8CF7F" box="[549,717,244,267]" county="northwestern Kimberley" latitude="-14.094999" longLatPrecision="21" longitude="125.74166" municipality="Bonaparte Archipelago" name="Admiralty Gulf" pageId="12" pageNumber="229" stateProvince="Western Australia">Admiralty Gulf</location>
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||
,
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<location id="8E8F6008FF9DF75000A210B14C46CF5D" LSID="urn:lsid:plazi:treatment:03F987C5FF9DF751035110DF4EB8C9D8:8E8F6008FF9DF75000A210B14C46CF5D" county="northwestern Kimberley" latitude="-14.094999" longLatPrecision="21" longitude="125.74166" municipality="Bonaparte Archipelago" name="Montesquieu Islands" pageId="12" pageNumber="229" stateProvince="Western Australia">Montesquieu Islands</location>
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,
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||
<location id="8E8F6008FF9DF750031411574CE1CF5D" LSID="urn:lsid:plazi:treatment:03F987C5FF9DF751035110DF4EB8C9D8:8E8F6008FF9DF750031411574CE1CF5D" box="[353,500,274,297]" county="northwestern Kimberley" latitude="-14.094999" longLatPrecision="21" longitude="125.74166" municipality="Bonaparte Archipelago" name="Oliver Island" pageId="12" pageNumber="229" stateProvince="Western Australia">Oliver Island</location>
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,
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<geoCoordinate id="EE645014FF9DF750007611574F68CF5D" box="[515,637,274,297]" degrees="14" direction="south" minutes="05" orientation="latitude" pageId="12" pageNumber="229" precision="15" seconds="42" value="-14.094999">14°05'42"S</geoCoordinate>
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<geoCoordinate id="EE645014FF9DF75000FD11574E00CF5D" box="[648,789,274,297]" degrees="125" direction="east" minutes="44" orientation="longitude" pageId="12" pageNumber="229" precision="15" seconds="30" value="125.74166">125°44'30"E</geoCoordinate>
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||
; KC-062 (leg.
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||
<collectionCode id="ED41AE16FF9DF750034D116A4C5ECF32" box="[312,331,303,326]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="229" type="Museum">V</collectionCode>
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||
.
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||
<collectorName id="26A55305FF9DF7500320116A4CBFCF32" box="[341,426,303,326]" pageId="12" pageNumber="229">Kessner</collectorName>
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||
&
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||
<collectorName id="26A55305FF9DF75003BE116A4F67CF32" box="[459,626,303,326]" pageId="12" pageNumber="229">A. Longbottom</collectorName>
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||
,
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||
<date id="FFEE1013FF9DF7500009116A4E1BCF32" box="[636,782,303,326]" pageId="12" pageNumber="229" value="1988-06-18">
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||
<collectingDate id="EFAAE9FBFF9DF7500009116A4E1BCF32" box="[636,782,303,326]" pageId="12" pageNumber="229" value="1988-06-18">18 June 1988</collectingDate>
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||
</date>
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||
)
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||
</materialsCitation>
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.
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||
</paragraph>
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||
<paragraph id="8BEF36D3FF9DF75002D311224FCBCFE7" blockId="12.[166,793,359,403]" pageId="12" pageNumber="229">
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||
<emphasis id="B924EAC1FF9DF75002D311224C30CF0E" bold="true" box="[166,293,359,378]" pageId="12" pageNumber="229">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B383C8EFF9DF750035B112D4F51CF0F" accessionNumber="S34604" box="[302,580,360,379]" collectionCode="WAM" pageId="12" pageNumber="229" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54EB8871FF9DF750035B112D4C68CF0F" box="[302,381,360,379]" pageId="12" pageNumber="229" type="holotype">Holotype</typeStatus>
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<collectionCode id="ED41AE16FF9DF75003F4112D4CA0CF0F" box="[385,437,360,379]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="229" type="Museum">WAM</collectionCode>
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<accessionNumber id="9403AB30FF9DF75003CD112D4CEFCF0F" box="[440,506,360,379]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34604" pageId="12" pageNumber="229" type="EnaNcbi">S34604</accessionNumber>
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(Pl. 1.8)
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||
</materialsCitation>
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.
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<materialsCitation id="3B383C8EFF9DF750003E112D4CABCFE7" accessionNumber="S41488" collectionCode="WAM, FMNH" pageId="12" pageNumber="229" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54EB8871FF9DF750003E112D4F88CF0F" box="[587,669,360,379]" pageId="12" pageNumber="229" type="paratype">Paratypes</typeStatus>
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<collectionCode id="ED41AE16FF9DF75000D5112D4FC1CF0F" box="[672,724,360,379]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="229" type="Museum">WAM</collectionCode>
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<accessionNumber id="9403AB30FF9DF75000AD112D4E0CCF0F" box="[728,793,360,379]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41488" pageId="12" pageNumber="229" type="EnaNcbi">S41488</accessionNumber>
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(3 preserved specimens),
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<collectionCode id="ED41AE16FF9DF75003F511C54CABCFE7" box="[384,446,384,403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="229" type="Museum">FMNH</collectionCode>
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</materialsCitation>
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219253 (3 preserved specimens).
|
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</paragraph>
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<paragraph id="8BEF36D3FF9DF75002D311F04F44CC64" blockId="12.[166,794,437,528]" pageId="12" pageNumber="229">
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<emphasis id="B924EAC1FF9DF75002D311F04CC3CFBC" bold="true" box="[166,470,437,456]" pageId="12" pageNumber="229">Additional, non-type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B383C8EFF9DF750039011F04F58CC64" accessionNumber="S41487" collectionCode="WAM" county="northwestern Kimberley" latitude="-14.098611" location="Admiralty Gulf" longLatPrecision="21" longitude="125.747215" municipality="Bonaparte Archipelago" pageId="12" pageNumber="229" specimenCode="FMNH 219250" specimenCount="1" stateProvince="Western Australia">
|
||
AMS C463727,
|
||
<specimenCode id="DBF69EA8FF9DF75000F111F04E03CFBC" box="[644,790,437,456]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="229" type="Museum">FMNH 219250</specimenCode>
|
||
,
|
||
<collectionCode id="ED41AE16FF9DF75002D311884DCFCF94" box="[166,218,461,480]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="229" type="Museum">WAM</collectionCode>
|
||
S41487 (
|
||
<collectingRegion id="4994F831FF9DF750034611884CC1CF94" box="[307,468,461,480]" country="Australia" name="Western Australia" pageId="12" pageNumber="229">Western Australia</collectingRegion>
|
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,
|
||
<collectingCounty id="628E4E5FFF9DF75003AA11884FA0CF94" box="[479,693,461,480]" pageId="12" pageNumber="229">northwestern Kimberley</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B8BACA9FF9DF75000B411884C07CF8C" pageId="12" pageNumber="229">Bonaparte Archipelago</collectingMunicipality>
|
||
,
|
||
<location id="8E8F6008FF9DF750036D11A04C8BCF8C" LSID="urn:lsid:plazi:treatment:03F987C5FF9DF751035110DF4EB8C9D8:8E8F6008FF9DF750036D11A04C8BCF8C" box="[280,414,485,504]" county="northwestern Kimberley" latitude="-14.098611" longLatPrecision="21" longitude="125.747215" municipality="Bonaparte Archipelago" name="Admiralty Gulf" pageId="12" pageNumber="229" stateProvince="Western Australia">Admiralty Gulf</location>
|
||
,
|
||
<location id="8E8F6008FF9DF75003D311A04F4CCF8C" LSID="urn:lsid:plazi:treatment:03F987C5FF9DF751035110DF4EB8C9D8:8E8F6008FF9DF75003D311A04F4CCF8C" box="[422,601,485,504]" county="northwestern Kimberley" latitude="-14.098611" longLatPrecision="21" longitude="125.747215" municipality="Bonaparte Archipelago" name="Montesquieu Islands" pageId="12" pageNumber="229" stateProvince="Western Australia">Montesquieu Islands</location>
|
||
, unnamed island
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||
<locationDeviation id="9A5D2FF1FF9DF750009F11A04C02CC64" location="Oliver Island" pageId="12" pageNumber="229">SE of Oliver Island</locationDeviation>
|
||
,
|
||
