281 lines
40 KiB
XML
281 lines
40 KiB
XML
<document id="07DC625E4CF6E44953EECA8B29634FC5" ID="10.11646/zootaxa.4150.3.3" ID-DOI="10.11646/zootaxa.4150.3.3" ID-GBIF-Dataset="e98edafd-7489-488d-ab8a-276a4a4926a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="265618" ID-ZooBank="79657D7A-324F-4539-A80F-E11BB4189753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470984303137" checkinUser="plazi" docAuthor="Patitucci, Luciano Damián, Mulieri, Pablo Ricardo & Mariluis, Juan Carlos" docDate="2016" docId="03F65F6EF047FFD9FF6CFAFAFB762CE8" docLanguage="en" docName="zootaxa.4150.3.3.pdf" docOrigin="Zootaxa 4150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina xena Malloch 1934" docType="treatment" docVersion="7" lastPageNumber="310" masterDocId="FFCF2716F05CFFC4FFFBFF88FFAE2F04" masterDocTitle="Taxonomic review of the species of Helina R. - D. (Diptera: Muscidae) from Andean-Patagonian forests" masterLastPageNumber="313" masterPageNumber="281" pageNumber="308" updateTime="1698381830647" updateUser="plazi">
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<mods:titleInfo id="768BA808132CDDC8E725F9E971E26440">
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<mods:title id="4A654FC346F5C2D390A28727608A9A32">Taxonomic review of the species of Helina R. - D. (Diptera: Muscidae) from Andean-Patagonian forests</mods:title>
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<mods:name id="7DF011B949A4BA9DBEFB8B216F61AAF7" type="personal">
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<mods:namePart id="4FE0AEF26664A52C9F8C5DA59AD2E345">Patitucci, Luciano Damián</mods:namePart>
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<mods:namePart id="C4BB94A39706078BC38FD54948D5A17A">Mulieri, Pablo Ricardo</mods:namePart>
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<mods:roleTerm id="A1633A2152C55BC423EF00ED7F4122EF">Author</mods:roleTerm>
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<mods:namePart id="27B92A1634890FE0FBCE8CD5970EF38A">Mariluis, Juan Carlos</mods:namePart>
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<mods:typeOfResource id="74E3DEEF23BB0A2012481677F585B0B9">text</mods:typeOfResource>
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<mods:title id="5302E29596DB23C4C2625D6D4D4E3E87">Zootaxa</mods:title>
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<mods:date id="DAE5E299ED9FBD1871EB992E7C8D04E5">2016</mods:date>
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<mods:number id="F1E0B7A59F758CFBE3B4E7EAA6A35883">4150</mods:number>
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<mods:identifier id="998E3B6D4B87A44F5D20B002210DBA9A" type="DOI">10.11646/zootaxa.4150.3.3</mods:identifier>
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<treatment id="03F65F6EF047FFD9FF6CFAFAFB762CE8" ID-DOI="http://doi.org/10.5281/zenodo.6088881" ID-GBIF-Taxon="124329292" ID-Zenodo-Dep="6088881" LSID="urn:lsid:plazi:treatment:03F65F6EF047FFD9FF6CFAFAFB762CE8" httpUri="http://treatment.plazi.org/id/03F65F6EF047FFD9FF6CFAFAFB762CE8" lastPageId="29" lastPageNumber="310" pageId="27" pageNumber="308">
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<subSubSection id="C345BDF3F047FFDFFF6CFAFAFE932AAA" pageId="27" pageNumber="308" type="nomenclature">
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<paragraph id="8BE0EE78F047FFDFFF6CFAFAFE4D2A89" blockId="27.[151,483,1394,1454]" box="[151,483,1394,1421]" pageId="27" pageNumber="308">
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<heading id="D0A85914F047FFDFFF6CFAFAFE4D2A89" bold="true" box="[151,483,1394,1421]" fontSize="11" level="1" pageId="27" pageNumber="308" reason="1">
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<taxonomicName id="4C5F95FBF047FFDFFF6CFAFAFE4D2A89" authority="Malloch, 1934" authorityName="Malloch" authorityYear="1934" box="[151,483,1394,1421]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="308" phylum="Arthropoda" rank="species" species="xena">
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||
<emphasis id="B92B326AF047FFDFFF6CFAFAFE4D2A89" bold="true" box="[151,483,1394,1421]" pageId="27" pageNumber="308">
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<emphasis id="B92B326AF047FFDFFF6CFAFAFE822A88" bold="true" box="[151,300,1394,1420]" italics="true" pageId="27" pageNumber="308">Helina xena</emphasis>
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<bibRefCitation id="EFCE9389F047FFDFFEC8FAFBFE4D2A89" author="Malloch" box="[307,483,1395,1421]" pageId="27" pageNumber="308" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch, 1934</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE0EE78F047FFDFFF6CFA1DFE932AAA" blockId="27.[151,483,1394,1454]" box="[151,317,1429,1454]" pageId="27" pageNumber="308">
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||
(
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<figureCitation id="1364F2FDF047FFDFFF64FA1DFF522AAA" box="[159,252,1429,1454]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="27" pageNumber="308">Figs. 14</figureCitation>
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,
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<figureCitation id="1364F2FDF047FFDFFEF3FA1DFE8D2AAA" box="[264,291,1429,1454]" captionStart="FIGURE 15" captionStartId="30.[151,250,1475,1497]" captionTargetBox="[152,1436,267,1447]" captionTargetId="figure@30.[151,1436,267,1453]" captionTargetPageId="30" captionText="FIGURE 15. Geographical distribution. A. Helina neosimplex Snyder. B. Helina nigrimana (Macquart). C. Helina ouina sp. nov. D. Helina rufoapicata Malloch. E. Helina viola Malloch. G. Helina xena Malloch. (Green area: Andean-Patagonian forests; orange circle: new records; blue triangle: previous distribution.)" httpUri="https://zenodo.org/record/265633/files/figure.png" pageId="27" pageNumber="308">15</figureCitation>
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F)
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</paragraph>
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</subSubSection>
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<subSubSection id="C345BDF3F047FFD9FF6CFA56FD222DAC" lastPageId="29" lastPageNumber="310" pageId="27" pageNumber="308" type="description">
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<paragraph id="8BE0EE78F047FFDFFF6CFA56FC222AF7" blockId="27.[151,1078,1501,1584]" box="[151,908,1501,1523]" pageId="27" pageNumber="308">
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For a complete list of references, see the catalog by
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<bibRefCitation id="EFCE9389F047FFDFFD5EFA55FC292AF7" box="[677,903,1501,1523]" pageId="27" pageNumber="308" refString="Carvalho, C. J. B. de, Couri, M. S., Pont, A. C., Pamplona, D. & Lopes, S. M. (2005) A Catalogue of the Muscidae (Diptera) of the Neotropical Region. Zootaxa, 860, 1 - 282. http: // dx. doi. org / 10.11646 / zootaxa. 860.1.1" type="journal article">
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Carvalho
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<emphasis id="B92B326AF047FFDFFCF3FA56FC922AF7" box="[776,828,1501,1523]" italics="true" pageId="27" pageNumber="308">et al.</emphasis>
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(2005)
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</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BE0EE78F047FFDFFF6CFA74FB982934" blockId="27.[151,1078,1501,1584]" pageId="27" pageNumber="308">
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Information subsequent to this catalog:
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<bibRefCitation id="EFCE9389F047FFDFFDD3FA74FC292916" author="Lowerberg-Neto" box="[552,903,1532,1554]" pageId="27" pageNumber="308" refString="Lowerberg-Neto, P. & Carvalho, C. J. B. de (2009). Areas of endemism and spatial diversification of the Muscidae (Insecta: Diptera) in the Andean and Neotropical regions. Journal of Biogeography, 36, 1750 - 1759. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2009.02112. x" type="journal article" year="2009">Löwerberg-Neto & Carvalho 2009</bibRefCitation>
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(endemism).
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<bibRefCitation id="EFCE9389F047FFDFFF6CF992FE1E2934" author="Lowerberg-Neto" box="[151,432,1562,1584]" pageId="27" pageNumber="308" refString="Lowerberg-Neto, P., Carvalho, C. J. B. de & Hawkins, B. A. (2011) Tropical niche conservatism as a historical narrative hypothesis for the Neotropics: a case study using the fly family Muscidae. Journal of Biogeography, 38, 1936 - 1947. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2011.02540. x" type="journal article" year="2011">
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Löwerberg-Neto
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<emphasis id="B92B326AF047FFDFFEBEF993FEDC2934" box="[325,370,1562,1584]" italics="true" pageId="27" pageNumber="308">et al</emphasis>
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. 2011
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</bibRefCitation>
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(biogeography);
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<bibRefCitation id="EFCE9389F047FFDFFD9BF992FC102934" author="Lowerberg-Neto" box="[608,958,1562,1584]" pageId="27" pageNumber="308" refString="Lowerberg-Neto, P. & Carvalho, C. J. B. de (2013) Muscidae (Insecta: Diptera) of Latin America and the Caribbean: geographic distribution and check-list by country. Zootaxa, 3650 (1), 01 - 147. http: // dx. doi. org / 10.11646 / zootaxa. 3650.1.1" type="journal article" year="2013">Löwerberg-Neto & Carvalho 2013</bibRefCitation>
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(checklist).
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</paragraph>
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||
<paragraph id="8BE0EE78F047FFDFFF6CF9D5FC702971" blockId="27.[151,1436,1628,2014]" box="[151,990,1628,1654]" pageId="27" pageNumber="308">
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<emphasis id="B92B326AF047FFDFFF6CF9D5FE932972" bold="true" box="[151,317,1629,1654]" pageId="27" pageNumber="308">Redescription</emphasis>
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.
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<emphasis id="B92B326AF047FFDFFEB0F9D4FE292971" bold="true" box="[331,391,1628,1653]" italics="true" pageId="27" pageNumber="308">Male</emphasis>
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(
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<figureCitation id="1364F2FDF047FFDFFE6EF9D5FE492972" box="[405,487,1629,1654]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="27" pageNumber="308">Fig. 14</figureCitation>
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A). Length. Body:
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<quantity id="4CA7439DF047FFDFFD41F9D5FC8E2971" box="[698,800,1629,1654]" metricMagnitude="-3" metricUnit="m" metricValue="6.31" pageId="27" pageNumber="308" unit="mm" value="6.31">6.31 mm</quantity>
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, wing:
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<quantity id="4CA7439DF047FFDFFC89F9D5FC772971" box="[882,985,1629,1654]" metricMagnitude="-3" metricUnit="m" metricValue="5.94" pageId="27" pageNumber="308" unit="mm" value="5.94">5.94 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BE0EE78F047FFDFFF3CF909FB5829E6" blockId="27.[151,1436,1628,2014]" pageId="27" pageNumber="308">
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<emphasis id="B92B326AF047FFDFFF3CF909FEAA299E" box="[199,260,1665,1690]" italics="true" pageId="27" pageNumber="308">Head</emphasis>
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(
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<figureCitation id="1364F2FDF047FFDFFEEFF909FEC6299E" box="[276,360,1665,1690]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="27" pageNumber="308">Fig. 14</figureCitation>
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B). Black with silver pollinosity. Holoptic, the shortest distance between eyes is
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<quantity id="4CA7439DF047FFDFFB05F909FAC8299D" box="[1278,1382,1665,1690]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="308" unit="mm" value="0.16">0.16 mm</quantity>
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. 4–5 pairs of frontal setae. Eye bare. Lunule brown; fronto-orbital plate, parafacial, gena, postgena, and occiput black with silver pollinosity. Antenna black; arista with hairs shorter than its basal diameter. Palpus black.
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</paragraph>
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<paragraph id="8BE0EE78F047FFDFFF3CF965FE8D28D9" blockId="27.[151,1436,1628,2014]" pageId="27" pageNumber="308">
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<emphasis id="B92B326AF047FFDFFF3CF965FEB62802" box="[199,280,1773,1798]" italics="true" pageId="27" pageNumber="308">Thorax</emphasis>
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||
(
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<figureCitation id="1364F2FDF047FFDFFED1F965FED12802" box="[298,383,1773,1798]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="27" pageNumber="308">Fig. 14</figureCitation>
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C). Scutum black with four brownish-grey pollinose vittae; scutellum black with brownishgrey pollinosity; anepisternum, anepimeron, katepimeron, katepisternum, proepisternum, proepimeron and meron black with grey pollinosity; anterior spiracle black and posterior spiracle brown. Chaetotaxy: acrostichal setae short, the seta before the scutellum longer than previous seta; dorsocentral setae 2+3; humeral setae 2; notopleural setae 2, of similar size. Prealar seta strong, shorter than one fourth of the length of the anterior supra-alar postsutural seta. Scutellum with strong and short basal and subapical setae. Anepisternum with a series of 6–8 strong setae.
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</paragraph>
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<caption id="DF20BEF0F040FFD8FF6CFA3FFA862963" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="28" pageNumber="309" targetBox="[173,1413,183,1428]" targetPageId="28">
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||
<paragraph id="8BE0EE78F040FFD8FF6CFA3FFA862963" blockId="28.[151,1436,1463,1639]" pageId="28" pageNumber="309">
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||
<emphasis id="B92B326AF040FFD8FF6CFA3FFE8E2AC9" bold="true" box="[151,288,1463,1485]" pageId="28" pageNumber="309">FIGURE 14.</emphasis>
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<taxonomicName id="4C5F95FBF040FFD8FED3FA3FFD8F2AC9" authority="Malloch. A" authorityName="Malloch. A" box="[296,545,1463,1486]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="309" phylum="Arthropoda" rank="species" species="xena">
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<emphasis id="B92B326AF040FFD8FED3FA3FFE0A2ACA" box="[296,420,1463,1486]" italics="true" pageId="28" pageNumber="309">Helina xena</emphasis>
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Malloch.
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<emphasis id="B92B326AF040FFD8FDF5FA3FFD8F2AC9" bold="true" box="[526,545,1463,1485]" pageId="28" pageNumber="309">A</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="B92B326AF040FFD8FDDBFA3FFDE92AC9" bold="true" box="[544,583,1463,1485]" pageId="28" pageNumber="309">–H.</emphasis>
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Male:
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<emphasis id="B92B326AF040FFD8FD6AFA3FFD042AC9" bold="true" box="[657,682,1463,1485]" pageId="28" pageNumber="309">A.</emphasis>
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Lateral view (scale bar: 1 mm).
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<emphasis id="B92B326AF040FFD8FC07FA3FFBBA2AC9" bold="true" box="[1020,1044,1463,1485]" pageId="28" pageNumber="309">B.</emphasis>
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Head, frontal view.
|
||
<emphasis id="B92B326AF040FFD8FB1CFA3FFB512AC9" bold="true" box="[1255,1279,1463,1485]" pageId="28" pageNumber="309">C.</emphasis>
|
||
Thorax, dorsal view.
|
||
<emphasis id="B92B326AF040FFD8FF2DFA5EFF402AE8" bold="true" box="[214,238,1494,1516]" pageId="28" pageNumber="309">D.</emphasis>
|
||
Abdomen, dorsal view (scale bars: 0.5 mm).
|
||
<emphasis id="B92B326AF040FFD8FD29FA5EFD472AE8" bold="true" box="[722,745,1494,1516]" pageId="28" pageNumber="309">E.</emphasis>
|
||
Sternite 5 (scale bar: 0.2 mm).
|
||
<emphasis id="B92B326AF040FFD8FBC0FA5EFBE22AE8" bold="true" box="[1083,1100,1494,1516]" pageId="28" pageNumber="309">F.</emphasis>
|
||
Cercal plate, posterior view.
|
||
<emphasis id="B92B326AF040FFD8FA7EFA5EFA372AE8" bold="true" box="[1413,1433,1494,1516]" pageId="28" pageNumber="309">G.</emphasis>
|
||
Cercal plate and surstylus, lateral view (scale bar: 0.1 mm).
|
||
<emphasis id="B92B326AF040FFD8FD02FA7DFCBA290F" bold="true" box="[761,788,1525,1547]" pageId="28" pageNumber="309">H.</emphasis>
|
||
Phallic complex, lateral view (scale bar: 0.1 mm).
|
||
<emphasis id="B92B326AF040FFD8FAE0FA7DFAEB290F" bold="true" box="[1307,1349,1525,1547]" pageId="28" pageNumber="309">I–J.</emphasis>
|
||
Female, apical portion of ovipositor:
|
||
<emphasis id="B92B326AF040FFD8FE47F99BFE62292D" bold="true" box="[444,460,1555,1577]" pageId="28" pageNumber="309">I.</emphasis>
|
||
Dorsal view.
|
||
<emphasis id="B92B326AF040FFD8FDA0F99BFDC0292D" bold="true" box="[603,622,1555,1577]" pageId="28" pageNumber="309">J.</emphasis>
|
||
Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE0EE78F040FFD8FF3CF91DFC3A29F0" blockId="28.[151,1436,1683,2035]" pageId="28" pageNumber="309">
|
||
<emphasis id="B92B326AF040FFD8FF3CF91DFF5129A8" box="[199,255,1685,1708]" italics="true" pageId="28" pageNumber="309">Wing</emphasis>
|
||
. Yellowish-brown; costal spine longer than the costal bristles; the cross-veins with cloud; the cross-vein dm-cu straight, an extra little cross-vein close to cross-vein r-m; vein R 4+5 and vein M straight; cross vein dm-cu in basal half of cell r4+5. Both calypters hyaline with yellow margins.
|
||
</paragraph>
|
||
<paragraph id="8BE0EE78F040FFD8FF3CF88BFA9228F7" blockId="28.[151,1436,1683,2035]" pageId="28" pageNumber="309">
|
||
<emphasis id="B92B326AF040FFD8FF3CF88BFF52281E" box="[199,252,1795,1818]" italics="true" pageId="28" pageNumber="309">Legs</emphasis>
|
||
. Coxa and trochanter black; fore femur black, yellow at base; mid and hind femora yellow, with black on apical third; tibiae yellow, tarsi black. Fore femur with rows of dorsal, posterodorsal, and posteroventral setae; fore tibia with one posterior seta, 4 preapical setae. Mid femur with 6–7 setae in the basal middle on ventral surface, a row of anterodorsal seta, and 3–4 preapical setae on dorsal to posterior surface; mid tibia with 2 posterior setae and 4 apical setae (anterodorsal, posteroventral, ventral, and anteroventral). Hind femur with anterodorsal row, anteroventral row of setae and 3–4 setae in the apical middle of posteroventral surface; hind tibia with 2–3 anteroventral setae, and one anterodorsal, calcar absent. Claws and pulvilli of similar size in all three legs.
|
||
</paragraph>
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFF1FFCE02FD0" blockId="29.[151,1436,151,1005]" pageId="29" pageNumber="310">
|
||
<emphasis id="B92B326AF041FFD9FF3CFF1FFE9F2FB4" box="[199,305,151,176]" italics="true" pageId="29" pageNumber="310">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="1364F2FDF041FFD9FEBBFF1FFE3C2FB4" box="[320,402,151,176]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="29" pageNumber="310">Fig. 14</figureCitation>
|
||
D). Black with brown pollinosity, two dark brown spots on tergite 3 and 4. Sternite 5 longer than wide, setulose; posterior margin membranous (
|
||
<figureCitation id="1364F2FDF041FFD9FD27FF34FC832FD0" box="[732,813,188,213]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="29" pageNumber="310">Fig. 14</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFF57FCB52EA8" blockId="29.[151,1436,151,1005]" pageId="29" pageNumber="310">
|
||
<taxonomicName id="4C5F95FBF041FFD9FF3CFF57FEEF2FFC" box="[199,321,223,248]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="310" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B92B326AF041FFD9FF3CFF57FEEF2FFC" box="[199,321,223,248]" italics="true" pageId="29" pageNumber="310">Terminalia</emphasis>
|
||
</taxonomicName>
|
||
. Cercal plate with a similar length and width, with only a distal incision; and with four protuberances at apex: inner protuberance short and rounded, outer protuberance lobed shape, and straight in lateral view; epandrium with lateral projection (
|
||
<figureCitation id="1364F2FDF041FFD9FD9CFEAFFD6D2E44" box="[615,707,295,320]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="29" pageNumber="310">Figs. 14</figureCitation>
|
||
F–G). Surstylus broad and curving forwards distally, with some setae on inner surface (
|
||
<figureCitation id="1364F2FDF041FFD9FE4DFEC4FDB72E60" box="[438,537,332,357]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="29" pageNumber="310">Figs. 14</figureCitation>
|
||
F–G). Aedeagus with aedeagal apodeme straight and strongly sclerotized; epiphalus slightly sclerotized, paramere slightly bent downward with setula; gonopod with short setulae; and distiphallus campanullate, slightly sclerotized (
|
||
<figureCitation id="1364F2FDF041FFD9FD5DFE1CFD592EA8" box="[678,759,404,429]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="29" pageNumber="310">Fig. 14</figureCitation>
|
||
H).
|
||
</paragraph>
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFE3FFD562ECB" blockId="29.[151,1436,151,1005]" box="[199,760,439,464]" pageId="29" pageNumber="310">
|
||
<emphasis id="B92B326AF041FFD9FF3CFE3FFEB02ED4" bold="true" box="[199,286,439,464]" italics="true" pageId="29" pageNumber="310">Female</emphasis>
|
||
. Length. Body:
|
||
<quantity id="4CA7439DF041FFD9FE2EFE3FFD952ECB" box="[469,571,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" pageId="29" pageNumber="310" unit="mm" value="4.95">4.95 mm</quantity>
|
||
, wing:
|
||
<quantity id="4CA7439DF041FFD9FD76FE3FFD5A2ECB" box="[653,756,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" pageId="29" pageNumber="310" unit="mm" value="5.15">5.15 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFE54FDA32D39" blockId="29.[151,1436,151,1005]" pageId="29" pageNumber="310">
|
||
Differs from male as follows:
|
||
<emphasis id="B92B326AF041FFD9FDDBFE54FDF32EF1" box="[544,605,476,501]" italics="true" pageId="29" pageNumber="310">Head</emphasis>
|
||
: dichoptic, the shortest distance between eyes is
|
||
<quantity id="4CA7439DF041FFD9FB6BFE54FB562EF0" box="[1168,1272,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="29" pageNumber="310" unit="mm" value="0.69">0.69 mm</quantity>
|
||
; fronto-orbital plate setulose with 2 reclinate orbital setae. Parafrontal plate setulose.
|
||
<emphasis id="B92B326AF041FFD9FC56FD89FC4C2D1C" box="[941,994,513,536]" italics="true" pageId="29" pageNumber="310">Legs</emphasis>
|
||
. Mid femur with 3–4 strong of setae in the basal third on ventral surface.
|
||
</paragraph>
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFDCFFD222DAC" blockId="29.[151,1436,151,1005]" pageId="29" pageNumber="310">
|
||
<taxonomicName id="4C5F95FBF041FFD9FF3CFDCFFEEF2D64" box="[199,321,583,608]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="310" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B92B326AF041FFD9FF3CFDCFFEEF2D64" box="[199,321,583,608]" italics="true" pageId="29" pageNumber="310">Terminalia</emphasis>
|
||
</taxonomicName>
|
||
. Tergite 8 with 2 parallel sclerotized plates with setulae on distal margin; epiproct with proximal margin straight, and a row of setula on distal margin, cercus digitiform with long setulae (
|
||
<figureCitation id="1364F2FDF041FFD9FB55FDE4FAAD2D80" box="[1198,1283,620,645]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="29" pageNumber="310">
|
||
Fig.
|
||
<date id="FFE1C8B8F041FFD9FB1EFDE4FAAD2D80" box="[1253,1283,620,644]" pageId="29" pageNumber="310" value="2014-01">14</date>
|
||
</figureCitation>
|
||
I). Hypoproct rounded, sclerotized, and setulose (
|
||
<figureCitation id="1364F2FDF041FFD9FDDAFD07FDDD2DAC" box="[545,627,655,680]" captionStart="FIGURE 14" captionStartId="28.[151,250,1463,1485]" captionTargetBox="[173,1413,183,1428]" captionTargetId="figure@28.[173,1413,183,1454]" captionTargetPageId="28" captionText="FIGURE 14. Helina xena Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; e lat prot, epandrium lateral projection; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; i protub, inner protuberance; o protub, outer protuberance; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265632/files/figure.png" pageId="29" pageNumber="310">Fig. 14</figureCitation>
|
||
J).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345BDF3F041FFD9FF3CFD3CFD502C10" pageId="29" pageNumber="310" type="materials_examined">
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFD3CFD502C10" blockId="29.[151,1436,151,1005]" pageId="29" pageNumber="310">
|
||
<materialsCitation id="3B37E425F041FFD9FF3CFD3CFD382DF4" ID-GBIF-Occurrence="1291551015" collectingDate="2013-02" collectionCode="MACN" collectorName="PN Los Alerces & Cabana La Cascada & Patitucci" country="Argentina" location="Material" pageId="29" pageNumber="310" specimenCount="1" specimenCount-female="1" stateProvince="Chubut">
|
||
<emphasis id="B92B326AF041FFD9FF3CFD3CFD7F2DC9" bold="true" box="[199,721,692,717]" pageId="29" pageNumber="310">
|
||
<location id="8E80B8A3F041FFD9FF3CFD3CFE812DC9" LSID="urn:lsid:plazi:treatment:03F65F6EF047FFD9FF6CFAFAFB762CE8:8E80B8A3F041FFD9FF3CFD3CFE812DC9" box="[199,303,692,717]" country="Argentina" name="Material" pageId="29" pageNumber="310" stateProvince="Chubut">Material</location>
|
||
examined.
|
||
<collectingCountry id="F348AEE8F041FFD9FE42FD3CFDC72DC9" box="[441,617,692,717]" name="Argentina" pageId="29" pageNumber="310">ARGENTINA</collectingCountry>
|
||
:
|
||
<collectingRegion id="499B209AF041FFD9FD8DFD3CFD7F2DC9" box="[630,721,692,717]" country="Argentina" name="Chubut" pageId="29" pageNumber="310">Chubut</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D5925F1F041FFD9FD1AFD3CFCE82DC9" box="[737,838,692,717]" pageId="29" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<collectorName id="26AA8BAEF041FFD9FCA9FD3DFBA62DC9" box="[850,1032,692,717]" pageId="29" pageNumber="310">PN Los Alerces</collectorName>
|
||
,
|
||
<collectorName id="26AA8BAEF041FFD9FBEEFD3CFB592DC9" box="[1045,1271,692,717]" pageId="29" pageNumber="310">Cabaña La Cascada</collectorName>
|
||
, -42.888499, - 71.592376,
|
||
<date id="FFE1C8B8F041FFD9FEDAFD50FED92DF4" box="[289,375,728,752]" pageId="29" pageNumber="310" value="2013-02">
|
||
<collectingDate id="EFA53150F041FFD9FEDAFD50FED92DF4" box="[289,375,728,752]" pageId="29" pageNumber="310" value="2013-02">II-2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26AA8BAEF041FFD9FE73FD5FFE442DF4" box="[392,490,727,752]" pageId="29" pageNumber="310">Patitucci</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED4E76BDF041FFD9FDC8FD50FD202DF4" box="[563,654,728,752]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="29" pageNumber="310">MACN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
; 1 male, PN Los Alerces, Puerto Limonao, Lago Futalaufquen, - 42.862656, -71.624142,
|
||
<date id="FFE1C8B8F041FFD9FE51FD74FDAF2C11" box="[426,513,764,789]" pageId="29" pageNumber="310" value="1959-02">II-1959</date>
|
||
, Trelles leg. (MACN).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345BDF3F041FFD9FF3CFC97FE2A2C59" pageId="29" pageNumber="310" type="distribution">
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFC97FE2A2C59" blockId="29.[151,1436,151,1005]" pageId="29" pageNumber="310">
|
||
<emphasis id="B92B326AF041FFD9FF3CFC97FEF72C3C" bold="true" box="[199,345,799,824]" pageId="29" pageNumber="310">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1364F2FDF041FFD9FE90FC97FE112C3C" box="[363,447,799,824]" captionStart="FIGURE 15" captionStartId="30.[151,250,1475,1497]" captionTargetBox="[152,1436,267,1447]" captionTargetId="figure@30.[151,1436,267,1453]" captionTargetPageId="30" captionText="FIGURE 15. Geographical distribution. A. Helina neosimplex Snyder. B. Helina nigrimana (Macquart). C. Helina ouina sp. nov. D. Helina rufoapicata Malloch. E. Helina viola Malloch. G. Helina xena Malloch. (Green area: Andean-Patagonian forests; orange circle: new records; blue triangle: previous distribution.)" httpUri="https://zenodo.org/record/265633/files/figure.png" pageId="29" pageNumber="310">Fig. 15</figureCitation>
|
||
F).
|
||
<collectingCountry id="F348AEE8F041FFD9FE1CFCA8FD212C3C" box="[487,655,800,824]" name="Argentina" pageId="29" pageNumber="310">ARGENTINA</collectingCountry>
|
||
:
|
||
<collectingRegion id="499B209AF041FFD9FD67FC97FD402C3C" box="[668,750,799,824]" country="Argentina" name="Chubut" pageId="29" pageNumber="310">Chubut</collectingRegion>
|
||
(new record),
|
||
<collectingRegion id="499B209AF041FFD9FC61FCA8FBB82C3C" box="[922,1046,800,824]" country="Argentina" name="Rio Negro" pageId="29" pageNumber="310">Río Negro</collectingRegion>
|
||
.
|
||
<collectingCountry id="F348AEE8F041FFD9FBD9FCA8FBD72C3C" box="[1058,1145,800,824]" name="Chile" pageId="29" pageNumber="310">CHILE</collectingCountry>
|
||
:
|
||
<collectingRegion id="499B209AF041FFD9FB7DFC97FA362C3C" box="[1158,1432,799,824]" country="Chile" name="Araucania" pageId="29" pageNumber="310">Región de la Araucanía</collectingRegion>
|
||
,
|
||
<collectingRegion id="499B209AF041FFD9FF6CFCCCFE2E2C59" box="[151,384,836,861]" country="Chile" name="Los Lagos" pageId="29" pageNumber="310">Región de los Lagos</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345BDF3F041FFD9FF3CFCEFFC1F2CCC" pageId="29" pageNumber="310" type="discussion">
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFCEFFC1F2CCC" blockId="29.[151,1436,151,1005]" pageId="29" pageNumber="310">
|
||
<emphasis id="B92B326AF041FFD9FF3CFCEFFE952C84" bold="true" box="[199,315,871,896]" pageId="29" pageNumber="310">Remarks.</emphasis>
|
||
<bibRefCitation id="EFCE9389F041FFD9FEB9FCEFFE5F2C84" author="Malloch" box="[322,497,871,896]" pageId="29" pageNumber="310" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch (1934)</bibRefCitation>
|
||
described the species from male and female specimens from several locations from
|
||
<collectingCountry id="F348AEE8F041FFD9FF6CFC04FEA82CA1" box="[151,262,908,933]" name="Argentina" pageId="29" pageNumber="310">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="F348AEE8F041FFD9FEB3FC04FE262CA1" box="[328,392,908,933]" name="Chile" pageId="29" pageNumber="310">Chile</collectingCountry>
|
||
. The male specimen studied here has an extra little cross-vein close to cross-vein r-m; postocular setae, inner vertical setae, and outer vertical setae damaged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345BDF3F041FFD9FF3CFC5CFB762CE8" box="[199,1240,980,1005]" pageId="29" pageNumber="310" type="biology_ecology">
|
||
<paragraph id="8BE0EE78F041FFD9FF3CFC5CFB762CE8" blockId="29.[151,1436,151,1005]" box="[199,1240,980,1005]" pageId="29" pageNumber="310">
|
||
<emphasis id="B92B326AF041FFD9FF3CFC5CFEB12CE9" bold="true" box="[199,287,980,1005]" pageId="29" pageNumber="310">Biology</emphasis>
|
||
. Unknown. The specimen collected by the authors was captured with a Malaise trap.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |