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<document id="3438405B383DF4FCC6B43038EB5BAD14" ID="10.11646/zootaxa.4125.1.1" ID-CLB-Dataset="38807" ID-DOI="10.11646/zootaxa.4125.1.1" ID-GBIF-Dataset="1af13aec-3c54-4a92-abdf-e4e29b97e82f" ID-ISSN="1175-5326" ID-Zenodo-Dep="271704" ID-ZooBank="118F4865-D89E-45EA-A210-8D61946CC37F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465971881907" checkinUser="plazi" docAuthor="Matz, Jess &amp; Brower, Andrew V. Z." docDate="2016" docId="03F187D7FFCF8439FF11FD84FD84BDCF" docLanguage="en" docName="zootaxa.4125.1.1.pdf" docOrigin="Zootaxa 4125 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tetraphlebia C. Felder &amp; R. Felder 1867" docType="treatment" docVersion="9" lastPageNumber="69" masterDocId="FFC8FFAFFF8B847DFF86FFA0FFD9B918" masterDocTitle="The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes" masterLastPageNumber="108" masterPageNumber="1" pageNumber="69" updateTime="1698662999642" updateUser="ExternalLinkService">
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<mods:title id="D854B45DD2566D11A475FCD1786B1AE3">The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes</mods:title>
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<mods:namePart id="71FE6E7F5C9C0D535F30A2A5FC4431A9">Matz, Jess</mods:namePart>
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<mods:date id="F98DA339AA51C44AFB6EDF15A7B8B1F1">2016</mods:date>
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<treatment id="03F187D7FFCF8439FF11FD84FD84BDCF" ID-DOI="http://doi.org/10.5281/zenodo.6070109" ID-GBIF-Taxon="120828433" ID-Zenodo-Dep="6070109" LSID="urn:lsid:plazi:treatment:03F187D7FFCF8439FF11FD84FD84BDCF" httpUri="http://treatment.plazi.org/id/03F187D7FFCF8439FF11FD84FD84BDCF" lastPageNumber="69" pageId="68" pageNumber="69">
<subSubSection id="C342654AFFCF8439FF11FD84FD56BB26" box="[151,655,548,574]" pageId="68" pageNumber="69" type="nomenclature">
<paragraph id="8BE736C1FFCF8439FF11FD84FD56BB26" blockId="68.[151,655,548,574]" box="[151,655,548,574]" pageId="68" pageNumber="69">
<heading id="D0AF81ADFFCF8439FF11FD84FD56BB26" bold="true" box="[151,655,548,574]" fontSize="11" level="1" pageId="68" pageNumber="69" reason="1">
<taxonomicName id="4C584D42FFCF8439FF11FD84FD56BB26" ID-CoL="92P4X" authority="C. Felder &amp; R. Felder, 1867" authorityName="C. Felder &amp; R. Felder" authorityYear="1867" box="[151,655,548,574]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCF8439FF11FD84FD56BB26" bold="true" box="[151,655,548,574]" pageId="68" pageNumber="69">
<emphasis id="B92CEAD3FFCF8439FF11FD84FEF6BB26" bold="true" box="[151,303,548,574]" italics="true" pageId="68" pageNumber="69">Tetraphlebia</emphasis>
<bibRefCitation id="EFC94B30FFCF8439FEB0FD84FD56BB26" author="Felder" box="[310,655,548,574]" pageId="68" pageNumber="69" refString="Felder, C. &amp; Felder, R. (1867) Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1859, 1859 unter den Befehlen des Commodore B. von Wullensdorf - Urbair. Zoologischer Thiel. Zweiter Band. Zweite Abteilung: Lepidoptera. Carl Gerold's Sohn, Wien, 536 + 9 pp., 74 pls." type="book" year="1867">C. Felder &amp; R. Felder, 1867</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCF8439FF11FDCAFD27BBA6" pageId="68" pageNumber="69" type="materials_examined">
<paragraph id="8BE736C1FFCF8439FF11FDCAFD5FBB87" blockId="68.[151,766,618,702]" pageId="68" pageNumber="69">
<emphasis id="B92CEAD3FFCF8439FF11FDCAFEFCBB98" bold="true" box="[151,293,618,640]" pageId="68" pageNumber="69">
<typeStatus id="54E38863FFCF8439FF11FDCAFF15BB98" box="[151,204,618,640]" pageId="68" pageNumber="69">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C584D42FFCF8439FEADFDCBFD11BB98" authority="C. Felder &amp; R. Felder, 1867" authorityName="C. Felder &amp; R. Felder" authorityYear="1867" box="[299,712,618,640]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="germainii">
<emphasis id="B92CEAD3FFCF8439FEADFDCBFE7EBB98" box="[299,423,619,640]" italics="true" pageId="68" pageNumber="69">T. germainii</emphasis>
<bibRefCitation id="EFC94B30FFCF8439FE2BFDCAFD11BB98" author="Felder" box="[429,712,618,640]" pageId="68" pageNumber="69" refString="Felder, C. &amp; Felder, R. (1867) Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1859, 1859 unter den Befehlen des Commodore B. von Wullensdorf - Urbair. Zoologischer Thiel. Zweiter Band. Zweite Abteilung: Lepidoptera. Carl Gerold's Sohn, Wien, 536 + 9 pp., 74 pls." type="book" year="1867">C. Felder &amp; R. Felder, 1867</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C584D42FFCF8439FF2DFD29FDFABB87" authority="C. Felder &amp; R. Felder, 1867" authorityName="C. Felder &amp; R. Felder" authorityYear="1867" box="[171,547,649,671]" class="Insecta" family="Nymphalidae" genus="Faunula" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCF8439FF2DFD29FED8BB87" box="[171,257,649,671]" italics="true" pageId="68" pageNumber="69">Faunula</emphasis>
<bibRefCitation id="EFC94B30FFCF8439FE81FD29FDFABB87" author="Felder" box="[263,547,649,671]" pageId="68" pageNumber="69" refString="Felder, C. &amp; Felder, R. (1867) Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1859, 1859 unter den Befehlen des Commodore B. von Wullensdorf - Urbair. Zoologischer Thiel. Zweiter Band. Zweite Abteilung: Lepidoptera. Carl Gerold's Sohn, Wien, 536 + 9 pp., 74 pls." type="book" year="1867">C. Felder &amp; R. Felder, 1867</bibRefCitation>
</taxonomicName>
<emphasis id="B92CEAD3FFCF8439FDAFFD29FD5FBB87" bold="true" box="[553,646,649,671]" pageId="68" pageNumber="69">syn. nov.</emphasis>
</paragraph>
<paragraph id="8BE736C1FFCF8439FF41FD09FD27BBA6" blockId="68.[151,766,618,702]" box="[199,766,680,702]" pageId="68" pageNumber="69">
<typeStatus id="54E38863FFCF8439FF41FD09FF20BBA6" box="[199,249,681,702]" pageId="68" pageNumber="69">Type</typeStatus>
species:
<taxonomicName id="4C584D42FFCF8439FED0FD09FD27BBA6" authority="C. Felder &amp; R. Felder, 1867" authorityName="C. Felder &amp; R. Felder" authorityYear="1867" box="[342,766,680,702]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="leucoglene">
<emphasis id="B92CEAD3FFCF8439FED0FD09FE05BBA6" box="[342,476,680,702]" italics="true" pageId="68" pageNumber="69">F. leucoglene</emphasis>
<bibRefCitation id="EFC94B30FFCF8439FE65FD08FD27BBA6" author="Felder" box="[483,766,680,702]" pageId="68" pageNumber="69" refString="Felder, C. &amp; Felder, R. (1867) Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1859, 1859 unter den Befehlen des Commodore B. von Wullensdorf - Urbair. Zoologischer Thiel. Zweiter Band. Zweite Abteilung: Lepidoptera. Carl Gerold's Sohn, Wien, 536 + 9 pp., 74 pls." type="book" year="1867">C. Felder &amp; R. Felder, 1867</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCF8439FF11FD4AFAF7BD73" pageId="68" pageNumber="69" type="diagnosis">
<paragraph id="8BE736C1FFCF8439FF11FD4AFAF7BD73" blockId="68.[151,1437,746,1239]" pageId="68" pageNumber="69">
<emphasis id="B92CEAD3FFCF8439FF11FD4AFEC8BA1B" bold="true" box="[151,273,746,771]" pageId="68" pageNumber="69">Diagnosis.</emphasis>
<taxonomicName id="4C584D42FFCF8439FE9DFD4AFE70BA1B" box="[283,425,746,771]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCF8439FE9DFD4AFE70BA1B" box="[283,425,746,771]" italics="true" pageId="68" pageNumber="69">Tetraphlebia</emphasis>
</taxonomicName>
has a well-developed M1-M3 ocellus on the ventral side of the forewing with variable pupillation. In
<taxonomicName id="4C584D42FFCF8439FEBAFCAFFE09BA3F" box="[316,464,782,807]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="leucoglene">
<emphasis id="B92CEAD3FFCF8439FEBAFCAFFE09BA3F" box="[316,464,782,807]" italics="true" pageId="68" pageNumber="69">T. leucoglene</emphasis>
</taxonomicName>
, males have heavy androconia, but these scales are sparse in
<taxonomicName id="4C584D42FFCF8439FBFCFCAFFADBBA3E" box="[1146,1282,783,806]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="germainii">
<emphasis id="B92CEAD3FFCF8439FBFCFCAFFADBBA3E" box="[1146,1282,783,806]" italics="true" pageId="68" pageNumber="69">T. germainii</emphasis>
</taxonomicName>
and absent in
<taxonomicName id="4C584D42FFCF8439FF11FC94FEDBBA53" box="[151,258,818,843]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="eleates">
<emphasis id="B92CEAD3FFCF8439FF11FC94FEDBBA53" box="[151,258,818,843]" italics="true" pageId="68" pageNumber="69">T. eleates</emphasis>
</taxonomicName>
. The hindwing is rectangular with a postmedian band on the ventral side that is without ocelli. Though specimens were unavailable for dissection,
<taxonomicName id="4C584D42FFCF8439FDF8FCF7FBECBA77" authority="(Mabille, 1855) (Erebia)" authorityName="Mabille" authorityYear="1855" baseAuthorityName="Mabille" baseAuthorityYear="1855" box="[638,1077,854,879]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="patagonica">
<emphasis id="B92CEAD3FFCF8439FDF8FCF7FCCEBA76" box="[638,791,855,878]" italics="true" pageId="68" pageNumber="69">T. patagonica</emphasis>
(Mabille, 1855) (
<emphasis id="B92CEAD3FFCF8439FC58FCF6FBF0BA77" box="[990,1065,854,879]" italics="true" pageId="68" pageNumber="69">Erebia</emphasis>
)
</taxonomicName>
, lately a species of
<taxonomicName id="4C584D42FFCF8439FA96FCF6FAB6BA77" box="[1296,1391,854,879]" class="Insecta" family="Nymphalidae" genus="Faunula" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCF8439FA96FCF6FAB6BA77" box="[1296,1391,854,879]" italics="true" pageId="68" pageNumber="69">Faunula</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFC94B30FFCF8439FF11FCDBFEC2BA8B" author="Pyrcz" box="[151,283,891,915]" pageId="68" pageNumber="69" refString="Pyrcz, T. W. (2012) A new species of satyrine butterfly from Patagonia in more than a century and revisional notes on the genus Faunula C. Felder &amp; R. Felder (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa, 3342, 60 - 68." type="journal article" year="2012">Pyrcz 2012</bibRefCitation>
), is very similar to
<taxonomicName id="4C584D42FFCF8439FE7EFCDCFD54BA8B" box="[504,653,890,915]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="leucoglene">
<emphasis id="B92CEAD3FFCF8439FE7EFCDCFD54BA8B" box="[504,653,890,915]" italics="true" pageId="68" pageNumber="69">T. leucoglene</emphasis>
</taxonomicName>
in that it bears an identical M1-M3 VFW ocellus with a large, white pupil, but
<taxonomicName id="4C584D42FFCF8439FE8DFC3FFE7FBAAE" box="[267,422,927,950]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="patagonica">
<emphasis id="B92CEAD3FFCF8439FE8DFC3FFE7FBAAE" box="[267,422,927,950]" italics="true" pageId="68" pageNumber="69">T. patagonica</emphasis>
</taxonomicName>
also bears two ocelli on the VHW, making it the only
<taxonomicName id="4C584D42FFCF8439FB93FC3EFB7ABAAF" box="[1045,1187,926,951]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCF8439FB93FC3EFB7ABAAF" box="[1045,1187,926,951]" italics="true" pageId="68" pageNumber="69">Tetraphlebia</emphasis>
</taxonomicName>
with hindwing ocelli. Antennae are round in
<taxonomicName id="4C584D42FFCF8439FE24FC64FDD4BAC3" box="[418,525,962,987]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="eleates">
<emphasis id="B92CEAD3FFCF8439FE24FC64FDD4BAC3" box="[418,525,962,987]" italics="true" pageId="68" pageNumber="69">T. eleates</emphasis>
</taxonomicName>
and
<taxonomicName id="4C584D42FFCF8439FDCCFC64FD38BAC3" box="[586,737,962,987]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="leucoglene">
<emphasis id="B92CEAD3FFCF8439FDCCFC64FD38BAC3" box="[586,737,962,987]" italics="true" pageId="68" pageNumber="69">T. leucoglene</emphasis>
</taxonomicName>
, spatulate in
<taxonomicName id="4C584D42FFCF8439FCF8FC64FBD0BAC3" box="[894,1033,964,987]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="germainii">
<emphasis id="B92CEAD3FFCF8439FCF8FC64FBD0BAC3" box="[894,1033,964,987]" italics="true" pageId="68" pageNumber="69">T. germainii</emphasis>
</taxonomicName>
. Eyes are naked and terminal palp segment is short and conical or oval. Foreleg tarsi are segmented with males having 2–3 tarsal segments and females 3–5 segments, those of
<taxonomicName id="4C584D42FFCF8439FE78FBACFD4ABD3B" box="[510,659,1034,1059]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="leucoglene">
<emphasis id="B92CEAD3FFCF8439FE78FBACFD4ABD3B" box="[510,659,1034,1059]" italics="true" pageId="68" pageNumber="69">T. leucoglene</emphasis>
</taxonomicName>
females also bearing spines. Male genitalia with the proximal end of the valvae triangular, a wide pedunculus, an aedeagus that is truncate at the proximal end, a saccus longer than it is wide, and an uncus that is widest where it meets the tegumen, narrowing gradually toward the distal end.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCF8439FF41FBD6FD84BDCF" pageId="68" pageNumber="69" type="discussion">
<paragraph id="8BE736C1FFCF8439FF41FBD6FD84BDCF" blockId="68.[151,1437,746,1239]" pageId="68" pageNumber="69">
<emphasis id="B92CEAD3FFCF8439FF41FBD6FEE2BD97" bold="true" box="[199,315,1142,1167]" pageId="68" pageNumber="69">Remarks.</emphasis>
Felder &amp; Felder described
<taxonomicName id="4C584D42FFCF8439FD0CFBD6FD30BD97" box="[650,745,1142,1167]" class="Insecta" family="Nymphalidae" genus="Faunula" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCF8439FD0CFBD6FD30BD97" box="[650,745,1142,1167]" italics="true" pageId="68" pageNumber="69">Faunula</emphasis>
</taxonomicName>
as a new genus immediately following the description of
<taxonomicName id="4C584D42FFCF8439FF11FB3AFEFCBDAB" box="[151,293,1178,1203]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCF8439FF11FB3AFEFCBDAB" box="[151,293,1178,1203]" italics="true" pageId="68" pageNumber="69">Tetraphlebia</emphasis>
</taxonomicName>
, noting similarities in wing venation between the two genera. These genera are here combined based on morphological and genetic similarity.
</paragraph>
</subSubSection>
</treatment>
</document>
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