<geoCoordinate id="EE645014FF9DF750035511B84C94CC64" box="[288,385,509,528]" degrees="14" direction="south" minutes="05" orientation="latitude" pageId="12" pageNumber="229" precision="15" seconds="55" value="-14.098611">14°05'55"S</geoCoordinate>
|
||
<geoCoordinate id="EE645014FF9DF75003F311B84CE3CC64" box="[390,502,509,528]" degrees="125" direction="east" minutes="44" orientation="longitude" pageId="12" pageNumber="229" precision="15" seconds="50" value="125.747215">125°44'50"E</geoCoordinate>
|
||
; KC-061)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A6558FF9DF75102D312714EB8C9D8" lastPageId="13" lastPageNumber="230" pageId="12" pageNumber="229" type="etymology">
|
||
<paragraph id="8BEF36D3FF9DF75002D312714CF3CCD7" blockId="12.[166,794,564,675]" pageId="12" pageNumber="229">
|
||
<emphasis id="B924EAC1FF9DF75002D312714C37CC38" bold="true" box="[166,290,564,588]" pageId="12" pageNumber="229">Etymology</emphasis>
|
||
. Named in honour of Norman L. McKenzie, Western Australian Department of Environment and Conservation, in recognition of his promotion of malacological research in
|
||
<collectingRegion id="4994F831FF9DF750035512C94CF4CCD7" box="[288,481,652,675]" country="Australia" name="Western Australia" pageId="12" pageNumber="229">Western Australia</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF36D3FF9DF75002D312824CF5CCAA" blockId="12.[166,480,711,734]" box="[166,480,711,734]" pageId="12" pageNumber="229">Sealing strategy. Rock sealer.</paragraph>
|
||
<paragraph id="8BEF36D3FF9DF75002D313444CD1CB7F" blockId="12.[166,795,769,1291]" pageId="12" pageNumber="229">
|
||
<emphasis id="B924EAC1FF9DF75002D313444DCBCD6D" bold="true" box="[166,222,769,793]" pageId="12" pageNumber="229">Shell</emphasis>
|
||
(
|
||
<figureCitation id="136B2A56FF9DF750028513474C61CD6D" box="[240,372,770,793]" captionStart="Figure 17" captionStartId="13.[121,184,1567,1588]" captionTargetBox="[121,1421,151,1555]" captionTargetId="figure-9@13.[121,1421,151,1555]" captionTargetPageId="13" captionText="Figure 17. SEM photographs of Amplirhagada mckenziei n.sp., paratypes WAM S41488. (A–C) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) first four whorls viewed from above (scale 200 µm); (C) details of axial sculpture on last whorl (scale 100 µm). (D) Jaw (scale 100 µm). (E–F) Radula: (E) close-up of central and inner lateral radular teeth (Scale 10 µm); (E) outer lateral and inner marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239123" httpUri="https://zenodo.org/record/5239123/files/figure.png" pageId="12" pageNumber="229">Fig. 17A–C</figureCitation>
|
||
, Pl. 1.8–9). Broadly conical to almost flat with low to medium spire. Thin to solid, translucent. Periphery rounded. Umbilicus concealed by columellar reflection to 50–100 percent, forming a chink or narrowly winding opening. Background colour brownish horn, ventrally lighter; with diffuse to well marked, thin, yellowish brown sub-sutural and mid-whorl bands that are most conspicuous on last whorl; outer lip colour same as shell; inner lip translucent, whitish. Protoconch c.
|
||
<quantity id="4CA89B36FF9DF75000B213A94E0CCA77" box="[711,793,1004,1027]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="229" unit="mm" value="2.2">2.2 mm</quantity>
|
||
in diameter with 1.5 whorls, sculptured by strong axial elements. Teleoconch with coarse axial lirae; curved when viewed from above; rounded in cross-section with irregular spacing; spaces equal to thickness of lirae; distributed evenly across shell and across whorl diameter, reduced underneath suture. Angle of aperture 30°; outer lip rounded, sharp to moderately thick, slightly expanded, not or slightly reflected; basal node absent or weak; palatal node absent. Parietal wall of inner lip inconspicuous.
|
||
</paragraph>
|
||
<paragraph id="8BEF36D3FF9DF75002D3156A4FACCBF5" blockId="12.[166,794,1327,1409]" pageId="12" pageNumber="229">
|
||
<emphasis id="B924EAC1FF9DF75002D3156A4C90CB33" bold="true" box="[166,389,1327,1351]" pageId="12" pageNumber="229">Pallial morphology</emphasis>
|
||
. Pallial cavity short, extending half whorl. Mantle pigmentation consists of blackish spots. Kidney extending more than half of pallial cavity.
|
||
</paragraph>
|
||
<paragraph id="8BEF36D3FF9DF750013210DF4ECACC88" blockId="12.[839,1467,154,764]" pageId="12" pageNumber="229">
|
||
<emphasis id="B924EAC1FF9DF750013210DF4939CEC6" bold="true" box="[839,1068,154,178]" pageId="12" pageNumber="229">Genital morphology</emphasis>
|
||
(
|
||
<figureCitation id="136B2A56FF9DF750064C10DF49A9CEC5" box="[1081,1212,154,177]" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStartId-0="12.[166,229,1912,1933]" captionStartId-1="12.[839,902,1913,1934]" captionTargetBox-0="[175,787,1498,1866]" captionTargetBox-1="[922,1399,789,1877]" captionTargetId-0="figure-295@12.[166,793,1481,1875]" captionTargetId-1="figure-319@12.[839,1466,780,1880]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionText-0="Figure 15. Genitalia of Amplirhagada mckenziei n.sp., paratype WAM S41488 (18 July, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 16. Interior of penial chamber of Amplirhagada mckenziei n.sp., paratype FMNH 219253 (18 July, scale 10 mm). Compare with Fig. 4 for labelling of structures." figureDoi-0="http://doi.org/10.5281/zenodo.5239119" figureDoi-1="http://doi.org/10.5281/zenodo.5239121" httpUri-0="https://zenodo.org/record/5239119/files/figure.png" httpUri-1="https://zenodo.org/record/5239121/files/figure.png" pageId="12" pageNumber="229">Figs. 15–16</figureCitation>
|
||
). Penis straight, slightly longer than anterior part of oviduct. Vas deferens forms simple loop before entering penis. Penial retractor muscle shorter than penis. Penial verge very short (barely visible) to short (<
|
||
<emphasis id="B924EAC1FF9DF75001EC114A4EB4CF69" bold="true" box="[921,929,271,285]" pageId="12" pageNumber="229">
|
||
<superScript id="7C259B9BFF9DF75001EC114A4EB4CF69" attach="none" box="[921,929,271,285]" fontSize="6" pageId="12" pageNumber="229">1</superScript>
|
||
</emphasis>
|
||
⁄
|
||
<emphasis id="B924EAC1FF9DF75001D011534EB8CF50" bold="true" box="[933,941,278,292]" pageId="12" pageNumber="229">
|
||
<subScript id="17D43496FF9DF75001D011534EB8CF50" attach="left" box="[933,941,278,292]" fontSize="6" pageId="12" pageNumber="229">8</subScript>
|
||
</emphasis>
|
||
penial chamber), slender to spatulate with pointed tip. Penial wall pustules small, arranged in rows over entire length of inner penial wall. Main stimulatory pilaster welldifferentiated, relatively thin, cone-shaped, sculptured by ridges with little hooks, on apical portion of inner penial wall. Vas deferens rather straight to undulated, rather thick, entering penial sheath in upper third of penis. Vagina rather thick, tubular to posteriorly inflated. Inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct wide, with smooth longitudinal pilasters inside. Spermathecal head globular to elongately inflated, connected with oviduct by connective tissue; inside entirely smooth with thin wall. Free oviduct more or less straight comprising less than half to about half of length anterior part of oviduct of anterior part of oviduct. Spermoviduct of same length as anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.
|
||
</paragraph>
|
||
<caption id="DF2F665BFF9DF75002D3173D4C6AC9B7" ID-DOI="http://doi.org/10.5281/zenodo.5239119" ID-Zenodo-Dep="5239119" httpUri="https://zenodo.org/record/5239119/files/figure.png" pageId="12" pageNumber="229" startId="12.[166,229,1912,1933]" targetBox="[175,787,1498,1866]" targetPageId="12">
|
||
<paragraph id="8BEF36D3FF9DF75002D3173D4C6AC9B7" blockId="12.[166,794,1912,1987]" pageId="12" pageNumber="229">
|
||
Figure 15. Genitalia of
|
||
<taxonomicName id="4C504D50FF9DF75003E4173D4F96C9F9" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[401,643,1912,1933]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="229" phylum="Mollusca" rank="species" species="mckenziei" status="sp. nov.">
|
||
<emphasis id="B924EAC1FF9DF75003E4173D4F96C9F9" box="[401,643,1912,1933]" italics="true" pageId="12" pageNumber="229">Amplirhagada mckenziei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21757BAFF9DF75000FE173D4FAEC9F9" box="[651,699,1912,1933]" pageId="12" pageNumber="229" rank="species">n.sp.</taxonomicNameLabel>
|
||
, paratype WAM
|
||
<accessionNumber id="9403AB30FF9DF750029C17D64C27C9DC" box="[233,306,1939,1960]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41488" pageId="12" pageNumber="229" type="EnaNcbi">S41488</accessionNumber>
|
||
(18 July, scale 10 mm). Compare with Fig. 3 for labelling of structures.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2F665BFF9DF7500132173C49A5C9B7" ID-DOI="http://doi.org/10.5281/zenodo.5239121" ID-Zenodo-Dep="5239121" httpUri="https://zenodo.org/record/5239121/files/figure.png" pageId="12" pageNumber="229" startId="12.[839,902,1913,1934]" targetBox="[922,1399,789,1877]" targetPageId="12">
|
||
<paragraph id="8BEF36D3FF9DF7500132173C49A5C9B7" blockId="12.[839,1467,1913,1987]" pageId="12" pageNumber="229">
|
||
Figure 16. Interior of penial chamber of
|
||
<taxonomicName id="4C504D50FF9DF75006B9173C48AEC9FA" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1228,1467,1913,1934]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="229" phylum="Mollusca" rank="species" species="mckenziei" status="sp. nov.">
|
||
<emphasis id="B924EAC1FF9DF75006B9173C48AEC9FA" box="[1228,1467,1913,1934]" italics="true" pageId="12" pageNumber="229">Amplirhagada mckenziei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21757BAFF9DF750013217D64E62C9DC" box="[839,887,1939,1960]" pageId="12" pageNumber="229" rank="species">n.sp.</taxonomicNameLabel>
|
||
, paratype FMNH 219253 (18 July, scale 10 mm). Compare with Fig. 4 for labelling of structures.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2F665BFF9CF751020C165A4C87C8F0" ID-DOI="http://doi.org/10.5281/zenodo.5239123" ID-Zenodo-Dep="5239123" httpUri="https://zenodo.org/record/5239123/files/figure.png" pageId="13" pageNumber="230" startId="13.[121,184,1567,1588]" targetBox="[121,1421,151,1555]" targetPageId="13">
|
||
<paragraph id="8BEF36D3FF9CF751020C165A4C87C8F0" blockId="13.[121,1422,1567,1668]" pageId="13" pageNumber="230">
|
||
Figure 17. SEM photographs of
|
||
<taxonomicName id="4C504D50FF9CF75103CC165A4FBCC840" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[441,681,1567,1588]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="230" phylum="Mollusca" rank="species" species="mckenziei" status="sp. nov.">
|
||
<emphasis id="B924EAC1FF9CF75103CC165A4FBCC840" box="[441,681,1567,1588]" italics="true" pageId="13" pageNumber="230">Amplirhagada mckenziei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21757BAFF9CF75100C4165A4FF5C840" box="[689,736,1567,1588]" pageId="13" pageNumber="230" rank="species">n.sp.</taxonomicNameLabel>
|
||
, paratypes WAM
|
||
<accessionNumber id="9403AB30FF9CF75101E4165A4EC8C840" box="[913,989,1567,1588]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41488" pageId="13" pageNumber="230" type="EnaNcbi">S41488</accessionNumber>
|
||
. (
|
||
<emphasis id="B924EAC1FF9CF7510185165A490EC840" box="[1008,1051,1567,1588]" italics="true" pageId="13" pageNumber="230">A–C</emphasis>
|
||
) Shell: (
|
||
<emphasis id="B924EAC1FF9CF7510606165A4997C840" box="[1139,1154,1567,1588]" italics="true" pageId="13" pageNumber="230">A</emphasis>
|
||
) apical whorl viewed from above (scale 200 µm); (
|
||
<emphasis id="B924EAC1FF9CF751032D167F4C72C83B" box="[344,359,1594,1615]" italics="true" pageId="13" pageNumber="230">B</emphasis>
|
||
) first four whorls viewed from above (scale 200 µm); (
|
||
<emphasis id="B924EAC1FF9CF751011B167F4E6BC83B" box="[878,894,1594,1615]" italics="true" pageId="13" pageNumber="230">C</emphasis>
|
||
) details of axial sculpture on last whorl (scale 100 µm). (
|
||
<emphasis id="B924EAC1FF9CF75102F416104D87C81E" box="[129,146,1621,1642]" italics="true" pageId="13" pageNumber="230">D</emphasis>
|
||
) Jaw (scale 100 µm). (
|
||
<emphasis id="B924EAC1FF9CF751030416104C8EC81E" box="[369,411,1621,1642]" italics="true" pageId="13" pageNumber="230">E–F</emphasis>
|
||
) Radula: (
|
||
<emphasis id="B924EAC1FF9CF751007716104F04C81E" box="[514,529,1621,1642]" italics="true" pageId="13" pageNumber="230">E</emphasis>
|
||
) close-up of central and inner lateral radular teeth (Scale 10 µm); (
|
||
<emphasis id="B924EAC1FF9CF75106EE161049BFC81E" box="[1179,1194,1621,1642]" italics="true" pageId="13" pageNumber="230">E</emphasis>
|
||
) outer lateral and inner marginal teeth (scale 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF36D3FF9CF751020C16824FA0C9BD" blockId="13.[121,749,1735,1993]" pageId="13" pageNumber="230">
|
||
<emphasis id="B924EAC1FF9CF751020C16824C65C8AB" bold="true" box="[121,368,1735,1759]" pageId="13" pageNumber="230">Radular morphology</emphasis>
|
||
(
|
||
<figureCitation id="136B2A56FF9CF75103F7168D4F13C8AB" box="[386,518,1736,1759]" captionStart="Figure 17" captionStartId="13.[121,184,1567,1588]" captionTargetBox="[121,1421,151,1555]" captionTargetId="figure-9@13.[121,1421,151,1555]" captionTargetPageId="13" captionText="Figure 17. SEM photographs of Amplirhagada mckenziei n.sp., paratypes WAM S41488. (A–C) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) first four whorls viewed from above (scale 200 µm); (C) details of axial sculpture on last whorl (scale 100 µm). (D) Jaw (scale 100 µm). (E–F) Radula: (E) close-up of central and inner lateral radular teeth (Scale 10 µm); (E) outer lateral and inner marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239123" httpUri="https://zenodo.org/record/5239123/files/figure.png" pageId="13" pageNumber="230">Fig. 17E–F</figureCitation>
|
||
). Rectangular. Tooth formula C+12–18+2–3+18–20. With in average 135±12.5 rows of teeth, 31.2±1.2 rows per mm (n = 2). Central teeth with sharply pointed, triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of tooth; ectocones tiny, endocones vestigial. Marginal teeth with triangular to ovate mesocones; ectocones shorter and narrower than mesocones; endocones reduced in size.
|
||
</paragraph>
|
||
<paragraph id="8BEF36D3FF9CF751016F16824EB8C9D8" blockId="13.[794,1422,1735,1964]" pageId="13" pageNumber="230">
|
||
<emphasis id="B924EAC1FF9CF751016F16824900C8AB" bold="true" box="[794,1045,1735,1759]" pageId="13" pageNumber="230">Comparative remarks</emphasis>
|
||
. The studied museum material was labelled as “
|
||
<taxonomicName id="4C504D50FF9CF75101EA16A04975C888" box="[927,1120,1765,1788]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="230" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B924EAC1FF9CF75101EA16A0492CC888" box="[927,1081,1765,1788]" italics="true" pageId="13" pageNumber="230">Amplirhagada</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
72” by Solem. This species is closely related to
|
||
<taxonomicName id="4C504D50FF9CF7510184174649AAC96E" authorityName="Köhler" authorityYear="2010" box="[1009,1215,1795,1818]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="230" phylum="Mollusca" rank="species" species="montesquieuana">
|
||
<emphasis id="B924EAC1FF9CF7510184174649AAC96E" box="[1009,1215,1795,1818]" italics="true" pageId="13" pageNumber="230">A. montesquieuana</emphasis>
|
||
</taxonomicName>
|
||
(see below) and
|
||
<taxonomicName id="4C504D50FF9CF751070317464E9CC943" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="230" phylum="Mollusca" rank="species" species="combeana">
|
||
<emphasis id="B924EAC1FF9CF751070317464E9CC943" italics="true" pageId="13" pageNumber="230">A. combeana</emphasis>
|
||
</taxonomicName>
|
||
, both inhabiting islands in relative proximity.
|
||
<taxonomicName id="4C504D50FF9CF751016F17784930C920" box="[794,1061,1853,1876]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="230" phylum="Mollusca" rank="species" species="combeana">
|
||
<emphasis id="B924EAC1FF9CF751016F17784930C920" box="[794,1061,1853,1876]" italics="true" pageId="13" pageNumber="230">Amplirhagada combeana</emphasis>
|
||
</taxonomicName>
|
||
is similar in exhibiting a relatively pronounced but irregular axial sculpture but differs in having smaller main stimulatory pilaster, penial verge and no penial wall pustules.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |