332 lines
50 KiB
XML
332 lines
50 KiB
XML
<document id="346ED442562EE3E251BA5F08B56CF5E1" ID-CLB-Dataset="32554" ID-DOI="10.11646/zootaxa.4300.4.3" ID-GBIF-Dataset="c47ff596-c049-46e1-b6c6-d0cd9ded095e" ID-ISSN="1175-5326" ID-Zenodo-Dep="839214" ID-ZooBank="6557F0CC-1558-431F-81AB-A4D6191FCB15" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501933676727" checkinUser="plazi" docAuthor="Paxton, Hannelore & Arias, Andres" docDate="2017" docId="03DD0635D62FFFE8FF0FFEB1FB3BFDE7" docLanguage="en" docName="zoobank.4300.4.3.pdf.imf" docOrigin="Zootaxa 4300 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diopatra mariae Paxton & Arias 2017, sp. nov." docType="treatment" docVersion="10" lastPageNumber="523" masterDocId="FFE47E4DD63FFFFAFF98FF96FFA0FF87" masterDocTitle="Unveiling a surprising diversity of the genus Diopatra Audouin & Milne Edwards, 1833 (Annelida: Onuphidae) in the Macaronesian region (eastern North Atlantic) with the description of four new species" masterLastPageNumber="535" masterPageNumber="505" pageNumber="521" updateTime="1698466220759" updateUser="ExternalLinkService">
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<mods:titleInfo id="5D972A7FAC3367129C25694C3B62F6B4">
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<mods:title id="E17FF380A950E159FB6F2C98C0AF887E">Unveiling a surprising diversity of the genus Diopatra Audouin & Milne Edwards, 1833 (Annelida: Onuphidae) in the Macaronesian region (eastern North Atlantic) with the description of four new species</mods:title>
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<mods:name id="1B9FD72AB1255B9B7F284B89E205F9B0" type="personal">
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<mods:roleTerm id="C2428B33EE8BED96F8E50C40C7D0E5C7">Author</mods:roleTerm>
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<mods:namePart id="58D428D56E5C1839A31439801B0A7566">Paxton, Hannelore</mods:namePart>
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<mods:name id="7BF11FEF6D5A7BB2AF5FE3E165C97530" type="personal">
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<mods:role id="2D205A5C456124382D53F1A6933B1AFB">
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<mods:roleTerm id="B9479B0132C12ED38299B4242F64E3DC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A731B47A0AF5F1DCE474E149F4EE95CD">Arias, Andres</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3B03FEAB40D3CCA10EE1A577C309B19A">text</mods:typeOfResource>
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<mods:title id="AFD56E0E591538771FC9D0EDF8C1BEBE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9E075E621D43E6114F41FE3CCA250AB8">
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<mods:date id="D715F9DBF7DA1D701704D3DDE25096E9">2017</mods:date>
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<mods:number id="88C8900D7DF34A5D629246F609F7816A">4300</mods:number>
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<mods:detail id="59305A4FFE22B6EC4A52262ED31BDF01" type="issue">
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<mods:number id="B099B4A5986FB2E4891F0F658C5724AB">4</mods:number>
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<mods:start id="40139A13BF80CE600DE39064B0B48B21">505</mods:start>
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<mods:classification id="B5DC5F39D7E35C67C9C49F77BCF0B58B">journal article</mods:classification>
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<mods:identifier id="15CDC9D20BFF72FD1E5DEEDEB07B7DAD" type="CLB-Dataset">32554</mods:identifier>
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<mods:identifier id="21EC202E42F09D280DB9429BFE6C8A50" type="DOI">10.11646/zootaxa.4300.4.3</mods:identifier>
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<treatment id="03DD0635D62FFFE8FF0FFEB1FB3BFDE7" ID-DOI="http://doi.org/10.5281/zenodo.5492081" ID-GBIF-Taxon="132630487" ID-Zenodo-Dep="5492081" LSID="urn:lsid:plazi:treatment:03DD0635D62FFFE8FF0FFEB1FB3BFDE7" httpUri="http://treatment.plazi.org/id/03DD0635D62FFFE8FF0FFEB1FB3BFDE7" lastPageId="18" lastPageNumber="523" pageId="16" pageNumber="521">
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<subSubSection id="C36EE4A8D62FFFEAFF0FFEB1FDA0FEE4" pageId="16" pageNumber="521" type="nomenclature">
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<paragraph id="8BCBB723D62FFFEAFF0FFEB1FE63FEC5" blockId="16.[151,512,295,355]" box="[151,451,295,322]" pageId="16" pageNumber="521">
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<heading id="D083004FD62FFFEAFF0FFEB1FE63FEC5" bold="true" box="[151,451,295,322]" fontSize="11" level="1" pageId="16" pageNumber="521" reason="1">
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<emphasis id="B9006B31D62FFFEAFF0FFEB1FE63FEC5" bold="true" box="[151,451,295,322]" pageId="16" pageNumber="521">
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<taxonomicName id="4C74CCA0D62FFFEAFF0FFEB1FEFFFEC6" ID-CoL="36BFL" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[151,351,295,321]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="521" phylum="Annelida" rank="species" species="mariae" status="sp. nov.">
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<emphasis id="B9006B31D62FFFEAFF0FFEB1FEFFFEC6" bold="true" box="[151,351,295,321]" italics="true" pageId="16" pageNumber="521">Diopatra mariae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A233D64AD62FFFEAFEFDFEBEFE63FEC5" box="[357,451,296,322]" pageId="16" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BCBB723D62FFFEAFF0FFEDCFDA0FEE4" blockId="16.[151,512,295,355]" box="[151,512,330,355]" pageId="16" pageNumber="521">
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<figureCitation id="134FABA6D62FFFEAFF0FFEDCFF5FFEE4" box="[151,255,330,355]" captionStart="FIGURE 7" captionStartId="10.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1668]" captionTargetId="figure@10.[151,1436,193,1668]" captionTargetPageId="10" captionText="FIGURE 7. Diopatra spp. photomicrographs. A, D. madeirensis (MNCN 16.01 / 17816), anterior end, dorsal view; B, D. madeirensis (AM W. 49213) anterior end, dorsal view; C, same, ventral view; D, tube of same; E, D. mariae sp. nov., paratype (MNCN 16.01 / 17819), anterior end, dorsal view; F, D. mariae sp. nov., holotype (MNCN 16.01 / 17818), anterior end, ventrolateral view; G, D. mariae sp. nov., paratype (MNCN 16.01 / 17819) everted maxillae, ventral view." httpUri="https://zenodo.org/record/839228/files/figure.png" pageId="16" pageNumber="521">Figures 7</figureCitation>
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E–G, 10, 11;
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<tableCitation id="C6F68298D62FFFEAFE0AFEDCFE44FEE4" box="[402,484,330,355]" captionStart="TABLE 1" captionStartId="14.[119,197,513,534]" captionText="TABLE 1. CƟmparisƟn Ɵf Diopatra speƇies treateđ in this paper." pageId="16" pageNumber="521">Table 1</tableCitation>
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,
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<tableCitation id="C6F68298D62FFFEAFE6AFEDDFDA0FEE4" box="[498,512,331,355]" captionStart="TABLE 2" captionStartId="25.[125,198,565,586]" captionText="TABLE 2. List Ɵf Diopatra speƇies with đƟuble pƟstƇhaetal lƟbes. PC, pseuđƟƇƟmpƟunđ hƟƟk." pageId="16" pageNumber="521">2</tableCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36EE4A8D62FFFEAFF0FFE04FE89FDBC" pageId="16" pageNumber="521" type="materials_examined">
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<paragraph id="8BCBB723D62FFFEAFF0FFE04FEF5FE74" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
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<materialsCitation id="3B1CBD7ED62FFFEAFF0FFE04FC7CFE48" ID-GBIF-Occurrence="1585879931" collectingDate="2011-08-10" collectionCode="MNCN" collectorName="A. Arias" latitude="28.842222" location="Lanzarote" longLatPrecision="20" longitude="-13.788334" municipality="Papagayo" pageId="16" pageNumber="521" specimenCode="MNCN 16.01" specimenCount="2" stateProvince="Canary Islands" typeStatus="holotype">
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<emphasis id="B9006B31D62FFFEAFF0FFE04FDE3FE2C" bold="true" box="[151,579,402,427]" pageId="16" pageNumber="521">
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Material examined.
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<typeStatus id="54CF0981D62FFFEAFE0DFE04FE70FE2C" box="[405,464,402,427]" pageId="16" pageNumber="521">Type</typeStatus>
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||
material
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||
</emphasis>
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||
.
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||
<emphasis id="B9006B31D62FFFEAFDC1FE04FD6FFE2C" bold="true" box="[601,719,402,427]" pageId="16" pageNumber="521">
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||
<typeStatus id="54CF0981D62FFFEAFDC1FE04FD6AFE2C" box="[601,714,402,427]" pageId="16" pageNumber="521" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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||
(
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<specimenCode id="DBD21F58D62FFFEAFD7DFE05FC28FE2C" box="[741,904,402,427]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="16" pageNumber="521">
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||
<collectionCode id="ED652FE6D62FFFEAFD7DFE05FC9BFE2C" box="[741,827,403,427]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="16" pageNumber="521">MNCN</collectionCode>
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16.01
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</specimenCode>
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/17818),
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<collectingMunicipality id="6BAF2D59D62FFFEAFC6CFE05FBC1FE2C" box="[1012,1121,403,427]" pageId="16" pageNumber="521">Papagayo</collectingMunicipality>
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||
beach,
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||
<location id="8EABE1F8D62FFFEAFB5EFE05FA99FE2D" LSID="urn:lsid:plazi:treatment:03DD0635D62FFFE8FF0FFEB1FB3BFDE7:8EABE1F8D62FFFEAFB5EFE05FA99FE2D" box="[1222,1337,403,426]" latitude="28.842222" longLatPrecision="20" longitude="-13.788334" municipality="Papagayo" name="Lanzarote" pageId="16" pageNumber="521" stateProvince="Canary Islands">Lanzarote</location>
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,
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||
<collectingRegion id="49B079C1D62FFFEAFAD3FE05FF49FE48" country="Spain" name="Canarias" pageId="16" pageNumber="521">Canary Islands</collectingRegion>
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||
,
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||
<geoCoordinate id="EE40D1E4D62FFFEAFF6CFE20FE25FE48" box="[244,389,438,463]" direction="north" orientation="latitude" pageId="16" pageNumber="521" precision="15" value="28.842222">28°50’32” N</geoCoordinate>
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–
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||
<geoCoordinate id="EE40D1E4D62FFFEAFE0DFE21FD84FE49" box="[405,548,439,463]" direction="west" orientation="longitude" pageId="16" pageNumber="521" precision="15" value="-13.788334">13°47’18” W</geoCoordinate>
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, intertidal, coll.
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<collectorName id="2681D2F5D62FFFEAFD43FE21FC9CFE48" box="[731,828,438,463]" pageId="16" pageNumber="521">A. Arias</collectorName>
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,
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||
<date id="FFCA91E3D62FFFEAFCD0FE21FC7CFE48" box="[840,988,439,463]" pageId="16" pageNumber="521" value="2011-08-10">
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||
<collectingDate id="EF8E680BD62FFFEAFCD0FE21FC7CFE48" box="[840,988,439,463]" pageId="16" pageNumber="521" value="2011-08-10">10 Aug 2011</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B1CBD7ED62FFFEAFC7FFE20FEF5FE74" ID-GBIF-Occurrence="1585879929" collectingDate="2011-08-10" collectionCode="MNCN" collectorName="A. Arias" latitude="28.842222" location="Lanzarote" longLatPrecision="20" longitude="-13.788334" municipality="Papagayo" pageId="16" pageNumber="521" specimenCode="MNCN 16.01" specimenCount="1" stateProvince="Canary Islands" typeStatus="paratype">
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||
<emphasis id="B9006B31D62FFFEAFC7FFE20FBF3FE48" bold="true" box="[999,1107,438,463]" pageId="16" pageNumber="521">
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||
<typeStatus id="54CF0981D62FFFEAFC7FFE20FBF3FE48" box="[999,1107,438,463]" pageId="16" pageNumber="521" type="paratype">paratype</typeStatus>
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||
</emphasis>
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||
(
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||
<specimenCode id="DBD21F58D62FFFEAFBF9FE21FB5DFE48" box="[1121,1277,438,463]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="16" pageNumber="521">
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||
<collectionCode id="ED652FE6D62FFFEAFBF9FE21FB17FE48" box="[1121,1207,439,463]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="16" pageNumber="521">MNCN</collectionCode>
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16.01
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||
</specimenCode>
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/17819), same data as holotype.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCBB723D62FFFEAFF5FFE68FE89FDBC" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
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||
<materialsCitation id="3B1CBD7ED62FFFEAFF5FFE68FE89FDBC" ID-GBIF-Occurrence="1585879932" county="Eastern North Atlantic" latitude="28.842222" location="Papagayo" longLatPrecision="20" longitude="-13.788334" municipality="Lanzarote" pageId="16" pageNumber="521" specimenCount="1" stateProvince="Canary Islands" typeStatus="holotype">
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||
<emphasis id="B9006B31D62FFFEAFF5FFE68FECBFD90" bold="true" box="[199,363,510,535]" pageId="16" pageNumber="521">
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||
<typeStatus id="54CF0981D62FFFEAFF5FFE68FEA2FD90" box="[199,258,510,535]" pageId="16" pageNumber="521">Type</typeStatus>
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locality:
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||
</emphasis>
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||
<collectingCounty id="62AACFAFD62FFFEAFEEDFE69FDDAFD90" box="[373,634,510,535]" pageId="16" pageNumber="521">Eastern North Atlantic</collectingCounty>
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||
,
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||
<collectingRegion id="49B079C1D62FFFEAFD10FE69FC95FD90" box="[648,821,510,535]" country="Spain" name="Canarias" pageId="16" pageNumber="521">Canary Islands</collectingRegion>
|
||
,
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||
<collectingMunicipality id="6BAF2D59D62FFFEAFCDBFE69FC16FD91" box="[835,950,511,534]" pageId="16" pageNumber="521">Lanzarote</collectingMunicipality>
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||
,
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||
<location id="8EABE1F8D62FFFEAFC5BFE69FB90FD90" LSID="urn:lsid:plazi:treatment:03DD0635D62FFFE8FF0FFEB1FB3BFDE7:8EABE1F8D62FFFEAFC5BFE69FB90FD90" box="[963,1072,511,535]" county="Eastern North Atlantic" latitude="28.842222" longLatPrecision="20" longitude="-13.788334" municipality="Lanzarote" name="Papagayo" pageId="16" pageNumber="521" stateProvince="Canary Islands">Papagayo</location>
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||
beach,
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||
<geoCoordinate id="EE40D1E4D62FFFEAFB13FE68FABFFD90" box="[1163,1311,510,535]" direction="north" orientation="latitude" pageId="16" pageNumber="521" precision="15" value="28.842222">28°50’32” N</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="EE40D1E4D62FFFEAFAB7FE69FF12FDBD" direction="west" orientation="longitude" pageId="16" pageNumber="521" precision="15" value="-13.788334">13°47’18” W</geoCoordinate>
|
||
, intertidal.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36EE4A8D62FFFEAFF5FFDD0FE75FC94" pageId="16" pageNumber="521" type="diagnosis">
|
||
<paragraph id="8BCBB723D62FFFEAFF5FFDD0FE75FC94" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
|
||
<emphasis id="B9006B31D62FFFEAFF5FFDD0FEE1FDD8" bold="true" box="[199,321,582,607]" pageId="16" pageNumber="521">Diagnosis.</emphasis>
|
||
Prostomium anteriorly rounded with two subulate frontal lips; antennae to chaetiger 13–15 with seven to nine ceratophoral rings, lateral projections absent; nuchal grooves crescentic; peristomial cirri present. Anterior four pairs of parapodia with bidentate pseudocompound hooks with pointed hoods; with upper subulate and lower knob-like postchaetal lobes. Ventral parapodial lobes absent, ventral cirri on five chaetigers. Subacicular hooks from chaetiger 17; pectinate chaetae with 18–22 teeth; spiralled branchiae, first on chaetiger 4, single filaments from chaetiger 50.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36EE4A8D62FFFEAFF5FFC88FC6BF814" pageId="16" pageNumber="521" type="description">
|
||
<paragraph id="8BCBB723D62FFFEAFF5FFC88FC7FFCF8" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
|
||
<emphasis id="B9006B31D62FFFEAFF5FFC88FEF9FCB0" bold="true" box="[199,345,798,823]" pageId="16" pageNumber="521">Description.</emphasis>
|
||
Both
|
||
<typeStatus id="54CF0981D62FFFEAFE05FC89FE78FCB0" box="[413,472,799,823]" pageId="16" pageNumber="521">types</typeStatus>
|
||
incomplete,
|
||
<typeStatus id="54CF0981D62FFFEAFDF1FC88FD6BFCB0" box="[617,715,798,823]" pageId="16" pageNumber="521" type="holotype">holotype</typeStatus>
|
||
<quantity id="4C8C1AC6D62FFFEAFD4AFC89FC81FCB1" box="[722,801,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="521" unit="mm" value="28.0">28 mm</quantity>
|
||
long for 86 chaetigers,
|
||
<quantity id="4C8C1AC6D62FFFEAFBB3FC88FB21FCB1" box="[1067,1153,798,823]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="521" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide;
|
||
<typeStatus id="54CF0981D62FFFEAFB54FC89FA8DFCB0" box="[1228,1325,799,823]" pageId="16" pageNumber="521" type="paratype">paratype</typeStatus>
|
||
<quantity id="4C8C1AC6D62FFFEAFAACFC88FAD5FCB1" box="[1332,1397,798,823]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="521" unit="mm" value="6.0">6 mm</quantity>
|
||
for 30 chaetigers,
|
||
<quantity id="4C8C1AC6D62FFFEAFEA4FCD5FE33FCDD" box="[316,403,835,859]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="521" unit="mm" value="0.9">0.9 mm</quantity>
|
||
wide. Ethanol stored specimen overall very pale brown with some darker pigmentation on prostomium and anterior chaetigers, no distinct colour pattern (
|
||
<figureCitation id="134FABA6D62FFFEAFCC2FCF0FC00FCF9" box="[858,928,870,895]" captionStart="FIGURE 7" captionStartId="10.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1668]" captionTargetId="figure@10.[151,1436,193,1668]" captionTargetPageId="10" captionText="FIGURE 7. Diopatra spp. photomicrographs. A, D. madeirensis (MNCN 16.01 / 17816), anterior end, dorsal view; B, D. madeirensis (AM W. 49213) anterior end, dorsal view; C, same, ventral view; D, tube of same; E, D. mariae sp. nov., paratype (MNCN 16.01 / 17819), anterior end, dorsal view; F, D. mariae sp. nov., holotype (MNCN 16.01 / 17818), anterior end, ventrolateral view; G, D. mariae sp. nov., paratype (MNCN 16.01 / 17819) everted maxillae, ventral view." httpUri="https://zenodo.org/record/839228/files/figure.png" pageId="16" pageNumber="521">Figs 7</figureCitation>
|
||
E, F).
|
||
</paragraph>
|
||
<paragraph id="8BCBB723D62FFFEAFF5FFC1CFAA7FBFD" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
|
||
Prostomium anteriorly rounded with two subulate frontal lips separated by a small gap (
|
||
<figureCitation id="134FABA6D62FFFEAFB03FC1CFB7EFC24" box="[1179,1246,906,931]" captionStart="FIGURE 7" captionStartId="10.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1668]" captionTargetId="figure@10.[151,1436,193,1668]" captionTargetPageId="10" captionText="FIGURE 7. Diopatra spp. photomicrographs. A, D. madeirensis (MNCN 16.01 / 17816), anterior end, dorsal view; B, D. madeirensis (AM W. 49213) anterior end, dorsal view; C, same, ventral view; D, tube of same; E, D. mariae sp. nov., paratype (MNCN 16.01 / 17819), anterior end, dorsal view; F, D. mariae sp. nov., holotype (MNCN 16.01 / 17818), anterior end, ventrolateral view; G, D. mariae sp. nov., paratype (MNCN 16.01 / 17819) everted maxillae, ventral view." httpUri="https://zenodo.org/record/839228/files/figure.png" pageId="16" pageNumber="521">Fig. 7</figureCitation>
|
||
E). Ventral upper lips oval with papilla-like median section between lips; lower lip with median section. Palps reaching to chaetiger 3, lateral antennae to chaetiger 15, median antenna to chaetiger 13; ceratophores with six to eight proximal and a longer distal ring, styles tapering to distal end, with fine tips. Ceratostyles with about 20 irregular longitudinal rows of flat sensory buds; buds forming semicircles (
|
||
<figureCitation id="134FABA6D62FFFEAFD50FB8CFCBCFBB4" box="[712,796,1050,1075]" captionStart="FIGURE 10" captionStartId="17.[151,250,1710,1732]" captionTargetBox="[151,1435,193,1689]" captionTargetId="figure@17.[151,1435,193,1689]" captionTargetPageId="17" captionText="FIGURE 10. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), scanning electron micrographs. A, Sensory buds of antenna; B, parapodium of chaetiger 4, posterior view; C, same magnified, note lower postchaetal lobe folded onto parapodium; D, parapodium of chaetiger 20, anterior view; E, parapodium of chaetiger 42, anterior view; F, pectinate chaetae. Abbreviations: a, aciculae, b, branchia, dc = dorsal cirrus, = limbate chaeta, lPCL = lower postchaetal lobe; p = pectinate chaeta, PrCL = prechaetal lobe; SAHs = subacicular hooks; uPCL = upper postchaetal lobe; vc = ventral cirrus." httpUri="https://zenodo.org/record/839234/files/figure.png" pageId="16" pageNumber="521">Fig. 10</figureCitation>
|
||
A) to almost full circles. Nuchal grooves very shallow crescents, hardly visible below contracted peristomium; peristomial cirri slender, twice as long as peristomium, reaching distal tip of palpophores, inserted distally on peristomium, almost lateral to lateral antennae.
|
||
</paragraph>
|
||
<paragraph id="8BCBB723D62FFFEAFF5FFB10FC8FF980" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
|
||
First four pairs of parapodia modified, slightly prolonged and directed anteroventrally (
|
||
<figureCitation id="134FABA6D62FFFEAFB20FB10FB5EFB18" box="[1208,1278,1158,1183]" captionStart="FIGURE 7" captionStartId="10.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1668]" captionTargetId="figure@10.[151,1436,193,1668]" captionTargetPageId="10" captionText="FIGURE 7. Diopatra spp. photomicrographs. A, D. madeirensis (MNCN 16.01 / 17816), anterior end, dorsal view; B, D. madeirensis (AM W. 49213) anterior end, dorsal view; C, same, ventral view; D, tube of same; E, D. mariae sp. nov., paratype (MNCN 16.01 / 17819), anterior end, dorsal view; F, D. mariae sp. nov., holotype (MNCN 16.01 / 17818), anterior end, ventrolateral view; G, D. mariae sp. nov., paratype (MNCN 16.01 / 17819) everted maxillae, ventral view." httpUri="https://zenodo.org/record/839228/files/figure.png" pageId="16" pageNumber="521">Fig. 7</figureCitation>
|
||
F). Prechaetal lobe rounded; large subulate upper and short, knob-like lower postchaetal lobes, lower postchaetal lobes equal in length to prechaetal lobe (
|
||
<figureCitation id="134FABA6D62FFFEAFE22FB58FDAFFB60" box="[442,527,1230,1255]" captionStart="FIGURE 10" captionStartId="17.[151,250,1710,1732]" captionTargetBox="[151,1435,193,1689]" captionTargetId="figure@17.[151,1435,193,1689]" captionTargetPageId="17" captionText="FIGURE 10. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), scanning electron micrographs. A, Sensory buds of antenna; B, parapodium of chaetiger 4, posterior view; C, same magnified, note lower postchaetal lobe folded onto parapodium; D, parapodium of chaetiger 20, anterior view; E, parapodium of chaetiger 42, anterior view; F, pectinate chaetae. Abbreviations: a, aciculae, b, branchia, dc = dorsal cirrus, = limbate chaeta, lPCL = lower postchaetal lobe; p = pectinate chaeta, PrCL = prechaetal lobe; SAHs = subacicular hooks; uPCL = upper postchaetal lobe; vc = ventral cirrus." httpUri="https://zenodo.org/record/839234/files/figure.png" pageId="16" pageNumber="521">Figs 10</figureCitation>
|
||
B, C, 11A). Prechaetal lobe becoming reduced, absent from chaetiger 8–9. Upper postchaetal lobe becoming smaller but remaining as little knob into posterior region; lower postchaetal knob absent from chaetiger 6. Dorsal cirri subulate, very long, attaining greatest length from chaetiger 5–10, almost half as long as branchial trunk (
|
||
<figureCitation id="134FABA6D62FFFEAFE18FAACFE68FAD4" box="[384,456,1338,1363]" captionStart="FIGURE 7" captionStartId="10.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1668]" captionTargetId="figure@10.[151,1436,193,1668]" captionTargetPageId="10" captionText="FIGURE 7. Diopatra spp. photomicrographs. A, D. madeirensis (MNCN 16.01 / 17816), anterior end, dorsal view; B, D. madeirensis (AM W. 49213) anterior end, dorsal view; C, same, ventral view; D, tube of same; E, D. mariae sp. nov., paratype (MNCN 16.01 / 17819), anterior end, dorsal view; F, D. mariae sp. nov., holotype (MNCN 16.01 / 17818), anterior end, ventrolateral view; G, D. mariae sp. nov., paratype (MNCN 16.01 / 17819) everted maxillae, ventral view." httpUri="https://zenodo.org/record/839228/files/figure.png" pageId="16" pageNumber="521">Fig. 7</figureCitation>
|
||
F), thereafter gradually shortening, becoming very slender posteriorly; ventral cirri subulate on anterior five chaetigers, with last being much shorter than preceding ones, thereafter replaced by ventral glandular pads. Ventral lobe absent. Spiralled branchiae (
|
||
<figureCitation id="134FABA6D62FFFEAFCE2FA14FC1EFA1D" box="[890,958,1410,1435]" captionStart="FIGURE 7" captionStartId="10.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1668]" captionTargetId="figure@10.[151,1436,193,1668]" captionTargetPageId="10" captionText="FIGURE 7. Diopatra spp. photomicrographs. A, D. madeirensis (MNCN 16.01 / 17816), anterior end, dorsal view; B, D. madeirensis (AM W. 49213) anterior end, dorsal view; C, same, ventral view; D, tube of same; E, D. mariae sp. nov., paratype (MNCN 16.01 / 17819), anterior end, dorsal view; F, D. mariae sp. nov., holotype (MNCN 16.01 / 17818), anterior end, ventrolateral view; G, D. mariae sp. nov., paratype (MNCN 16.01 / 17819) everted maxillae, ventral view." httpUri="https://zenodo.org/record/839228/files/figure.png" pageId="16" pageNumber="521">Fig. 7</figureCitation>
|
||
E, F) from chaetiger 4, best developed on chaetigers 5–10 with about 12 whorls of short, thin filaments on thin branchial trunk, with length of trunk almost equalling body width. Thereafter length of branchiae and number of filaments decreasing gradually (
|
||
<figureCitation id="134FABA6D62FFFEAFA9FFA5CFAF9FA64" box="[1287,1369,1482,1507]" captionStart="FIGURE 10" captionStartId="17.[151,250,1710,1732]" captionTargetBox="[151,1435,193,1689]" captionTargetId="figure@17.[151,1435,193,1689]" captionTargetPageId="17" captionText="FIGURE 10. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), scanning electron micrographs. A, Sensory buds of antenna; B, parapodium of chaetiger 4, posterior view; C, same magnified, note lower postchaetal lobe folded onto parapodium; D, parapodium of chaetiger 20, anterior view; E, parapodium of chaetiger 42, anterior view; F, pectinate chaetae. Abbreviations: a, aciculae, b, branchia, dc = dorsal cirrus, = limbate chaeta, lPCL = lower postchaetal lobe; p = pectinate chaeta, PrCL = prechaetal lobe; SAHs = subacicular hooks; uPCL = upper postchaetal lobe; vc = ventral cirrus." httpUri="https://zenodo.org/record/839234/files/figure.png" pageId="16" pageNumber="521">Fig. 10</figureCitation>
|
||
D, E), single filament from chaetiger 50, absent shortly thereafter.
|
||
</paragraph>
|
||
<paragraph id="8BCBB723D62FFFEAFF5FF984FC70F910" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
|
||
Modified parapodia (chaetigers 1–4) with one to two slender upper simple limbate chaetae and pseudocompound bidentate hooks with pointed hoods and shafts with two rows of tiny spines (
|
||
<figureCitation id="134FABA6D62FFFEAFB7CF9A0FA96F9C8" box="[1252,1334,1590,1615]" captionStart="FIGURE 11" captionStartId="18.[151,250,1426,1448]" captionTargetBox="[166,1415,674,1389]" captionTargetId="figure@18.[152,1435,663,1404]" captionTargetPageId="18" captionText="FIGURE 11. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), line drawings. A, Parapodium from chaetiger 2, anterior view; B, median robust pseudompound hook from chaetiger 2; C, upper slender pseudocompound hook from same; D, lower very slender pseudocompound hook from same; E, pectinate chaeta from chaetiger 42; F, subacicular hook from same. Abbreviations: lPCL = lower postchaetal lobe; uPCL = upper postchaetal lobe." httpUri="https://zenodo.org/record/839236/files/figure.png" pageId="16" pageNumber="521">Fig. 11</figureCitation>
|
||
A). Each parapodium with one median thicker hook (
|
||
<figureCitation id="134FABA6D62FFFEAFD17F9CCFD41F9F4" box="[655,737,1626,1651]" captionStart="FIGURE 11" captionStartId="18.[151,250,1426,1448]" captionTargetBox="[166,1415,674,1389]" captionTargetId="figure@18.[152,1435,663,1404]" captionTargetPageId="18" captionText="FIGURE 11. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), line drawings. A, Parapodium from chaetiger 2, anterior view; B, median robust pseudompound hook from chaetiger 2; C, upper slender pseudocompound hook from same; D, lower very slender pseudocompound hook from same; E, pectinate chaeta from chaetiger 42; F, subacicular hook from same. Abbreviations: lPCL = lower postchaetal lobe; uPCL = upper postchaetal lobe." httpUri="https://zenodo.org/record/839236/files/figure.png" pageId="16" pageNumber="521">Fig. 11</figureCitation>
|
||
B) and two to three more slender ones (
|
||
<figureCitation id="134FABA6D62FFFEAFB35F9CCFB5EF9F4" box="[1197,1278,1626,1651]" captionStart="FIGURE 11" captionStartId="18.[151,250,1426,1448]" captionTargetBox="[166,1415,674,1389]" captionTargetId="figure@18.[152,1435,663,1404]" captionTargetPageId="18" captionText="FIGURE 11. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), line drawings. A, Parapodium from chaetiger 2, anterior view; B, median robust pseudompound hook from chaetiger 2; C, upper slender pseudocompound hook from same; D, lower very slender pseudocompound hook from same; E, pectinate chaeta from chaetiger 42; F, subacicular hook from same. Abbreviations: lPCL = lower postchaetal lobe; uPCL = upper postchaetal lobe." httpUri="https://zenodo.org/record/839236/files/figure.png" pageId="16" pageNumber="521">Fig. 11</figureCitation>
|
||
C) from main pocket, and one to two very slender hooks
|
||
<figureCitation id="134FABA6D62FFFEAFDEFF9E8FD64F910" box="[631,708,1662,1687]" captionStart="FIGURE 11" captionStartId="18.[151,250,1426,1448]" captionTargetBox="[166,1415,674,1389]" captionTargetId="figure@18.[152,1435,663,1404]" captionTargetPageId="18" captionText="FIGURE 11. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), line drawings. A, Parapodium from chaetiger 2, anterior view; B, median robust pseudompound hook from chaetiger 2; C, upper slender pseudocompound hook from same; D, lower very slender pseudocompound hook from same; E, pectinate chaeta from chaetiger 42; F, subacicular hook from same. Abbreviations: lPCL = lower postchaetal lobe; uPCL = upper postchaetal lobe." httpUri="https://zenodo.org/record/839236/files/figure.png" pageId="16" pageNumber="521">Fig. 11</figureCitation>
|
||
D) from lower position.
|
||
</paragraph>
|
||
<paragraph id="8BCBB723D62FFFEAFF5FF934FF72F8A0" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
|
||
Unmodified parapodia (chaetigers 5 onwards) with pectinate and limbate chaetae. Pectinate chaetae slightly oblique with 18–22 teeth (
|
||
<figureCitation id="134FABA6D62FFFEAFE5FF950FDBDF958" box="[455,541,1734,1759]" captionStart="FIGURE 10" captionStartId="17.[151,250,1710,1732]" captionTargetBox="[151,1435,193,1689]" captionTargetId="figure@17.[151,1435,193,1689]" captionTargetPageId="17" captionText="FIGURE 10. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), scanning electron micrographs. A, Sensory buds of antenna; B, parapodium of chaetiger 4, posterior view; C, same magnified, note lower postchaetal lobe folded onto parapodium; D, parapodium of chaetiger 20, anterior view; E, parapodium of chaetiger 42, anterior view; F, pectinate chaetae. Abbreviations: a, aciculae, b, branchia, dc = dorsal cirrus, = limbate chaeta, lPCL = lower postchaetal lobe; p = pectinate chaeta, PrCL = prechaetal lobe; SAHs = subacicular hooks; uPCL = upper postchaetal lobe; vc = ventral cirrus." httpUri="https://zenodo.org/record/839234/files/figure.png" pageId="16" pageNumber="521">Figs 10</figureCitation>
|
||
F, 11E), about
|
||
<quantity id="4C8C1AC6D62FFFEAFD51F951FCA4F958" box="[713,772,1734,1759]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="521" unit="in" value="10.0">10 in</quantity>
|
||
median body region. Limbate chaetae becoming coarsely serrated by chaetiger 30–40; lower limbate chaetae replaced by bidentate subacicular hooks from chaetiger 17 (
|
||
<figureCitation id="134FABA6D62FFFEAFAF7F97CFF12F8A0" captionStart="FIGURE 11" captionStartId="18.[151,250,1426,1448]" captionTargetBox="[166,1415,674,1389]" captionTargetId="figure@18.[152,1435,663,1404]" captionTargetPageId="18" captionText="FIGURE 11. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), line drawings. A, Parapodium from chaetiger 2, anterior view; B, median robust pseudompound hook from chaetiger 2; C, upper slender pseudocompound hook from same; D, lower very slender pseudocompound hook from same; E, pectinate chaeta from chaetiger 42; F, subacicular hook from same. Abbreviations: lPCL = lower postchaetal lobe; uPCL = upper postchaetal lobe." httpUri="https://zenodo.org/record/839236/files/figure.png" pageId="16" pageNumber="521">Fig. 11</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
<paragraph id="8BCBB723D62FFFEAFF5FF8A4FC6BF814" blockId="16.[151,1437,402,1975]" pageId="16" pageNumber="521">
|
||
Mandibles and maxillae completely withdrawn in holotype; maxillary formula of paratype (
|
||
<figureCitation id="134FABA6D62FFFEAFB74F8A4FA92F8CC" box="[1260,1330,1842,1867]" captionStart="FIGURE 7" captionStartId="10.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1668]" captionTargetId="figure@10.[151,1436,193,1668]" captionTargetPageId="10" captionText="FIGURE 7. Diopatra spp. photomicrographs. A, D. madeirensis (MNCN 16.01 / 17816), anterior end, dorsal view; B, D. madeirensis (AM W. 49213) anterior end, dorsal view; C, same, ventral view; D, tube of same; E, D. mariae sp. nov., paratype (MNCN 16.01 / 17819), anterior end, dorsal view; F, D. mariae sp. nov., holotype (MNCN 16.01 / 17818), anterior end, ventrolateral view; G, D. mariae sp. nov., paratype (MNCN 16.01 / 17819) everted maxillae, ventral view." httpUri="https://zenodo.org/record/839228/files/figure.png" pageId="16" pageNumber="521">Fig. 7</figureCitation>
|
||
G): MI = 1+1; MII = 7+6; MIII = 6+0; MIV = 14+11; MV = 1+1. Tube typical of genus with inner secreted layer and outer layer of foreign particles consisting mainly of algae and vegetal material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36EE4A8D62FFFEAFF5FF808FAFDF830" box="[199,1373,1950,1975]" pageId="16" pageNumber="521" type="etymology">
|
||
<paragraph id="8BCBB723D62FFFEAFF5FF808FAFDF830" blockId="16.[151,1437,402,1975]" box="[199,1373,1950,1975]" pageId="16" pageNumber="521">
|
||
<emphasis id="B9006B31D62FFFEAFF5FF808FEEEF830" bold="true" box="[199,334,1950,1975]" pageId="16" pageNumber="521">Etymology.</emphasis>
|
||
It gives us great pleasure to name the new species in honour of Mrs. María Jesús Rodríguez.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0BE7ABD62EFFEBFF0FF938FB5FF8B9" httpUri="https://zenodo.org/record/839234/files/figure.png" pageId="17" pageNumber="522" targetBox="[151,1435,193,1689]" targetPageId="17">
|
||
<paragraph id="8BCBB723D62EFFEBFF0FF938FB5FF8B9" blockId="17.[151,1436,1710,1854]" pageId="17" pageNumber="522">
|
||
<emphasis id="B9006B31D62EFFEBFF0FF938FE81F944" bold="true" box="[151,289,1710,1732]" pageId="17" pageNumber="522">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C74CCA0D62EFFEBFEB0F939FE75F943" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[296,469,1711,1732]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="522" phylum="Annelida" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B9006B31D62EFFEBFEB0F939FE75F943" box="[296,469,1711,1732]" italics="true" pageId="17" pageNumber="522">Diopatra mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62EFFEBFE44F938FD8EF944" bold="true" box="[476,558,1710,1731]" pageId="17" pageNumber="522">
|
||
<taxonomicNameLabel id="A233D64AD62EFFEBFE44F938FD8EF944" box="[476,558,1710,1731]" pageId="17" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (MNCN 16.01/17819), scanning electron micrographs. A, Sensory buds of antenna; B, parapodium of chaetiger 4, posterior view; C, same magnified, note lower postchaetal lobe folded onto parapodium; D, parapodium of chaetiger 20, anterior view; E, parapodium of chaetiger 42, anterior view; F, pectinate chaetae. Abbreviations: a, aciculae, b, branchia, dc = dorsal cirrus, = limbate chaeta, lPCL = lower postchaetal lobe; p = pectinate chaeta, PrCL = prechaetal lobe; SAHs = subacicular hooks; uPCL = upper postchaetal lobe; vc = ventral cirrus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36EE4A8D62EFFE8FF5FF8FDFBC3FDBA" lastPageId="18" lastPageNumber="523" pageId="17" pageNumber="522" type="discussion">
|
||
<paragraph id="8BCBB723D62EFFE8FF5FF8FDFBC3FDBA" blockId="17.[151,1436,1899,2032]" lastBlockId="18.[151,1436,151,608]" lastPageId="18" lastPageNumber="523" pageId="17" pageNumber="522">
|
||
<emphasis id="B9006B31D62EFFEBFF5FF8FDFE9BF803" bold="true" box="[199,315,1899,1924]" pageId="17" pageNumber="522">Remarks.</emphasis>
|
||
Postchaetal lobes are well developed in the anterior parapodia of onuphids and occur in most cases singly in a median to dorsal or upper position. However, in some species of
|
||
<taxonomicName id="4C74CCA0D62EFFEBFC66F806FBC5F820" authorityName="Audouin & Milne Edwards" authorityYear="1833" box="[1022,1125,1936,1959]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="522" phylum="Annelida" rank="genus">
|
||
<emphasis id="B9006B31D62EFFEBFC66F806FBC5F820" box="[1022,1125,1936,1959]" italics="true" pageId="17" pageNumber="522">Diopatra</emphasis>
|
||
</taxonomicName>
|
||
a ventral knob may extend from the lower part of the postchaetal lobe to the prechaetal lobe, forming a ventral protrusion (or short lower postchaetal lobe) as in
|
||
<taxonomicName id="4C74CCA0D62EFFEBFE04F84EFDB0F868" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[412,528,2008,2031]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="522" phylum="Annelida" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B9006B31D62EFFEBFE04F84EFDB0F868" box="[412,528,2008,2031]" italics="true" pageId="17" pageNumber="522">D. mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62EFFEBFD8FF841FDD2F877" bold="true" box="[535,626,2007,2032]" pageId="17" pageNumber="522">
|
||
<taxonomicNameLabel id="A233D64AD62EFFEBFD8FF841FDD2F877" box="[535,626,2007,2032]" pageId="17" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="134FABA6D62EFFEBFD1BF841FD72F877" box="[643,722,2007,2032]" captionStart="FIGURE 11" captionStartId="18.[151,250,1426,1448]" captionTargetBox="[166,1415,674,1389]" captionTargetId="figure@18.[152,1435,663,1404]" captionTargetPageId="18" captionText="FIGURE 11. Diopatra mariae sp. nov., holotype (MNCN 16.01 / 17819), line drawings. A, Parapodium from chaetiger 2, anterior view; B, median robust pseudompound hook from chaetiger 2; C, upper slender pseudocompound hook from same; D, lower very slender pseudocompound hook from same; E, pectinate chaeta from chaetiger 42; F, subacicular hook from same. Abbreviations: lPCL = lower postchaetal lobe; uPCL = upper postchaetal lobe." httpUri="https://zenodo.org/record/839236/files/figure.png" pageId="17" pageNumber="522">Fig. 11</figureCitation>
|
||
A) or when longer and subulate, form a lower postchaetal lobe that is only slightly shorter than the upper postchaetal lobe (see
|
||
<taxonomicName id="4C74CCA0D62DFFE8FCFCFF0FFC71FF37" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[868,977,152,176]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="mellea" status="sp. nov.">
|
||
<emphasis id="B9006B31D62DFFE8FCFCFF0FFC71FF37" box="[868,977,152,176]" italics="true" pageId="18" pageNumber="523">D. mellea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62DFFE8FC40FF01FB92FF37" bold="true" box="[984,1074,151,176]" pageId="18" pageNumber="523">
|
||
<taxonomicNameLabel id="A233D64AD62DFFE8FC40FF01FB92FF37" box="[984,1074,151,176]" pageId="18" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
described below (
|
||
<figureCitation id="134FABA6D62DFFE8FB67FF01FAF1FF37" box="[1279,1361,151,176]" captionStart="FIGURE 15" captionStartId="22.[151,250,1307,1329]" captionTargetBox="[161,1421,195,1280]" captionTargetId="figure@22.[150,1436,193,1286]" captionTargetPageId="22" captionText="FIGURE 15. Diopatra mellea sp. nov., paratype (AM W. 94216), line drawings. A, Left parapodium from chaetiger 2, anterior view, chaetae omitted; B, right parapodium from chaetiger 2, chaetae omitted; C, median robust pseudocompound hook from chaetiger 4; D, upper slender pseudocompound hook from same; E, lower very slender pseudocompound hook from same; F, pectinate chaeta from chaetiger 21; G, pectinate chaeta from same; H, mandibles; I, maxillae. Abbreviations: lPCL = lower postchaetal lobe; uPCL = upper postchaetal lobe." httpUri="https://zenodo.org/record/839244/files/figure.png" pageId="18" pageNumber="523">Fig. 15</figureCitation>
|
||
B); for detailed discussion see
|
||
<bibRefCitation id="EFE5CAD2D62DFFE8FE32FF2BFDF2FF53" author="Paxton" box="[426,594,188,212]" pageId="18" pageNumber="523" refString="Paxton, H. (1998) The Diopatra chiliensis confusion-redescription of D. chiliensis (Polychaeta, Onuphidae) and implicated species. Zoologica Scripta, 27 (1), 31 - 48." type="journal article" year="1998">Paxton (1998)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C74CCA0D62DFFE8FDF8FF2BFC81FF53" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[608,801,189,212]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B9006B31D62DFFE8FDF8FF2BFC81FF53" box="[608,801,189,212]" italics="true" pageId="18" pageNumber="523">Diopatra mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62DFFE8FCB5FF2AFC2BFF52" bold="true" box="[813,907,188,213]" pageId="18" pageNumber="523">
|
||
<taxonomicNameLabel id="A233D64AD62DFFE8FCB5FF2AFC2BFF52" box="[813,907,188,213]" pageId="18" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C74CCA0D62DFFE8FC52FF2BFB9BFF53" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[970,1083,188,212]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="mellea" status="sp. nov.">
|
||
<emphasis id="B9006B31D62DFFE8FC52FF2BFB9BFF53" box="[970,1083,188,212]" italics="true" pageId="18" pageNumber="523">D. mellea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62DFFE8FBDEFF2AFB04FF52" bold="true" box="[1094,1188,188,213]" pageId="18" pageNumber="523">
|
||
<taxonomicNameLabel id="A233D64AD62DFFE8FBDEFF2AFB04FF52" box="[1094,1188,188,213]" pageId="18" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
bring the number of
|
||
<taxonomicName id="4C74CCA0D62DFFE8FF0FFF76FF5EFF70" authorityName="Audouin & Milne Edwards" authorityYear="1833" box="[151,254,224,247]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="genus">
|
||
<emphasis id="B9006B31D62DFFE8FF0FFF76FF5EFF70" box="[151,254,224,247]" italics="true" pageId="18" pageNumber="523">Diopatra</emphasis>
|
||
</taxonomicName>
|
||
species with double postchaetal lobes to eight (
|
||
<tableCitation id="C6F68298D62DFFE8FC89FF49FCC9FF7F" box="[785,873,223,248]" captionStart="TABLE 2" captionStartId="25.[125,198,565,586]" captionText="TABLE 2. List Ɵf Diopatra speƇies with đƟuble pƟstƇhaetal lƟbes. PC, pseuđƟƇƟmpƟunđ hƟƟk." pageId="18" pageNumber="523">Table 2</tableCitation>
|
||
). In four of these (
|
||
<emphasis id="B9006B31D62DFFE8FBA1FF77FEB9FE9B" italics="true" pageId="18" pageNumber="523">
|
||
<taxonomicName id="4C74CCA0D62DFFE8FBA1FF77FB7DFF7F" authorityName="Fauchald et al." authorityYear="2012" box="[1081,1245,224,248]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="biscayensis">D. biscayensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C74CCA0D62DFFE8FB70FF77FA38FF7F" authorityName="Paxton & Bailey-Brock" authorityYear="1986" box="[1256,1432,224,248]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="dexiognatha">D. dexiognatha</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C74CCA0D62DFFE8FF0FFE93FEB9FE9B" authorityName="Paxton" authorityYear="1998" box="[151,281,260,284]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="kristiani">D. kristiani</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C74CCA0D62DFFE8FECDFE93FE72FE9B" authorityName="Izuka" authorityYear="1907" box="[341,466,260,284]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="sugokai">
|
||
<emphasis id="B9006B31D62DFFE8FECDFE93FE72FE9B" box="[341,466,260,284]" italics="true" pageId="18" pageNumber="523">D. sugokai</emphasis>
|
||
</taxonomicName>
|
||
) the upper and lower postchaetal lobes are both subulate, while in
|
||
<emphasis id="B9006B31D62DFFE8FB79FE93FF77FEC7" italics="true" pageId="18" pageNumber="523">
|
||
<taxonomicName id="4C74CCA0D62DFFE8FB79FE93FAD2FE9B" box="[1249,1394,260,284]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="chiliensis">D. chiliensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C74CCA0D62DFFE8FA19FE93FF77FEC7" authorityName="Paxton" authorityYear="1998" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="gesae">D. gesae</taxonomicName>
|
||
</emphasis>
|
||
and the two new species the lower postchaetal lobes range from knob-like to subulate. The former two species can easily be distinguished from the latter two by their unique features:
|
||
<taxonomicName id="4C74CCA0D62DFFE8FB8AFEDBFB3EFEE3" box="[1042,1182,332,356]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="chiliensis">
|
||
<emphasis id="B9006B31D62DFFE8FB8AFEDBFB3EFEE3" box="[1042,1182,332,356]" italics="true" pageId="18" pageNumber="523">D. chiliensis</emphasis>
|
||
</taxonomicName>
|
||
has extremely oblique pectinate chaetae and
|
||
<taxonomicName id="4C74CCA0D62DFFE8FE17FEE7FE52FE0F" authorityName="Paxton" authorityYear="1998" box="[399,498,369,392]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="gesae">
|
||
<emphasis id="B9006B31D62DFFE8FE17FEE7FE52FE0F" box="[399,498,369,392]" italics="true" pageId="18" pageNumber="523">D. gesae</emphasis>
|
||
</taxonomicName>
|
||
has mandibles with a ventral bulge. Furthermore, the anterior hooks are uni- to bi- and even tridentate while those of the two new species are bidentate only; other differences are detailed in
|
||
<tableCitation id="C6F68298D62DFFE8FADAFE02FA38FE2B" box="[1346,1432,404,429]" captionStart="TABLE 2" captionStartId="25.[125,198,565,586]" captionText="TABLE 2. List Ɵf Diopatra speƇies with đƟuble pƟstƇhaetal lƟbes. PC, pseuđƟƇƟmpƟunđ hƟƟk." pageId="18" pageNumber="523">Table 2</tableCitation>
|
||
. The two new species from the
|
||
<collectingRegion id="49B079C1D62DFFE8FE75FE21FDEFFE57" box="[493,591,439,464]" country="Spain" name="Canarias" pageId="18" pageNumber="523">Canaries</collectingRegion>
|
||
can be distinguished from each other in that in
|
||
<taxonomicName id="4C74CCA0D62DFFE8FBF9FE2FFB72FE48" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[1121,1234,440,464]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B9006B31D62DFFE8FBF9FE2FFB72FE48" box="[1121,1234,440,464]" italics="true" pageId="18" pageNumber="523">D. mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62DFFE8FB41FE21FA92FE57" bold="true" box="[1241,1330,439,464]" pageId="18" pageNumber="523">
|
||
<taxonomicNameLabel id="A233D64AD62DFFE8FB41FE21FA92FE57" box="[1241,1330,439,464]" pageId="18" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
all lower postchaetal lobes are knob-like, ceratophoral rings number 7–9, pectinate chaetae have 18–22 teeth and limbate chaetae are coarsely serrated, while
|
||
<taxonomicName id="4C74CCA0D62DFFE8FDA8FD97FD3EFD9F" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[560,670,512,536]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="mellea" status="sp. nov.">
|
||
<emphasis id="B9006B31D62DFFE8FDA8FD97FD3EFD9F" box="[560,670,512,536]" italics="true" pageId="18" pageNumber="523">D. mellea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62DFFE8FD3DFE69FCA0FD9F" bold="true" box="[677,768,511,536]" pageId="18" pageNumber="523">
|
||
<taxonomicNameLabel id="A233D64AD62DFFE8FD3DFE69FCA0FD9F" box="[677,768,511,536]" pageId="18" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has knob-like and subulate lower postchaetal lobes, 12–13 ceratophoral rings, 4–11 teeth on pectinate chaetae and finely serrated limbate chaetae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36EE4A8D62DFFE8FF5FFDD1FB3BFDE7" box="[199,1179,583,608]" pageId="18" pageNumber="523" type="distribution">
|
||
<paragraph id="8BCBB723D62DFFE8FF5FFDD1FB3BFDE7" blockId="18.[151,1436,151,608]" box="[199,1179,583,608]" pageId="18" pageNumber="523">
|
||
<emphasis id="B9006B31D62DFFE8FF5FFDD1FEC0FDE7" bold="true" box="[199,352,583,608]" pageId="18" pageNumber="523">Distribution.</emphasis>
|
||
<taxonomicName id="4C74CCA0D62DFFE8FEFFFDDEFD84FDD8" authority="Paxton & Arias, 2017" authorityName="Paxton & Arias" authorityYear="2017" box="[359,548,584,607]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="523" phylum="Annelida" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B9006B31D62DFFE8FEFFFDDEFD84FDD8" box="[359,548,584,607]" italics="true" pageId="18" pageNumber="523">Diopatra mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9006B31D62DFFE8FDB3FDD1FD25FDE7" bold="true" box="[555,645,583,608]" pageId="18" pageNumber="523">
|
||
<taxonomicNameLabel id="A233D64AD62DFFE8FDB3FDD1FD25FDE7" box="[555,645,583,608]" pageId="18" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is only known from Lanzarote,
|
||
<collectingRegion id="49B079C1D62DFFE8FC75FDDEFB37FDE7" box="[1005,1175,583,608]" country="Spain" name="Canarias" pageId="18" pageNumber="523">Canary Islands</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |