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<document id="AE38386F6776390DD33619376E5A7E7F" ID-CLB-Dataset="46281" ID-DOI="10.11646/zootaxa.3774.3.5" ID-GBIF-Dataset="72c58303-3689-46f0-906c-9ece69e683da" ID-ISSN="1175-5326" ID-Zenodo-Dep="229649" ID-ZooBank="7DC18AE1-A029-4876-9EEC-CC23C08EA647" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460393102248" checkinUser="plazi" docAuthor="Artigas, Jorge N. &amp; Vieira, Rodrigo" docDate="2014" docId="03F987B3FFAB5A1AFF7AE8E2FEE8F899" docLanguage="en" docName="zt03774p288.pdf" docOrigin="Zootaxa 3774 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Papaverellus Artigas &amp; Vieira, gen. nov." docType="treatment" docVersion="9" lastPageNumber="283" masterDocId="FFC0FFCBFFAA5A1BFFEDE83CFFCFFFB4" masterDocTitle="New genus and species of Neotropical robber flies (Diptera, Asilidae, Asilinae)" masterLastPageNumber="288" masterPageNumber="282" pageNumber="283" updateTime="1698298429581" updateUser="ExternalLinkService">
<mods:mods id="FE1B8E91F8F80EE62489AE0669A190A8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="38E8FC3A1963AA82BD41587DF1ADA31E">
<mods:title id="8F9B7FF641C76C060742628F100B205A">New genus and species of Neotropical robber flies (Diptera, Asilidae, Asilinae)</mods:title>
</mods:titleInfo>
<mods:name id="AB811F304B1EF6746709BE65A1676593" type="personal">
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<mods:namePart id="4CE49C28BBFCCBD7AA5B4DCB27C0F3E5">Artigas, Jorge N.</mods:namePart>
</mods:name>
<mods:name id="B1C986830258A153CB26266E5AA2918A" type="personal">
<mods:role id="4AE5E32BB5EF1681A958B833CA6B1091">
<mods:roleTerm id="2649ABE223B69C6E4FAF4F4933D9C2C9">Author</mods:roleTerm>
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<mods:namePart id="746E42169B48DB7843AAFA6006FC49A8">Vieira, Rodrigo</mods:namePart>
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<mods:typeOfResource id="C0428A262F554322A30D4E0E04806899">text</mods:typeOfResource>
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<mods:title id="D42622874513CB48814264650E3AF8FE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3F22A7575837DD8827C93A337D31410F">
<mods:date id="300DFB85BFC5DB0C83FE568BBC63B894">2014</mods:date>
<mods:detail id="EA17E4773EFC0393EAA64F786CD11877" type="volume">
<mods:number id="7DF18A7E3FEAB86EA2B8DD67BF431A3F">3774</mods:number>
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<mods:detail id="C0BB598AB7C66C85BD6A7A89FD5C3850" type="issue">
<mods:number id="5CDC7B7E1C58CD30F3F5FCAD899E2D4A">3</mods:number>
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<mods:classification id="75CD1EAA3D9E549DAD062DD2EE6DA2B5">journal article</mods:classification>
<mods:identifier id="23A209528DF8858AFA59637C38304944" type="CLB-Dataset">46281</mods:identifier>
<mods:identifier id="8254D1209B2805FA73E20FA1AA96E058" type="DOI">10.11646/zootaxa.3774.3.5</mods:identifier>
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<mods:identifier id="83302AFB9C73A77C1CD85A37CE856143" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F987B3FFAB5A1AFF7AE8E2FEE8F899" ID-DOI="http://doi.org/10.5281/zenodo.6139381" ID-GBIF-Taxon="119417274" ID-Zenodo-Dep="6139381" LSID="urn:lsid:plazi:treatment:03F987B3FFAB5A1AFF7AE8E2FEE8F899" httpUri="http://treatment.plazi.org/id/03F987B3FFAB5A1AFF7AE8E2FEE8F899" lastPageNumber="283" pageId="1" pageNumber="283">
<subSubSection id="C34A652EFFAB5A1AFF7AE8E2FEEFFEAF" pageId="1" pageNumber="283" type="nomenclature">
<paragraph id="8BEF36A5FFAB5A1AFF7AE8E2FDA4FF43" blockId="1.[151,619,222,283]" box="[151,619,222,247]" pageId="1" pageNumber="283">
<heading id="D0A781C9FFAB5A1AFF7AE8E2FDA4FF43" bold="true" box="[151,619,222,247]" fontSize="10" level="2" pageId="1" pageNumber="283" reason="2">
<emphasis id="B924EAB7FFAB5A1AFF7AE8E2FDA4FF43" bold="true" box="[151,619,222,247]" pageId="1" pageNumber="283">
<taxonomicName id="4C504D26FFAB5A1AFF7AE8E2FE34FF43" ID-CoL="89QZF" authority="Artigas &amp; Vieira" authorityName="Artigas &amp; Vieira" box="[151,507,222,247]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B924EAB7FFAB5A1AFF7AE8E2FEE3FF43" bold="true" box="[151,300,222,247]" italics="true" pageId="1" pageNumber="283">Papaverellus</emphasis>
Artigas &amp; Vieira
</taxonomicName>
<taxonomicNameLabel id="A21757CCFFAB5A1AFDEFE8E2FDA4FF43" box="[514,619,222,247]" pageId="1" pageNumber="283" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF7AE93EFEEFFEAF" blockId="1.[151,619,222,283]" box="[151,288,258,283]" pageId="1" pageNumber="283">
(
<figureCitation id="136B2A20FFAB5A1AFF72E93EFED8FEAF" box="[159,279,258,283]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 13" captionStart-2="FIGURES 14 – 18" captionStartId-0="3.[151,264,1952,1975]" captionStartId-1="4.[151,264,1883,1906]" captionStartId-2="5.[151,264,1879,1902]" captionTargetBox-0="[224,1332,195,1891]" captionTargetBox-1="[262,1322,259,1815]" captionTargetBox-2="[221,1357,193,1812]" captionTargetId-0="figure@3.[224,1359,195,1919]" captionTargetId-1="figure@4.[246,1335,193,1839]" captionTargetId-2="figure@5.[215,1373,193,1840]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." captionText-1="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." captionText-2="FIGURES 14 – 18. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Female. 14, Habitus, lateral view; 15, Ovipositor, lateral view; 16, Ovipositor, dorsal view; 17, Ovipositor, ventral view; 18, Genital fork and spermathecae. Abbreviations: ar: arms of genital fork; cap duct: capsular ducts; com duct: common duct; exp duct: expulsory ducts; cerc: cercus; f apod: furcal apodeme; spmth: spermathecae; stVIII: sternite VIII; tVII: tergite VII; tVIII: tergite VIII; tIX + X: tergite IX + X." httpUri-0="https://zenodo.org/record/229650/files/figure.png" httpUri-1="https://zenodo.org/record/229651/files/figure.png" httpUri-2="https://zenodo.org/record/229652/files/figure.png" pageId="1" pageNumber="283">Figs. 1–18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A652EFFAB5A1AFF7AE976FBF7FED7" box="[151,1080,330,355]" pageId="1" pageNumber="283" type="materials_examined">
<paragraph id="8BEF36A5FFAB5A1AFF7AE976FBF7FED7" blockId="1.[151,1436,330,1837]" box="[151,1080,330,355]" pageId="1" pageNumber="283">
<emphasis id="B924EAB7FFAB5A1AFF7AE976FEF6FED7" bold="true" box="[151,313,330,355]" pageId="1" pageNumber="283">
<typeStatus id="54EB8807FFAB5A1AFF7AE976FF1FFED7" box="[151,208,330,355]" pageId="1" pageNumber="283">Type</typeStatus>
–species.
</emphasis>
<taxonomicName id="4C504D26FFAB5A1AFEADE977FC9FFED7" authority="Artigas &amp; Vieira" authorityName="Artigas &amp; Vieira" box="[320,848,330,355]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="aureocingulatus" status="sp. nov.">
<emphasis id="B924EAB7FFAB5A1AFEADE977FD40FED7" box="[320,655,331,355]" italics="true" pageId="1" pageNumber="283">Papaverellus aureocingulatus</emphasis>
Artigas &amp; Vieira
</taxonomicName>
<taxonomicNameLabel id="A21757CCFFAB5A1AFCBAE976FC7FFED6" box="[855,944,330,355]" pageId="1" pageNumber="283" rank="species">
<emphasis id="B924EAB7FFAB5A1AFCBAE976FC66FED7" bold="true" box="[855,937,330,355]" pageId="1" pageNumber="283">sp. nov</emphasis>
.
</taxonomicNameLabel>
(orig. des.).
</paragraph>
</subSubSection>
<subSubSection id="C34A652EFFAB5A1AFF2AE952FC32FE32" box="[199,1021,366,391]" pageId="1" pageNumber="283" type="etymology">
<paragraph id="8BEF36A5FFAB5A1AFF2AE952FC32FE32" blockId="1.[151,1436,330,1837]" box="[199,1021,366,391]" pageId="1" pageNumber="283">
<emphasis id="B924EAB7FFAB5A1AFF2AE952FE81FE33" bold="true" box="[199,334,366,391]" pageId="1" pageNumber="283">Etymology.</emphasis>
The name
<taxonomicName id="4C504D26FFAB5A1AFE26E952FD91FE32" box="[459,606,366,390]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B924EAB7FFAB5A1AFE26E952FD91FE32" box="[459,606,366,390]" italics="true" pageId="1" pageNumber="283">Papaverellus</emphasis>
</taxonomicName>
is dedicated to Dr. Nelson Papavero.
</paragraph>
</subSubSection>
<subSubSection id="C34A652EFFAB5A1AFF2AE9AEFD2EFE1F" box="[199,737,402,427]" pageId="1" pageNumber="283" type="distribution">
<paragraph id="8BEF36A5FFAB5A1AFF2AE9AEFD2EFE1F" blockId="1.[151,1436,330,1837]" box="[199,737,402,427]" pageId="1" pageNumber="283">
<emphasis id="B924EAB7FFAB5A1AFF2AE9AEFEAFFE1F" bold="true" box="[199,352,402,427]" pageId="1" pageNumber="283">Distribution.</emphasis>
<collectingCountry id="F3477635FFAB5A1AFE8AE9AEFE7FFE1F" box="[359,432,402,427]" name="Brazil" pageId="1" pageNumber="283">Brazil</collectingCountry>
: Pará State and Piauí State.
</paragraph>
</subSubSection>
<subSubSection id="C34A652EFFAB5A1AFF2AE98AFE3BFAEC" pageId="1" pageNumber="283" type="description">
<paragraph id="8BEF36A5FFAB5A1AFF2AE98AFBE4FD8F" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">
<emphasis id="B924EAB7FFAB5A1AFF2AE98AFE9EFE7B" bold="true" box="[199,337,438,463]" pageId="1" pageNumber="283">Description</emphasis>
. Head. Face (
<figureCitation id="136B2A20FFAB5A1AFE00E98AFDFEFE7B" box="[493,561,438,463]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 2</figureCitation>
) with only slightly exposed on lower ½ in lateral view, sparse black and short setae dorsally; remaining area and oral border with abundant long macrosetae directed anteriorly and ventrally (
<figureCitation id="136B2A20FFAB5A1AFF72E9C2FF2CFDA3" box="[159,227,510,535]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 3</figureCitation>
). Postpedicel of antenna as long as scape and pedicel combined (
<figureCitation id="136B2A20FFAB5A1AFC29E9C2FBC5FDA3" box="[964,1034,510,535]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 4</figureCitation>
); scape 1.5 times as long as pedicel (
<figureCitation id="136B2A20FFAB5A1AFF72EA1EFF2AFD8F" box="[159,229,546,571]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 4</figureCitation>
); stylus slightly longer than scape and pedicel; apex of stylus bare (
<figureCitation id="136B2A20FFAB5A1AFC34EA1EFBD1FD8F" box="[985,1054,546,571]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF2AEA7AFB41FD12" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">Thorax. Tumid, postsutural dorsocentral macrosetae, notopleural, supraalar and postalar macrosetae strong; anatergal setae absent; scutellum tumid as the thorax and coxae, without rim on posterior border; 2 scutellar macrosetae on posterior border, sparse, fine, short setae on disc and border; pleura asetose.</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF2AEA8EFD73FD46" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">
Wing (
<figureCitation id="136B2A20FFAB5A1AFEFEEA8EFE99FD7F" box="[275,342,690,715]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 5</figureCitation>
). Cell r1 closed before wing margin; stump vein supernumerary crossvein on R4 absent; R4 ending before the wing tip, and R5, after; m3 cell closed.
</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF2AEB3DFF29FC89" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">
Legs (
<figureCitation id="136B2A20FFAB5A1AFEE0EB3CFE80FCAC" box="[269,335,768,793]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 1</figureCitation>
). Coxae gray; femora shining black; tibiae yellow, except for brown stripe anteriorly; tarsus light brown.
</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF2AEB74FCABFCD4" blockId="1.[151,1436,330,1837]" box="[199,868,840,865]" pageId="1" pageNumber="283">Abdomen. Vestiture as in notum; sides of all tergites setose.</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF2AEB50FC18FB7C" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">
Male terminalia. Angled 45º with abdomen (
<figureCitation id="136B2A20FFAB5A1AFD26EB50FCC0FC31" box="[715,783,876,901]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[224,1332,195,1891]" captionTargetId="figure@3.[224,1359,195,1919]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 1, Habitus, lateral view; 2, Head, frontal view; 3, Head, lateral view; 4, Antennae, lateral view; 5, Wing; Male terminalia, lateral view. Abbreviations: pp: postpedicel; ped: pedicel; scp: scape; sty: stylus." httpUri="https://zenodo.org/record/229650/files/figure.png" pageId="1" pageNumber="283">Fig. 1</figureCitation>
); epandrium flat ventrally, looks like goat hoof (
<figureCitation id="136B2A20FFAB5A1AFAABEB50FA42FC31" box="[1350,1421,876,901]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">Fig. 7</figureCitation>
). Epandria broad and thick, subglobular on proximal 3/4, attenuated distally and meeting in an acute tip (
<figureCitation id="136B2A20FFAB5A1AFACFEBACFA42FC1C" box="[1314,1421,912,937]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">
Figs. 6
<emphasis id="B924EAB7FFAB5A1AFA82EBACFAB2FC1D" bold="true" box="[1391,1405,912,937]" pageId="1" pageNumber="283">–</emphasis>
8
</figureCitation>
). Gonocoxites globular on basal 3/4, distal 1/4 flattened, elongate, with 3
<emphasis id="B924EAB7FFAB5A1AFC0EEB88FC3EFC79" bold="true" box="[995,1009,948,973]" pageId="1" pageNumber="283">–</emphasis>
5 teeth or spines ventrally (
<figureCitation id="136B2A20FFAB5A1AFADBEB88FA41FC79" box="[1334,1422,948,973]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">Fig. 12</figureCitation>
). Median region of gonocoxite with round tumid protuberance with few short setae (
<figureCitation id="136B2A20FFAB5A1AFBDBEBE4FB03FC44" box="[1078,1228,984,1009]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">Figs. 7, 8, 12</figureCitation>
). Gonocoxite with 2 conspicuous patches of long setae, 1 of scattered long, fine, yellow setae directed posteriorly and other with patch of dense long, golden setae directed to opposite side and crossing over hypandrium tip as golden belt (
<figureCitation id="136B2A20FFAB5A1AFAC4EC1CFF7BFBE9" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">Figs. 7, 8, 11</figureCitation>
); gonostylus long, flattened towards apex, with abundant short, fine setae and few macrosetae (
<figureCitation id="136B2A20FFAB5A1AFAEFEC78FA41FBE9" box="[1282,1422,1092,1117]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">Figs. 11, 12</figureCitation>
); gonostylus hidden in closed genitalia; hypandrium subglobose and subtriangular, with 3 small tubercles on tip and patch of 12 long, fine setae on each side, directed posteriorly (
<figureCitation id="136B2A20FFAB5A1AFCB0ECB0FBDBFB11" box="[861,1044,1164,1189]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">Figs. 7, 8, 9, 11</figureCitation>
). Aedeagus long, ending in 2 long tubes, ejaculatory apodeme thin and long, not expanded apically (
<figureCitation id="136B2A20FFAB5A1AFC9BEC8CFC05FB7C" box="[886,970,1200,1225]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1883,1906]" captionTargetBox="[262,1322,259,1815]" captionTargetId="figure@4.[246,1335,193,1839]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Male. 6, Male terminalia, dorsal view; 7, Male terminalia, ventral view; 8, Male terminalia, ventral view; 9, Male terminalia, lateral view; 10, Inner margin of epandrium and subepandrial sclerite; 11, Hypandrium, gonocoxite and gonostylus, ventral view; 12, gonocoxite and gonostylus; 13, Aedeagus and ejaculatory apodeme, lateral view. Abbreviations: aed: aedeagus; cerc: cercus; ej apod: ejaculatory apodeme; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium." httpUri="https://zenodo.org/record/229651/files/figure.png" pageId="1" pageNumber="283">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF2AECE8FE3BFAEC" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">
Female terminalia. Ovipositor compressed, formed by segment VIII and the following (
<figureCitation id="136B2A20FFAB5A1AFB42ECE8FAF5FB59" box="[1199,1338,1236,1261]" captionStart="FIGURES 14 – 18" captionStartId="5.[151,264,1879,1902]" captionTargetBox="[221,1357,193,1812]" captionTargetId="figure@5.[215,1373,193,1840]" captionTargetPageId="5" captionText="FIGURES 14 – 18. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Female. 14, Habitus, lateral view; 15, Ovipositor, lateral view; 16, Ovipositor, dorsal view; 17, Ovipositor, ventral view; 18, Genital fork and spermathecae. Abbreviations: ar: arms of genital fork; cap duct: capsular ducts; com duct: common duct; exp duct: expulsory ducts; cerc: cercus; f apod: furcal apodeme; spmth: spermathecae; stVIII: sternite VIII; tVII: tergite VII; tVIII: tergite VIII; tIX + X: tergite IX + X." httpUri="https://zenodo.org/record/229652/files/figure.png" pageId="1" pageNumber="283">Figs. 14–17</figureCitation>
); genital fork with long, thin arms, about 2.5 times longer than base, base much wider than arms (
<figureCitation id="136B2A20FFAB5A1AFB79ECC4FB25FAA4" box="[1172,1258,1272,1297]" captionStart="FIGURES 14 – 18" captionStartId="5.[151,264,1879,1902]" captionTargetBox="[221,1357,193,1812]" captionTargetId="figure@5.[215,1373,193,1840]" captionTargetPageId="5" captionText="FIGURES 14 – 18. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Female. 14, Habitus, lateral view; 15, Ovipositor, lateral view; 16, Ovipositor, dorsal view; 17, Ovipositor, ventral view; 18, Genital fork and spermathecae. Abbreviations: ar: arms of genital fork; cap duct: capsular ducts; com duct: common duct; exp duct: expulsory ducts; cerc: cercus; f apod: furcal apodeme; spmth: spermathecae; stVIII: sternite VIII; tVII: tergite VII; tVIII: tergite VIII; tIX + X: tergite IX + X." httpUri="https://zenodo.org/record/229652/files/figure.png" pageId="1" pageNumber="283">Fig. 18</figureCitation>
); common duct long (
<figureCitation id="136B2A20FFAB5A1AFF31ED20FEFCFA81" box="[220,307,1308,1333]" captionStart="FIGURES 14 – 18" captionStartId="5.[151,264,1879,1902]" captionTargetBox="[221,1357,193,1812]" captionTargetId="figure@5.[215,1373,193,1840]" captionTargetPageId="5" captionText="FIGURES 14 – 18. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Female. 14, Habitus, lateral view; 15, Ovipositor, lateral view; 16, Ovipositor, dorsal view; 17, Ovipositor, ventral view; 18, Genital fork and spermathecae. Abbreviations: ar: arms of genital fork; cap duct: capsular ducts; com duct: common duct; exp duct: expulsory ducts; cerc: cercus; f apod: furcal apodeme; spmth: spermathecae; stVIII: sternite VIII; tVII: tergite VII; tVIII: tergite VIII; tIX + X: tergite IX + X." httpUri="https://zenodo.org/record/229652/files/figure.png" pageId="1" pageNumber="283">Fig. 18</figureCitation>
); 2 expulsory and capsular ducts ending in 2 spherical capsules, as wide as the width tip of the apodeme of the furca (
<figureCitation id="136B2A20FFAB5A1AFE7EED7CFE28FAEC" box="[403,487,1344,1369]" captionStart="FIGURES 14 – 18" captionStartId="5.[151,264,1879,1902]" captionTargetBox="[221,1357,193,1812]" captionTargetId="figure@5.[215,1373,193,1840]" captionTargetPageId="5" captionText="FIGURES 14 – 18. Papaverellus aureocingulatus Artigas &amp; Vieira gen. nov. et sp. nov. Female. 14, Habitus, lateral view; 15, Ovipositor, lateral view; 16, Ovipositor, dorsal view; 17, Ovipositor, ventral view; 18, Genital fork and spermathecae. Abbreviations: ar: arms of genital fork; cap duct: capsular ducts; com duct: common duct; exp duct: expulsory ducts; cerc: cercus; f apod: furcal apodeme; spmth: spermathecae; stVIII: sternite VIII; tVII: tergite VII; tVIII: tergite VIII; tIX + X: tergite IX + X." httpUri="https://zenodo.org/record/229652/files/figure.png" pageId="1" pageNumber="283">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A652EFFAB5A1AFF2AED58FEE8F899" pageId="1" pageNumber="283" type="discussion">
<paragraph id="8BEF36A5FFAB5A1AFF2AED58FF24F975" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">
<emphasis id="B924EAB7FFAB5A1AFF2AED58FE83FAC9" bold="true" box="[199,332,1380,1405]" pageId="1" pageNumber="283">Discussion.</emphasis>
In Papavero
<emphasis id="B924EAB7FFAB5A1AFE15ED59FDF7FAC8" box="[504,568,1380,1404]" italics="true" pageId="1" pageNumber="283">et al.</emphasis>
(2009),
<taxonomicName id="4C504D26FFAB5A1AFD4AED58FCF5FAC8" box="[679,826,1380,1404]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B924EAB7FFAB5A1AFD4AED58FCF5FAC8" box="[679,826,1380,1404]" italics="true" pageId="1" pageNumber="283">Papaverellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757CCFFAB5A1AFCA4ED58FC75FAC9" box="[841,954,1380,1405]" pageId="1" pageNumber="283" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFCA4ED58FC75FAC9" bold="true" box="[841,954,1380,1405]" pageId="1" pageNumber="283">gen. nov.</emphasis>
</taxonomicNameLabel>
, keys to
<taxonomicName id="4C504D26FFAB5A1AFBDAED58FA57FAC9" authority="Papavero, 2009" authorityName="Papavero" authorityYear="2009" box="[1079,1432,1380,1405]" class="Insecta" family="Asilidae" genus="Scarbroughia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFBDAED58FB1FFAC8" box="[1079,1232,1380,1404]" italics="true" pageId="1" pageNumber="283">Scarbroughia</emphasis>
Papavero, 2009
</taxonomicName>
. However,
<taxonomicName id="4C504D26FFAB5A1AFEFFEDB4FE6AFA14" box="[274,421,1416,1440]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B924EAB7FFAB5A1AFEFFEDB4FE6AFA14" box="[274,421,1416,1440]" italics="true" pageId="1" pageNumber="283">Papaverellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757CCFFAB5A1AFE5EEDB4FDECFA15" box="[435,547,1416,1441]" pageId="1" pageNumber="283" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFE5EEDB4FDECFA15" bold="true" box="[435,547,1416,1441]" pageId="1" pageNumber="283">gen. nov.</emphasis>
</taxonomicNameLabel>
differs from it by the following characters: stylus 1½ times longer than postpedicel; 2 apical scutellar macrosetae; setae present only in the fore and middle coxae, and sparse black macrosetae on the femora, tibiae and tarsi; basal 1/2 of epandrium globose; hypandrium with abundant macrosetae and setae laterally; aedeagus with 2 very long, pointed prongs. By contrast,
<taxonomicName id="4C504D26FFAB5A1AFC19EDC8FB42F9B8" box="[1012,1165,1524,1548]" class="Insecta" family="Asilidae" genus="Scarbroughia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFC19EDC8FB42F9B8" box="[1012,1165,1524,1548]" italics="true" pageId="1" pageNumber="283">Scarbroughia</emphasis>
</taxonomicName>
has postpedicel as long as the scape and pedicel combined and as long as the stylus; 4 or more apical scutellar macrosetae; apical 1/2 of the fore and middle femora medially with stout bristles, all tibiae and tarsi, and coxae (mainly fore and middle ones) have long white macrosetae, in addition to more or less long, white, dense, appressed short setae; epandrium narrow, elongate; hypandrium with patches of short, sparse macrosetae lateroventrally; aedeagus with 2 long prongs.
</paragraph>
<paragraph id="8BEF36A5FFAB5A1AFF2AEEF0FEE8F899" blockId="1.[151,1436,330,1837]" pageId="1" pageNumber="283">
The junior author included only
<taxonomicName id="4C504D26FFAB5A1AFDD2EEF0FD1DF950" box="[575,722,1740,1764]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B924EAB7FFAB5A1AFDD2EEF0FD1DF950" box="[575,722,1740,1764]" italics="true" pageId="1" pageNumber="283">Papaverellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757CCFFAB5A1AFD36EEF0FC89F951" box="[731,838,1740,1765]" pageId="1" pageNumber="283" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFD36EEF0FC89F951" bold="true" box="[731,838,1740,1765]" pageId="1" pageNumber="283">gen. nov.</emphasis>
</taxonomicNameLabel>
in the cladistic analysis of Asilinae (Vieira
<emphasis id="B924EAB7FFAB5A1AFAA9EEF0FAB2F950" box="[1348,1405,1740,1764]" italics="true" pageId="1" pageNumber="283">et.al.</emphasis>
in prep.), i.e.
<taxonomicName id="4C504D26FFAB5A1AFEF9EECCFE68F8BC" box="[276,423,1776,1800]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B924EAB7FFAB5A1AFEF9EECCFE68F8BC" box="[276,423,1776,1800]" italics="true" pageId="1" pageNumber="283">Papaverellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757CCFFAB5A1AFE42EECCFDD4F8BD" box="[431,539,1776,1801]" pageId="1" pageNumber="283" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFE42EECCFDD4F8BD" bold="true" box="[431,539,1776,1801]" pageId="1" pageNumber="283">gen. nov.</emphasis>
</taxonomicNameLabel>
with
<taxonomicName id="4C504D26FFAB5A1AFDB0EECCFCD4F8BD" authority="Bigot" authorityName="Bigot" box="[605,795,1776,1801]" class="Insecta" family="Asilidae" genus="Cerozodus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFDB0EECCFD1BF8BC" box="[605,724,1776,1800]" italics="true" pageId="1" pageNumber="283">Cerozodus</emphasis>
Bigot
</taxonomicName>
and
<taxonomicName id="4C504D26FFAB5A1AFCB8EECDFC3AF8BD" authority="Hull" authorityName="Hull" box="[853,1013,1776,1801]" class="Insecta" family="Asilidae" genus="Myaptex" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFCB8EECDFC7AF8BC" box="[853,949,1777,1800]" italics="true" pageId="1" pageNumber="283">Myaptex</emphasis>
Hull
</taxonomicName>
: (
<taxonomicName id="4C504D26FFAB5A1AFBE7EECDFBA4F8BC" box="[1034,1131,1777,1800]" class="Insecta" family="Asilidae" genus="Myaptex" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFBE7EECDFBA4F8BC" box="[1034,1131,1777,1800]" italics="true" pageId="1" pageNumber="283">Myaptex</emphasis>
</taxonomicName>
(
<taxonomicName id="4C504D26FFAB5A1AFB90EECCFADFF8BC" box="[1149,1296,1776,1800]" class="Insecta" family="Asilidae" genus="Papaverellus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B924EAB7FFAB5A1AFB90EECCFADFF8BC" box="[1149,1296,1776,1800]" italics="true" pageId="1" pageNumber="283">Papaverellus</emphasis>
</taxonomicName>
<emphasis id="B924EAB7FFAB5A1AFAF5EECCFA4BF8BD" bold="true" box="[1304,1412,1776,1801]" pageId="1" pageNumber="283">
<taxonomicNameLabel id="A21757CCFFAB5A1AFAF5EECCFA4BF8BD" box="[1304,1412,1776,1801]" pageId="1" pageNumber="283" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
+
<taxonomicName id="4C504D26FFAB5A1AFF7AEF28FEC1F898" box="[151,270,1812,1836]" class="Insecta" family="Asilidae" genus="Cerozodus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAB7FFAB5A1AFF7AEF28FEC1F898" box="[151,270,1812,1836]" italics="true" pageId="1" pageNumber="283">Cerozodus</emphasis>
</taxonomicName>
)).
</paragraph>
</subSubSection>
</treatment>
</document>
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