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<document id="C357AFC26E17FEC10606E22C93BE9F1D" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FF80F74F01F011AB49C8CFA3" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada descartesana Köhler 2010, n.sp." docType="treatment" docUuid="907DB6AF-C085-451A-A575-BBF57138188D" docUuidSource="ZooBank" docVersion="3" lastPageNumber="236" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="234" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="34E387E56AE4C15E53DB5E58F2830C6B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ACEAC67A2FE6321AE1175199ABB9E663">
<mods:title id="7F50DFDDF37FA40D721F5FB847F8C54E">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="8ECA98D39B2EA4FC61FE8E7AB0D3581D" type="personal">
<mods:role id="CF5BAB3719C59FBDBD0073EF35C9C42A">
<mods:roleTerm id="49F139F5CDBA6DD2A14D942BF200586A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B9A1D13E63703C79E9A813A28FAA6CF">Köhler, Frank</mods:namePart>
</mods:name>
<mods:typeOfResource id="F23AA4C64EB4842A09710A89FDC8FAC1">text</mods:typeOfResource>
<mods:relatedItem id="B7A790DA024CED3099A88B101D654FDC" type="host">
<mods:titleInfo id="73999FECF109455636FB2FE6924B607A">
<mods:title id="2BD74834D2E2A38F89EDE32AFC631F9E">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="3186E1F05D25AFD9C25F6159D5DBF64C">
<mods:date id="1996C99B5082413C0886E16D42946872">2010</mods:date>
<mods:detail id="9596C689BF008E973826A27D252659D5" type="pubDate">
<mods:number id="A1855097BA7AFFBED93FA3ACD9B007B6">2010-11-24</mods:number>
</mods:detail>
<mods:detail id="C256260355850CC5C499C58B03F5F027" type="volume">
<mods:number id="0047009B81CAC18185D1966053734C32">62</mods:number>
</mods:detail>
<mods:detail id="DF481FD4E0436BA057D914DCFD827657" type="issue">
<mods:number id="D6FCCB8708E1B1D08C14C737F29B9308">2</mods:number>
</mods:detail>
<mods:extent id="B324E94D287ED2EA47709ECB815E1511" unit="page">
<mods:start id="D47A3CCE1539140FC430428B2EF8442E">217</mods:start>
<mods:end id="8144F173228A7525B252850409F0C02E">284</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="54339A430FB89B3BE3B2FCBEA1941764">
<mods:url id="5F46EAFA647C809A2C59F02B197EEB51">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
</mods:location>
<mods:classification id="C7D20B8270F3E2B242A4087985EBCAB4">journal article</mods:classification>
<mods:identifier id="77E38D69EF369619762B88F2DADADC86" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="008DB907F63AE13A478F1ACD3D3AB372" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="EFF81E5986C0BE8B469D978A729125E6" type="Zenodo-Dep">5239086</mods:identifier>
<mods:identifier id="AD2513DE7C836F4D1F739E3F80E1D2C2" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
</mods:mods>
<treatment id="03F987C5FF80F74F01F011AB49C8CFA3" ID-DOI="http://doi.org/10.5281/zenodo.5239021" ID-Zenodo-Dep="5239021" LSID="urn:lsid:zoobank.org:act:907DB6AF-C085-451A-A575-BBF57138188D" httpUri="http://treatment.plazi.org/id/03F987C5FF80F74F01F011AB49C8CFA3" lastPageId="19" lastPageNumber="236" pageId="17" pageNumber="234">
<subSubSection id="C34A6558FF80F74D01F011AB4837CC7C" box="[901,1314,494,520]" pageId="17" pageNumber="234" type="nomenclature">
<paragraph id="8BEF36D3FF80F74D01F011AB4837CC7C" blockId="17.[901,1314,494,520]" box="[901,1314,494,520]" pageId="17" pageNumber="234">
<heading id="D0A781BFFF80F74D01F011AB4837CC7C" box="[901,1314,494,520]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="234" reason="2">
<emphasis id="B924EAC1FF80F74D01F011AB4837CC7C" bold="true" box="[901,1314,494,520]" pageId="17" pageNumber="234">
<taxonomicName id="4C504D50FF80F74D01F011AB49CBCC7C" ID-CoL="D7QP" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[901,1246,494,520]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="234" phylum="Mollusca" rank="species" species="descartesana" status="sp. nov.">
<emphasis id="B924EAC1FF80F74D01F011AB49CBCC7C" bold="true" box="[901,1246,494,520]" italics="true" pageId="17" pageNumber="234">Amplirhagada descartesana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF80F74D069311AB4837CC7C" box="[1254,1314,494,520]" pageId="17" pageNumber="234" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF80F74D016F126F4EEFCDE0" pageId="17" pageNumber="234" type="materials_examined">
<paragraph id="8BEF36D3FF80F74D016F126F496ACCED" blockId="17.[794,1421,554,665]" pageId="17" pageNumber="234">
<materialsCitation id="3B383C8EFF80F74D016F126F496ECCED" collectingDate="1988-07-19" collectionCode="V" collectorName="Kessner &amp; A. Longbottom" county="northwestern Kimberley" latitude="-14.173889" location="Institute Islands" longLatPrecision="21" longitude="125.67722" municipality="Admiralty Gulf" pageId="17" pageNumber="234" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FF80F74D016F126F4EA6CC36" bold="true" box="[794,947,554,578]" pageId="17" pageNumber="234">
<typeStatus id="54EB8871FF80F74D016F126F4E41CC36" box="[794,852,554,578]" pageId="17" pageNumber="234">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FF80F74D01B2126F491ECC35" box="[967,1035,554,577]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="17" pageNumber="234">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FF80F74D0650126F49FACC35" box="[1061,1263,554,577]" country="Australia" name="Western Australia" pageId="17" pageNumber="234">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFF80F74D068B126F4E84CC2B" pageId="17" pageNumber="234">northwestern Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FF80F74D01E8120D495BCC2B" box="[925,1102,584,607]" pageId="17" pageNumber="234">Admiralty Gulf</collectingMunicipality>
,
<location id="8E8F6008FF80F74D0628120D4806CC2B" LSID="urn:lsid:plazi:treatment:03F987C5FF80F74F01F011AB49C8CFA3:8E8F6008FF80F74D0628120D4806CC2B" box="[1117,1299,584,607]" county="northwestern Kimberley" latitude="-14.173889" longLatPrecision="21" longitude="125.67722" municipality="Admiralty Gulf" name="Institute Islands" pageId="17" pageNumber="234" stateProvince="Western Australia">Institute Islands</location>
,
<location id="8E8F6008FF80F74D0757120D4E4BCC08" LSID="urn:lsid:plazi:treatment:03F987C5FF80F74F01F011AB49C8CFA3:8E8F6008FF80F74D0757120D4E4BCC08" county="northwestern Kimberley" latitude="-14.173889" longLatPrecision="21" longitude="125.67722" municipality="Admiralty Gulf" name="Descartes Island" pageId="17" pageNumber="234" stateProvince="Western Australia">Descartes Island</location>
,
<geoCoordinate id="EE645014FF80F74D011F12204EF6CC08" box="[874,995,613,636]" degrees="14" direction="south" minutes="10" orientation="latitude" pageId="17" pageNumber="234" precision="15" seconds="26" value="-14.173889">14°10'26&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF80F74D019912204963CC08" box="[1004,1142,613,636]" degrees="125" direction="east" minutes="40" orientation="longitude" pageId="17" pageNumber="234" precision="15" seconds="38" value="125.67722">125°40'38&quot;E</geoCoordinate>
; KC-071 (leg.
<collectionCode id="ED41AE16FF80F74D076C12204839CC08" box="[1305,1324,613,636]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="234" type="Museum">V</collectionCode>
.
<collectorName id="26A55305FF80F74D074D12204898CC08" box="[1336,1421,613,636]" pageId="17" pageNumber="234">Kessner</collectorName>
&amp;
<collectorName id="26A55305FF80F74D014112C74ECECCED" box="[820,987,642,665]" pageId="17" pageNumber="234">A. Longbottom</collectorName>
,
<date id="FFEE1013FF80F74D019012C74966CCED" box="[997,1139,642,665]" pageId="17" pageNumber="234" value="1988-07-19">
<collectingDate id="EFAAE9FBFF80F74D019012C74966CCED" box="[997,1139,642,665]" pageId="17" pageNumber="234" value="1988-07-19">19 July 1988</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<caption id="DF2F665BFF80F74D020C123D4CA4CCB7" ID-DOI="http://doi.org/10.5281/zenodo.5239133" ID-Zenodo-Dep="5239133" httpUri="https://zenodo.org/record/5239133/files/figure.png" pageId="17" pageNumber="234" startId="17.[121,186,632,653]" targetBox="[126,736,149,618]" targetPageId="17">
<paragraph id="8BEF36D3FF80F74D020C123D4CA4CCB7" blockId="17.[121,749,632,707]" pageId="17" pageNumber="234">
Figure 22. Genitalia of
<taxonomicName id="4C504D50FF80F74D0305123D4FB8CCF9" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[368,685,632,653]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="234" phylum="Mollusca" rank="species" species="montesquieuana" status="sp. nov.">
<emphasis id="B924EAC1FF80F74D0305123D4FB8CCF9" box="[368,685,632,653]" italics="true" pageId="17" pageNumber="234">Amplirhagada montesquieuana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF80F74D00C2123D4FFDCCF9" box="[695,744,632,653]" pageId="17" pageNumber="234" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463748 (28 August, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FF80F74D016F12FE49D0CD32" blockId="17.[794,1422,699,838]" pageId="17" pageNumber="234">
<emphasis id="B924EAC1FF80F74D016F12FE4EB4CCBA" bold="true" box="[794,929,699,718]" pageId="17" pageNumber="234">Type material</emphasis>
.
<materialsCitation id="3B383C8EFF80F74D01C512FE49FACCBA" accessionNumber="S34607" box="[944,1263,699,718]" collectionCode="WAM" pageId="17" pageNumber="234" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FF80F74D01C512FE4916CCBA" box="[944,1027,699,718]" pageId="17" pageNumber="234" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FF80F74D067E12FE4955CCBA" box="[1035,1088,699,718]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="234" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF80F74D063D12FE4999CCBA" box="[1096,1164,699,718]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34607" pageId="17" pageNumber="234" type="EnaNcbi">S34607</accessionNumber>
(Pl. 1.12)
</materialsCitation>
.
<materialsCitation id="3B383C8EFF80F74D068F12FE498DCC92" accessionNumber="S41491" collectionCode="WAM" pageId="17" pageNumber="234" specimenCount="2" typeStatus="paratype">
<typeStatus id="54EB8871FF80F74D068F12FE4845CCBA" box="[1274,1360,699,718]" pageId="17" pageNumber="234" type="paratype">Paratypes</typeStatus>
<collectionCode id="ED41AE16FF80F74D072D12FE4898CCBA" box="[1368,1421,699,718]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="234" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF80F74D016F12964E4ECC92" box="[794,859,723,742]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41491" pageId="17" pageNumber="234" type="EnaNcbi">S41491</accessionNumber>
(4 preserved specimens, as
<typeStatus id="54EB8871FF80F74D063112964987CC92" box="[1092,1170,723,742]" pageId="17" pageNumber="234" type="holotype">holotype</typeStatus>
)
</materialsCitation>
,
<materialsCitation id="3B383C8EFF80F74D06EA12964EF4CC8A" collectionCode="FMNH" pageId="17" pageNumber="234" specimenCode="FMNH 219276" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBF69EA8FF80F74D06EA12964834CC92" box="[1183,1313,723,742]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="234" type="Museum">FMNH 219276</specimenCode>
(5 preserved specimens, as
<typeStatus id="54EB8871FF80F74D01FA12AE4ECECC8A" box="[911,987,747,766]" pageId="17" pageNumber="234" type="holotype">holotype</typeStatus>
)
</materialsCitation>
,
<materialsCitation id="3B383C8EFF80F74D019D12AE4844CD62" collectionCode="FMNH" latitude="-14.179167" location="Descartes Island" longLatPrecision="21" longitude="125.666664" pageId="17" pageNumber="234" specimenCode="FMNH 219272" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBF69EA8FF80F74D019D12AE497DCC8A" box="[1000,1128,747,766]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="234" type="Museum">FMNH 219272</specimenCode>
(16 preserved specimens, unnamed island SW of
<location id="8E8F6008FF80F74D01FA13464908CD62" LSID="urn:lsid:plazi:treatment:03F987C5FF80F74F01F011AB49C8CFA3:8E8F6008FF80F74D01FA13464908CD62" box="[911,1053,771,790]" latitude="-14.179167" longLatPrecision="21" longitude="125.666664" name="Descartes Island" pageId="17" pageNumber="234">Descartes Island</location>
,
<geoCoordinate id="EE645014FF80F74D065313464992CD62" box="[1062,1159,771,790]" degrees="14" direction="south" minutes="10" orientation="latitude" pageId="17" pageNumber="234" precision="15" seconds="45" value="-14.179167">14°10'45&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF80F74D06F8134649E9CD62" box="[1165,1276,771,790]" degrees="125" direction="east" minutes="40" orientation="longitude" pageId="17" pageNumber="234" precision="15" seconds="00" value="125.666664">125°40'00&quot;E</geoCoordinate>
, KC/070)
</materialsCitation>
,
<materialsCitation id="3B383C8EFF80F74D072C13464812CD5A" accessionNumber="S41490" collectionCode="WAM" pageId="17" pageNumber="234" specimenCode="FMNH 219272" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED41AE16FF80F74D072C13464898CD62" box="[1369,1421,771,790]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="234" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF80F74D016F135E4E4ECD5A" box="[794,859,795,814]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41490" pageId="17" pageNumber="234" type="EnaNcbi">S41490</accessionNumber>
(10 preserved specimens, same as
<specimenCode id="DBF69EA8FF80F74D0608135E4814CD5A" box="[1149,1281,795,814]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="234" type="Museum">FMNH 219272</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B383C8EFF80F74D0778135E49D4CD32" collectionCode="FMNH" pageId="17" pageNumber="234" specimenCode="FMNH 219272" specimenCount="1" typeStatus="paratype">
AMS C463749 (7 preserved specimens, same as
<specimenCode id="DBF69EA8FF80F74D0640137649A9CD32" box="[1077,1212,819,838]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="234" type="Museum">FMNH 219272</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FF80F74D016F132D4EEFCDE0" blockId="17.[794,1422,872,916]" pageId="17" pageNumber="234">
<emphasis id="B924EAC1FF80F74D016F132D4953CD0F" bold="true" box="[794,1094,872,891]" pageId="17" pageNumber="234">Additional, non-type material</emphasis>
.
<materialsCitation id="3B383C8EFF80F74D0620132C4EE3CDE0" accessionNumber="S41452" collectionCode="WAM" latitude="-14.163889" location="Descartes Island" longLatPrecision="21" longitude="125.67972" pageId="17" pageNumber="234" specimenCount="1">
<collectionCode id="ED41AE16FF80F74D0620132C4999CD08" box="[1109,1164,873,892]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="234" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF80F74D06E1132C49CFCD08" box="[1172,1242,873,892]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41452" pageId="17" pageNumber="234" type="EnaNcbi">S41452</accessionNumber>
(
<location id="8E8F6008FF80F74D069F132C489FCD08" LSID="urn:lsid:plazi:treatment:03F987C5FF80F74F01F011AB49C8CFA3:8E8F6008FF80F74D069F132C489FCD08" box="[1258,1418,873,892]" latitude="-14.163889" longLatPrecision="21" longitude="125.67972" name="Descartes Island" pageId="17" pageNumber="234">Descartes Island</location>
,
<geoCoordinate id="EE645014FF80F74D016F13C44E93CDE0" box="[794,902,897,916]" degrees="125" direction="east" minutes="40" orientation="longitude" pageId="17" pageNumber="234" precision="15" seconds="47" value="125.67972">125°40'47&quot;E</geoCoordinate>
<geoCoordinate id="EE645014FF80F74D01F913C44EE5CDE0" box="[908,1008,897,916]" degrees="14" direction="south" minutes="09" orientation="latitude" pageId="17" pageNumber="234" precision="15" seconds="50" value="-14.163889">14°09'50&quot;S</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF80F74F016F13F249C8CFA3" lastPageId="19" lastPageNumber="236" pageId="17" pageNumber="234" type="etymology">
<paragraph id="8BEF36D3FF80F74D016F13F24EAFCD98" blockId="17.[794,1421,951,1004]" pageId="17" pageNumber="234">
<emphasis id="B924EAC1FF80F74D016F13F24E83CDBB" bold="true" box="[794,918,951,975]" pageId="17" pageNumber="234">Etymology</emphasis>
. In reference to Descartes Island, where this species occurs.
</paragraph>
<paragraph id="8BEF36D3FF80F74D016F144A49ABCB88" blockId="17.[794,1422,1039,1532]" pageId="17" pageNumber="234">
<emphasis id="B924EAC1FF80F74D016F144A4E44CA53" bold="true" box="[794,849,1039,1063]" pageId="17" pageNumber="234">Shell</emphasis>
(
<figureCitation id="136B2A56FF80F74D012914554ECFCA53" box="[860,986,1040,1063]" captionStart="Figure 24" captionStartId="18.[166,229,1933,1954]" captionTargetBox="[167,1467,151,1926]" captionTargetId="figure-8@18.[167,1467,151,1926]" captionTargetPageId="18" captionText="Figure 24. SEM photographs of Amplirhagada descartesana n.sp., paratype AMS C463749. (A–D) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on first four whorls viewed from above (scale 1 mm); (C) sculpture, lateral view (scale 100 µm); (D) close-up of sculpture, obliquely from above (scale 100 µm). (E–G) Radula: (E) central and lateral teeth (Scale 20 µm); (F) close-up of central and inner lateral teeth (scale 10 µm); (G) outer lateral and inner marginal teeth (scale 10 µm). (H) Jaw (scale 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239137" httpUri="https://zenodo.org/record/5239137/files/figure.png" pageId="17" pageNumber="234">Fig. 24A–D</figureCitation>
, Pl. 1.12). Semi-globose to broadly conical with medium high spire; thin (translucent) to solid. Periphery evenly rounded to slightly angulate; upper and basal sectors of whorls rounded. Umbilicus forming a chink to narrowly winding opening, 30–100 percent concealed by columellar reflection. Background colour brownish horn; uniform or banded; if present sub-sutural and mid-whorl bands diffuse, thin, brown, on last whorl(s) only; ventral colour, outer and inner lip colour whitish to cream. Protoconch c.
<quantity id="4CA89B36FF80F74D076A14BF4864CB65" box="[1311,1393,1274,1297]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="17" pageNumber="234" unit="mm" value="2.8">2.8 mm</quantity>
in diameter with 1.7 whorls, almost smooth, sculptured by faint axial lirae. Teleoconch sculptured by coarse lirae, rounded in cross-section, regularly spaced, spaces equal to thickness of lirae, evenly distributed across shell and whorl diameter even, reduced underneath suture.Angle of aperture 45°, outer lip rounded, moderately thick, slightly expanded, slightly reflected; basal node absent or weak, palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<paragraph id="8BEF36D3FF80F74D016F165A4822C805" blockId="17.[794,1421,1567,1649]" pageId="17" pageNumber="234">
<emphasis id="B924EAC1FF80F74D016F165A4912C843" bold="true" box="[794,1031,1567,1591]" pageId="17" pageNumber="234">Pallial morphology</emphasis>
. Pallial cavity moderately deep, extending
<emphasis id="B924EAC1FF80F74D01FE16784E86C83F" bold="true" box="[907,915,1597,1611]" pageId="17" pageNumber="234">
<superScript id="7C259B9BFF80F74D01FE16784E86C83F" attach="right" box="[907,915,1597,1611]" fontSize="6" pageId="17" pageNumber="234">3</superScript>
</emphasis>
⁄
<emphasis id="B924EAC1FF80F74D01ED16014EB5C826" bold="true" box="[920,928,1604,1618]" pageId="17" pageNumber="234">
<subScript id="17D43496FF80F74D01ED16014EB5C826" attach="left" box="[920,928,1604,1618]" fontSize="6" pageId="17" pageNumber="234">4</subScript>
</emphasis>
whorl. Mottled pigmentation on mantle dark grey. Kidney extending about half of pallial cavity.
</paragraph>
<caption id="DF2F665BFF80F74D020C16184CC8C806" ID-DOI="http://doi.org/10.5281/zenodo.5239135" ID-Zenodo-Dep="5239135" box="[121,477,1629,1650]" httpUri="https://zenodo.org/record/5239135/files/figure.png" pageId="17" pageNumber="234" startId="17.[121,184,1629,1650]" targetBox="[121,728,734,1757]" targetPageId="17">
<paragraph id="8BEF36D3FF80F74D020C16184CC8C806" blockId="17.[121,477,1629,1757]" box="[121,477,1629,1650]" pageId="17" pageNumber="234">Figure 23. Interior of penial chamber</paragraph>
</caption>
<paragraph id="8BEF36D3FF80F74F016F16D04C8ECDCD" blockId="17.[794,1422,1685,2001]" lastBlockId="19.[121,749,695,953]" lastPageId="19" lastPageNumber="236" pageId="17" pageNumber="234">
<emphasis id="B924EAC1FF80F74D016F16D04905C8D9" bold="true" box="[794,1040,1685,1709]" pageId="17" pageNumber="234">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FF80F74D065016D049A3C8D8" box="[1061,1206,1685,1708]" captionStart-0="Figure 25" captionStart-1="Figure 26" captionStartId-0="19.[121,183,572,593]" captionStartId-1="19.[121,193,1919,1940]" captionTargetBox-0="[130,739,144,556]" captionTargetBox-1="[134,744,1270,1902]" captionTargetId-0="figure-9@19.[121,748,134,564]" captionTargetId-1="figure-33@19.[121,748,1262,1910]" captionTargetPageId-0="19" captionTargetPageId-1="19" captionText-0="Figure 25. Genitalia of Amplirhagada descartesana n.sp., paratype WAM S41491 (20 July, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 26. Interior of penial chamber of Amplirhagada descartesana n.sp., paratype FMNH 219276 (20 July, scale 5 mm). Compare with Fig. 4 for labelling of structures." figureDoi-0="http://doi.org/10.5281/zenodo.5239139" figureDoi-1="http://doi.org/10.5281/zenodo.5239141" httpUri-0="https://zenodo.org/record/5239139/files/figure.png" httpUri-1="https://zenodo.org/record/5239141/files/figure.png" pageId="17" pageNumber="234">Figs. 25–26</figureCitation>
). Penis straight to slightly curved, longer than anterior part of oviduct. Vas deferens forms simple loop before entering penis. Penial retractor muscle shorter than penis. Penial verge short (&lt;
<emphasis id="B924EAC1FF80F74D0147174F4E2FC96C" bold="true" box="[818,826,1802,1816]" pageId="17" pageNumber="234">
<superScript id="7C259B9BFF80F74D0147174F4E2FC96C" attach="none" box="[818,826,1802,1816]" fontSize="6" pageId="17" pageNumber="234">1</superScript>
</emphasis>
⁄
<emphasis id="B924EAC1FF80F74D014A17544E52C96B" bold="true" box="[831,839,1809,1823]" pageId="17" pageNumber="234">
<subScript id="17D43496FF80F74D014A17544E52C96B" attach="left" box="[831,839,1809,1823]" fontSize="6" pageId="17" pageNumber="234">4</subScript>
</emphasis>
penial chamber), slender to spatulate, with pointed tip. Penial wall pustules present, of average size, slightly elongated, arranged in rows across entire length of inner penial wall. Main stimulatory pilaster well-differentiated, large, cone-shaped, sculptured by smooth horizontal ridges, comprising entire length of penial chamber. Vas deferens entering penial sheath in upper third. Vagina comparatively short to moderately long, posteriorly inflated. Inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct comparatively wide, internally with smooth longitudinal pilasters. Spermathecal head elongately inflated, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprising about half of anterior part of oviduct, rather straight. Spermoviduct clearly longer than anterior part of oviduct. Talon embedded in albumen gland close to anterior end.
</paragraph>
<caption id="DF2F665BFF83F74E02D317C8484FC986" ID-DOI="http://doi.org/10.5281/zenodo.5239137" ID-Zenodo-Dep="5239137" httpUri="https://zenodo.org/record/5239137/files/figure.png" pageId="18" pageNumber="235" startId="18.[166,229,1933,1954]" targetBox="[167,1467,151,1926]" targetPageId="18">
<paragraph id="8BEF36D3FF83F74E02D317C8484FC986" blockId="18.[166,1467,1933,2034]" pageId="18" pageNumber="235">
Figure 24. SEM photographs of
<taxonomicName id="4C504D50FF83F74E03A917C84FF9C9D6" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[476,748,1933,1954]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="235" phylum="Mollusca" rank="species" species="descartesana" status="sp. nov.">
<emphasis id="B924EAC1FF83F74E03A917C84FF9C9D6" box="[476,748,1933,1954]" italics="true" pageId="18" pageNumber="235">Amplirhagada descartesana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF83F74E008417C84E34C9D6" box="[753,801,1933,1954]" pageId="18" pageNumber="235" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463749. (
<emphasis id="B924EAC1FF83F74E065D17C84941C9D6" box="[1064,1108,1933,1954]" italics="true" pageId="18" pageNumber="235">A–D</emphasis>
) Shell: (
<emphasis id="B924EAC1FF83F74E06D217C849A3C9D6" box="[1191,1206,1933,1954]" italics="true" pageId="18" pageNumber="235">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF83F74E03F317ED4C80C9C9" box="[390,405,1960,1981]" italics="true" pageId="18" pageNumber="235">B</emphasis>
) sculpture on first four whorls viewed from above (scale 1 mm); (
<emphasis id="B924EAC1FF83F74E067C17ED490CC9C9" box="[1033,1049,1960,1981]" italics="true" pageId="18" pageNumber="235">C</emphasis>
) sculpture, lateral view (scale 100 µm); (
<emphasis id="B924EAC1FF83F74E07D417ED48A7C9C9" box="[1441,1458,1960,1981]" italics="true" pageId="18" pageNumber="235">D</emphasis>
) close-up of sculpture, obliquely from above (scale 100 µm). (
<emphasis id="B924EAC1FF83F74E008C17864E31C9AC" box="[761,804,1987,2008]" italics="true" pageId="18" pageNumber="235">E–G</emphasis>
) Radula: (
<emphasis id="B924EAC1FF83F74E01F817864E89C9AC" box="[909,924,1987,2008]" italics="true" pageId="18" pageNumber="235">E</emphasis>
) central and lateral teeth (Scale 20 µm); (
<emphasis id="B924EAC1FF83F74E074417864855C9AC" box="[1329,1344,1987,2008]" italics="true" pageId="18" pageNumber="235">F</emphasis>
) close-up of central and inner lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FF83F74E002E17984F79C986" box="[603,620,2013,2034]" italics="true" pageId="18" pageNumber="235">G</emphasis>
) outer lateral and inner marginal teeth (scale 10 µm). (
<emphasis id="B924EAC1FF83F74E060F1798499EC986" box="[1146,1163,2013,2034]" italics="true" pageId="18" pageNumber="235">H</emphasis>
) Jaw (scale 100 µm).
</paragraph>
</caption>
<caption id="DF2F665BFF82F74F020C12794C44CCF2" ID-DOI="http://doi.org/10.5281/zenodo.5239139" ID-Zenodo-Dep="5239139" httpUri="https://zenodo.org/record/5239139/files/figure.png" pageId="19" pageNumber="236" startId="19.[121,183,572,593]" targetBox="[130,739,144,556]" targetPageId="19">
<paragraph id="8BEF36D3FF82F74F020C12794C44CCF2" blockId="19.[121,748,572,646]" pageId="19" pageNumber="236">
Figure 25. Genitalia of
<taxonomicName id="4C504D50FF82F74F032712794F4BCC25" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[338,606,572,593]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="descartesana" status="sp. nov.">
<emphasis id="B924EAC1FF82F74F032712794F4BCC25" box="[338,606,572,593]" italics="true" pageId="19" pageNumber="236">Amplirhagada descartesana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF82F74F001612794F87CC25" box="[611,658,572,593]" pageId="19" pageNumber="236" rank="species">n.sp.</taxonomicNameLabel>
, paratype WAM
<accessionNumber id="9403AB30FF82F74F02CE12124C10CC18" box="[187,261,599,620]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41491" pageId="19" pageNumber="236" type="EnaNcbi">S41491</accessionNumber>
(20 July, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FF82F74F020C13994C0BCA8F" blockId="19.[121,749,988,1275]" pageId="19" pageNumber="236">
<emphasis id="B924EAC1FF82F74F020C13994C7BCD80" bold="true" box="[121,366,988,1012]" pageId="19" pageNumber="236">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FF82F74F03F513994F12CD87" box="[384,519,988,1011]" captionStart="Figure 24" captionStartId="18.[166,229,1933,1954]" captionTargetBox="[167,1467,151,1926]" captionTargetId="figure-8@18.[167,1467,151,1926]" captionTargetPageId="18" captionText="Figure 24. SEM photographs of Amplirhagada descartesana n.sp., paratype AMS C463749. (A–D) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on first four whorls viewed from above (scale 1 mm); (C) sculpture, lateral view (scale 100 µm); (D) close-up of sculpture, obliquely from above (scale 100 µm). (E–G) Radula: (E) central and lateral teeth (Scale 20 µm); (F) close-up of central and inner lateral teeth (scale 10 µm); (G) outer lateral and inner marginal teeth (scale 10 µm). (H) Jaw (scale 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239137" httpUri="https://zenodo.org/record/5239137/files/figure.png" pageId="19" pageNumber="236">Fig. 24E–G</figureCitation>
). Rectangular. Tooth formula C+16–17+3–4+18–20. In average with 139±5.5 rows of teeth, 31.4±0.2 rows per mm (n = 2). Central teeth with bluntly pointed, triangular mesocones, shorter than base of tooth; ectocones small. Lateral teeth with bluntly pointed, triangular mesocones, length equal to base of tooth; ectocones small, endocones vestigial. Marginal teeth with elongate mesocones; ectocones shorter and narrower than mesocones, divided into two denticles; endocones smaller than ectocones.
</paragraph>
<paragraph id="8BEF36D3FF82F74F016F10DF49C8CFA3" blockId="19.[794,1422,154,471]" pageId="19" pageNumber="236">
<emphasis id="B924EAC1FF82F74F016F10DF4937CEC6" bold="true" box="[794,1058,154,178]" pageId="19" pageNumber="236">Comparative remarks</emphasis>
. The present material has been labelled as “
<taxonomicName id="4C504D50FF82F74F01D410FD4977CEBB" box="[929,1122,184,207]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FF82F74F01D410FD492ECEBB" box="[929,1083,184,207]" italics="true" pageId="19" pageNumber="236">Amplirhagada</emphasis>
sp.
</taxonomicName>
74” by Solem. The species is most readily distinguished from other species from the Montesquieu Islands by its more turreted shell. Similar to
<taxonomicName id="4C504D50FF82F74F070310B64EDDCF53" authorityName="Köhler" authorityYear="2010" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="montesquieuana">
<emphasis id="B924EAC1FF82F74F070310B64EDDCF53" italics="true" pageId="19" pageNumber="236">A. montesquieuana</emphasis>
</taxonomicName>
, its axial sculpture is more regular than that of
<taxonomicName id="4C504D50FF82F74F014011684ED5CF30" box="[821,960,301,324]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="combeana">
<emphasis id="B924EAC1FF82F74F014011684ED5CF30" box="[821,960,301,324]" italics="true" pageId="19" pageNumber="236">A. combeana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FF82F74F018411684963CF30" authorityName="Köhler" authorityYear="2010" box="[1009,1142,301,324]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="mckenziei">
<emphasis id="B924EAC1FF82F74F018411684963CF30" box="[1009,1142,301,324]" italics="true" pageId="19" pageNumber="236">A. mckenziei</emphasis>
</taxonomicName>
. From
<taxonomicName id="4C504D50FF82F74F06B711684898CF30" authorityName="Köhler" authorityYear="2010" box="[1218,1421,301,324]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="montesquieuana">
<emphasis id="B924EAC1FF82F74F06B711684898CF30" box="[1218,1421,301,324]" italics="true" pageId="19" pageNumber="236">A. montesquieuana</emphasis>
</taxonomicName>
it differs by its larger size and more conical shape. The main stimulatory pilaster is similar to that in
<taxonomicName id="4C504D50FF82F74F06B5112D4898CF0B" authorityName="Köhler" authorityYear="2010" box="[1216,1421,360,383]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="montesquieuana">
<emphasis id="B924EAC1FF82F74F06B5112D4898CF0B" box="[1216,1421,360,383]" italics="true" pageId="19" pageNumber="236">A. montesquieuana</emphasis>
</taxonomicName>
in shape but lacks “hooked toes”; broader than pilaster in
<taxonomicName id="4C504D50FF82F74F016F11E64E9FCFCE" authorityName="Köhler" authorityYear="2010" box="[794,906,419,442]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="ponderi">
<emphasis id="B924EAC1FF82F74F016F11E64E9FCFCE" box="[794,906,419,442]" italics="true" pageId="19" pageNumber="236">A. ponderi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FF82F74F01C811E64957CFCE" authorityName="Köhler" authorityYear="2010" box="[957,1090,419,442]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="mckenziei">
<emphasis id="B924EAC1FF82F74F01C811E64957CFCE" box="[957,1090,419,442]" italics="true" pageId="19" pageNumber="236">A. mckenziei</emphasis>
</taxonomicName>
; penial wall pustulation differs markedly from that in
<taxonomicName id="4C504D50FF82F74F067C118549C3CFA3" authorityName="Köhler" authorityYear="2010" box="[1033,1238,448,471]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="236" phylum="Mollusca" rank="species" species="montesquieuana">
<emphasis id="B924EAC1FF82F74F067C118549C3CFA3" box="[1033,1238,448,471]" italics="true" pageId="19" pageNumber="236">A. montesquieuana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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