338 lines
29 KiB
XML
338 lines
29 KiB
XML
<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683667246730" checkinUser="felipe" docAuthor="., D] D, Qgudnrwr, Csuzdi, Cs. & Blanchart, E." docDate="2011" docId="03DC87D4FF8FFF97FDD0FDC9FD504656" docLanguage="en" docName="AfrInvertebr.52.2.285-294.pdf" docOrigin="African Invertebrates 52 (2)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Kynotus giganteus . & D]D & Qgudnrwr & Csuzdi & Blanchart 2011, sp. n." docType="treatment" docVersion="1" lastPageNumber="290" masterDocId="FFE5FFACFF8BFF91FFD5FF85FF8F434E" masterDocTitle="New and little known giant earthworms from Madagascar (Oligochaeta: Kynotidae)" masterLastPageNumber="294" masterPageNumber="285" pageNumber="288" updateTime="1683667307928" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New and little known giant earthworms from Madagascar (Oligochaeta: Kynotidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>D] D</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Qgudnrwr</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Csuzdi, Cs.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Blanchart, E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2011-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>52</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>285</mods:start>
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<mods:end>294</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03DC87D4FF8FFF97FDD0FDC9FD504656" httpUri="http://treatment.plazi.org/id/03DC87D4FF8FFF97FDD0FDC9FD504656" lastPageId="6" lastPageNumber="290" pageId="4" pageNumber="288">
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<subSubSection box="[517,806,587,614]" pageId="4" pageNumber="288" type="nomenclature">
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<paragraph blockId="4.[181,1142,587,1164]" box="[517,806,587,614]" pageId="4" pageNumber="288">
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<heading box="[517,806,587,614]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="288" reason="2">
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<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[517,739,587,614]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="288" phylum="Annelida" rank="species" species="giganteus" status="sp. n.">
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<emphasis bold="true" box="[517,739,587,614]" pageId="4" pageNumber="288">Kynotus giganteus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[746,806,588,614]" pageId="4" pageNumber="288" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="290" pageId="4" pageNumber="288" type="description">
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<paragraph blockId="4.[181,1142,587,1164]" box="[611,711,627,653]" pageId="4" pageNumber="288">
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<figureCitation box="[611,711,627,653]" captionStart-0="Fig" captionStart-1="Figs 4, 5" captionStartId-0="4.[371,404,1654,1675]" captionStartId-1="5.[144,183,1630,1651]" captionTargetBox-0="[191,1136,1201,1632]" captionTargetBox-1="[151,1096,574,1612]" captionTargetId-0="figure-316@4.[191,1136,1201,1632]" captionTargetId-1="figure-156@5.[151,1096,955,1612]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 3. K. giganteus sp. n., body view and enlarged anterior end." captionText-1="Figs 4, 5. K. giganteus sp. n.: (4) spermathecae; (5) penial seta: (A) whole seta, (B) ornamentation under the tip, (C) tip of the seta." pageId="4" pageNumber="288">Figs 3–5</figureCitation>
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</paragraph>
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<paragraph blockId="4.[181,1142,587,1164]" pageId="4" pageNumber="288">
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Etymology: From Latin
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<taxonomicName authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[468,577,670,694]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="288" phylum="Annelida" rank="species" species="giganteus">
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<emphasis box="[468,577,670,694]" italics="true" pageId="4" pageNumber="288">giganteus</emphasis>
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</taxonomicName>
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(giant), in reference to the size of the captured worms.
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</paragraph>
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<paragraph blockId="4.[181,1142,587,1164]" pageId="4" pageNumber="288">
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Diagnosis: Length alive
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<quantity box="[478,666,739,766]" metricMagnitude="0" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="4" pageNumber="288" unit="mm" value="1375.0" valueMax="1400.0" valueMin="1350.0">1350–1400 mm</quantity>
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, diameter
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<quantity box="[797,930,739,766]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="288" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20–25 mm</quantity>
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; preserved length
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<quantity box="[181,362,771,798]" metricMagnitude="0" metricUnit="m" metricValue="1.46" metricValueMax="1.8399999999999999" metricValueMin="1.08" pageId="4" pageNumber="288" unit="mm" value="1460.0" valueMax="1840.0" valueMin="1080.0">1080–1840 mm</quantity>
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diameter
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<quantity box="[479,606,771,798]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="288" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
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. Colour slightly reddish at the head, pale after the clitellum. Segments 1–2 simple, 3–16 double ringed. Setae
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<emphasis box="[890,918,806,830]" italics="true" pageId="4" pageNumber="288">ab</emphasis>
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and
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<emphasis box="[972,998,806,830]" italics="true" pageId="4" pageNumber="288">cd</emphasis>
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appear from segment 17. Male pore on 16, on the second ringlet.
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<taxonomicName box="[764,870,835,861]" class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aplousobranchia" pageId="4" pageNumber="293" phylum="Chordata" rank="genus">Clitellum</taxonomicName>
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indistinct on 22–47, 48. Setae closely paired, both
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<emphasis box="[474,502,870,894]" italics="true" pageId="4" pageNumber="288">ab</emphasis>
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and
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<emphasis box="[556,582,870,894]" italics="true" pageId="4" pageNumber="288">cd</emphasis>
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lateral. Spermathecal pores from above
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<emphasis box="[1033,1061,870,894]" italics="true" pageId="4" pageNumber="288">ab</emphasis>
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to Md, 9–14 on each side in 13/14, 14/15, 15/16. Spermathecae intramuscular. Genital setal glands in 14, 15, 16. Genital setae spoon-shaped, slightly tetrahedral. Length
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<emphasis box="[1038,1064,934,958]" italics="true" pageId="4" pageNumber="288">ca</emphasis>
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<quantity box="[1071,1137,931,958]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="288" unit="mm" value="6.0">6 mm</quantity>
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, diameter 0.04 mm, ornamentation dense serrations.
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</paragraph>
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<paragraph blockId="4.[181,1142,587,1164]" box="[181,321,1004,1030]" pageId="4" pageNumber="288">Description:</paragraph>
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<paragraph blockId="4.[181,1142,587,1164]" lastBlockId="5.[144,1104,203,550]" lastPageId="5" lastPageNumber="289" pageId="4" pageNumber="288">
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||
<emphasis box="[181,284,1043,1067]" italics="true" pageId="4" pageNumber="288">
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<typeStatus box="[181,284,1043,1067]" pageId="4" pageNumber="288">Holotype</typeStatus>
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</emphasis>
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: Preserved length
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<quantity box="[499,610,1041,1068]" metricMagnitude="0" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="288" unit="mm" value="1450.0">1450 mm</quantity>
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, diameter after clitellum
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<quantity box="[900,981,1041,1068]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="288" unit="mm" value="15.0">15 mm</quantity>
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(
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<figureCitation box="[998,1068,1041,1067]" captionStart="Fig" captionStartId="4.[371,404,1654,1675]" captionTargetBox="[191,1136,1201,1632]" captionTargetId="figure-316@4.[191,1136,1201,1632]" captionTargetPageId="4" captionText="Fig. 3. K. giganteus sp. n., body view and enlarged anterior end." pageId="4" pageNumber="288">Fig. 3</figureCitation>
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).
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<emphasis italics="true" pageId="4" pageNumber="288">Segment number</emphasis>
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: 1330.
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<emphasis box="[411,525,1075,1099]" italics="true" pageId="4" pageNumber="288">
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<typeStatus box="[411,525,1075,1099]" pageId="4" pageNumber="288">Paratypes</typeStatus>
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</emphasis>
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:
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<quantity box="[539,716,1073,1100]" metricMagnitude="0" metricUnit="m" metricValue="1.46" metricValueMax="1.8399999999999999" metricValueMin="1.08" pageId="4" pageNumber="288" unit="mm" value="1460.0" valueMax="1840.0" valueMin="1080.0">1080–1840 mm</quantity>
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in length
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||
<quantity box="[826,948,1073,1100]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="288" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
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in diameter.
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<emphasis italics="true" pageId="4" pageNumber="288">Segment number</emphasis>
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: 1300–1350. Weight of preserved worms ranges between
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<quantity box="[969,1090,1105,1132]" metricMagnitude="-1" metricUnit="kg" metricValue="1.895" metricValueMax="1.95" metricValueMin="1.8399999999999999" pageId="4" pageNumber="288" unit="g" value="189.5" valueMax="195.0" valueMin="184.0">184–195g</quantity>
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.
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<emphasis italics="true" pageId="4" pageNumber="288">Colour</emphasis>
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||
: Alive slightly reddish at head, whitish after clitellum; preserved greyish.
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<emphasis box="[1073,1134,1139,1163]" italics="true" pageId="4" pageNumber="288">Head</emphasis>
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: Prolobous.
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<emphasis box="[271,377,206,230]" italics="true" pageId="5" pageNumber="289">Segments</emphasis>
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||
: 1–2 simple, 3–16 clearly double ringed.
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<emphasis box="[844,990,206,230]" italics="true" pageId="5" pageNumber="289">Dorsal pores</emphasis>
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: Lacking.
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<emphasis box="[144,205,238,262]" italics="true" pageId="5" pageNumber="289">Setae</emphasis>
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: Small, both
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<emphasis box="[367,395,238,262]" italics="true" pageId="5" pageNumber="289">ab</emphasis>
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and
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<emphasis box="[458,484,238,262]" italics="true" pageId="5" pageNumber="289">bc</emphasis>
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became observable on segment 17. All setae lateral, setal ratio
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<emphasis box="[268,296,270,294]" italics="true" pageId="5" pageNumber="289">aa</emphasis>
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:
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<emphasis box="[304,332,270,294]" italics="true" pageId="5" pageNumber="289">ab</emphasis>
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:
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<emphasis box="[340,366,270,294]" italics="true" pageId="5" pageNumber="289">bc</emphasis>
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:
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<emphasis box="[374,400,270,294]" italics="true" pageId="5" pageNumber="289">cd</emphasis>
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:
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<emphasis box="[408,436,270,294]" italics="true" pageId="5" pageNumber="289">dd</emphasis>
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= 100:1:10:1:90.
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<emphasis box="[649,842,270,294]" italics="true" pageId="5" pageNumber="289">Nephridial pores</emphasis>
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: Begin on segment 2, somewhat dorsal to setal line
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<emphasis box="[476,504,302,326]" italics="true" pageId="5" pageNumber="289">ab</emphasis>
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.
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</paragraph>
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<caption box="[371,953,1654,1675]" pageId="4" pageNumber="288" startId="4.[371,404,1654,1675]" targetBox="[191,1136,1201,1632]" targetPageId="4" targetType="figure">
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<paragraph blockId="4.[371,953,1654,1675]" box="[371,953,1654,1675]" pageId="4" pageNumber="288">
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Fig. 3.
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<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[435,550,1656,1675]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="288" phylum="Annelida" rank="species" species="giganteus" status="sp. n.">
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<emphasis box="[435,550,1656,1675]" italics="true" pageId="4" pageNumber="288">K. giganteus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[555,606,1654,1675]" pageId="4" pageNumber="288" rank="species">sp. n.</taxonomicNameLabel>
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, body view and enlarged anterior end.
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</paragraph>
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</caption>
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<paragraph blockId="5.[144,1104,203,550]" pageId="5" pageNumber="289">
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<taxonomicName authority=": Only" authorityName="Only" box="[144,322,331,358]" class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aplousobranchia" pageId="5" pageNumber="289" phylum="Chordata" rank="genus">
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<emphasis box="[144,248,334,358]" italics="true" pageId="5" pageNumber="289">Clitellum</emphasis>
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: Only
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</taxonomicName>
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indistinctly developed on segments 22–47, 48.
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<emphasis box="[886,1004,334,358]" italics="true" pageId="5" pageNumber="289">Male pore</emphasis>
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: Ventral on 16, large oval slit on second ringlet.
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<emphasis box="[610,765,366,390]" italics="true" pageId="5" pageNumber="289">Female pores</emphasis>
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: Small slits on 14, in line of male pores.
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<emphasis box="[284,508,398,422]" italics="true" pageId="5" pageNumber="289">Spermathecal pores</emphasis>
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: In intersegmental furrows 13/14, 14/15, 15/16, numerous, 9–14 on each side, begin around the line of nephridial pores and scattered until mid-dorsal line. Spermathecal lines sometimes interrupted and continue not in intersegmental furrow but in furrows of two ringlets.
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<emphasis box="[749,963,494,518]" italics="true" pageId="5" pageNumber="289">Genital setal pores</emphasis>
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: Segmental, irregularly occur on lateral sides of segments 14, 15 and 16.
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</paragraph>
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<caption pageId="5" pageNumber="289" startId="5.[144,183,1630,1651]" targetBox="[151,1096,574,1612]" targetPageId="5" targetType="figure">
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<paragraph blockId="5.[144,1104,1630,1675]" pageId="5" pageNumber="289">
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Figs 4, 5.
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<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[236,351,1632,1651]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="289" phylum="Annelida" rank="species" species="giganteus" status="sp. n.">
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<emphasis box="[236,351,1632,1651]" italics="true" pageId="5" pageNumber="289">K. giganteus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[358,410,1630,1651]" pageId="5" pageNumber="289" rank="species">sp. n.</taxonomicNameLabel>
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: (4) spermathecae; (5) penial seta: (A) whole seta, (B) ornamentation under the tip, (C) tip of the seta.
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</paragraph>
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</caption>
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<paragraph blockId="6.[181,1142,203,797]" pageId="6" pageNumber="290">
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Internal characters:
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<emphasis box="[410,609,206,230]" italics="true" pageId="6" pageNumber="290">Muscular gizzard</emphasis>
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: In 5, well-developed, cylindrical.
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<taxonomicName authorityName="Perry" authorityYear="1810" box="[1019,1081,206,230]" class="Gastropoda" family="Cymatiidae" genus="Septa" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="290" phylum="Mollusca" rank="genus">
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<emphasis box="[1019,1081,206,230]" italics="true" pageId="6" pageNumber="290">Septa</emphasis>
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</taxonomicName>
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: 5/6, 12/13, 13/14 thickened, 6/7–11/12 highly strengthened.
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<emphasis box="[815,1027,238,262]" italics="true" pageId="6" pageNumber="290">Calciferous glands</emphasis>
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: Lacking, even lamellar thickening of oesophagus missing.
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<emphasis box="[730,951,270,294]" italics="true" pageId="6" pageNumber="290">Dorsal blood vessel</emphasis>
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: Simple.
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<emphasis box="[1058,1134,270,294]" italics="true" pageId="6" pageNumber="290">Hearts</emphasis>
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: In 6–11, large moniliform, especially last two pairs.
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<emphasis box="[778,845,303,327]" italics="true" pageId="6" pageNumber="290">Testes</emphasis>
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and
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<emphasis box="[902,1059,303,327]" italics="true" pageId="6" pageNumber="290">sperm funnels</emphasis>
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: In 10, 11 enclosed in peritoneal sack.
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<emphasis box="[533,717,335,359]" italics="true" pageId="6" pageNumber="290">Seminal vesicles</emphasis>
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: Lacking.
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||
<emphasis box="[838,927,335,359]" italics="true" pageId="6" pageNumber="290">Ovaries</emphasis>
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: In 13.
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<emphasis italics="true" pageId="6" pageNumber="290">Copulatory chamber</emphasis>
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: Large, oval, occupies the ventral place of segments 15–17. Each copulatory chamber bears irregular prostate-like gland (pseudoprostate) bulging up to segment 13; gland’s position highly variable, sometimes dorsal but sometimes running towards tail.
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<emphasis box="[230,389,464,488]" italics="true" pageId="6" pageNumber="290">Spermathecae</emphasis>
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: Small, numerous, completely embedded in body wall, only shining sperm can be seen from inside. Cutting body wall at intersegmental furrow 13/14, several small oval spermathecae can be seen that open to surface with small duct (
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<figureCitation box="[1060,1128,526,552]" captionStart="Figs 4, 5" captionStartId="5.[144,183,1630,1651]" captionTargetBox="[151,1096,574,1612]" captionTargetId="figure-156@5.[151,1096,955,1612]" captionTargetPageId="5" captionText="Figs 4, 5. K. giganteus sp. n.: (4) spermathecae; (5) penial seta: (A) whole seta, (B) ornamentation under the tip, (C) tip of the seta." pageId="6" pageNumber="290">Fig. 4</figureCitation>
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).
|
||
<emphasis box="[182,398,561,585]" italics="true" pageId="6" pageNumber="290">Genital setal organ</emphasis>
|
||
: Three pairs in segments 14, 15, 16. They consist of large, slightly spiral gland similar in shape to pseudoprostates and genital setal sac containing several genital setae.
|
||
<emphasis box="[404,554,625,649]" italics="true" pageId="6" pageNumber="290">Genital setae</emphasis>
|
||
: Spoon-shaped, slightly tetrahedral,
|
||
<emphasis box="[974,1000,625,649]" italics="true" pageId="6" pageNumber="290">ca</emphasis>
|
||
<quantity box="[1009,1076,623,649]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="290" unit="mm" value="6.0">6 mm</quantity>
|
||
long, 0.04mm in diameter and provided with characteristic serrated ornamentation (
|
||
<figureCitation box="[1059,1128,655,681]" captionStart="Figs 4, 5" captionStartId="5.[144,183,1630,1651]" captionTargetBox="[151,1096,574,1612]" captionTargetId="figure-156@5.[151,1096,955,1612]" captionTargetPageId="5" captionText="Figs 4, 5. K. giganteus sp. n.: (4) spermathecae; (5) penial seta: (A) whole seta, (B) ornamentation under the tip, (C) tip of the seta." pageId="6" pageNumber="290">Fig. 5</figureCitation>
|
||
).
|
||
<typeStatus box="[182,270,694,715]" pageId="6" pageNumber="290">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry box="[282,447,694,715]" name="Madagascar" pageId="6" pageNumber="290">MADAGASCAR</collectingCountry>
|
||
: Atsimo-Atsinanana region, Farafangana district,
|
||
<collectingRegion box="[916,1032,694,715]" country="Madagascar" name="Fianarantsoa" pageId="6" pageNumber="290">Fianarantsoa</collectingRegion>
|
||
town, near
|
||
<emphasis bold="true" box="[181,1141,716,740]" pageId="6" pageNumber="290">YLOODJH RI $QND]RPDIDLWV\ (22°38‵27.8̎6:47°49‵4Ɵ.8̎(), 4ƽ P, 23.L.2ƟlƟ, 0. 5D]D¿QGUDNRWR (+1+0</emphasis>
|
||
AF/5513).
|
||
</paragraph>
|
||
<paragraph blockId="6.[181,1142,203,797]" box="[181,885,776,797]" pageId="6" pageNumber="290">
|
||
<materialsCitation box="[181,882,776,797]" collectionCode="HNHM" pageId="6" pageNumber="290" specimenCount="4" typeStatus="paratype">
|
||
<typeStatus box="[181,273,776,797]" pageId="6" pageNumber="290">Paratypes</typeStatus>
|
||
: Same data as holotype (
|
||
<collectionCode box="[497,568,776,797]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="290">HNHM</collectionCode>
|
||
AF/5514
|
||
<specimenCount box="[658,705,776,797]" pageId="6" pageNumber="290" type="generic">1 ex.</specimenCount>
|
||
, ZMUA-030
|
||
<specimenCount box="[827,875,776,797]" pageId="6" pageNumber="290" type="generic">3 ex.</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="6.[181,1142,815,874]" pageId="6" pageNumber="290">
|
||
Habitat: The vegetation in the locality is formed by savannah and secondary forest with
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1891" box="[238,553,850,874]" class="Liliopsida" family="Strelitziaceae" genus="Ravenala" kingdom="Plantae" order="Zingiberales" pageId="6" pageNumber="290" phylum="Tracheophyta" rank="species" species="madagascariensis">
|
||
<emphasis box="[238,553,850,874]" italics="true" pageId="6" pageNumber="290">Ravenala madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
very dominant.
|
||
</paragraph>
|
||
<paragraph blockId="6.[181,1142,891,1304]" pageId="6" pageNumber="290">
|
||
Remarks: The new species exceeds in size all known
|
||
<taxonomicName box="[788,976,891,917]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis box="[788,880,893,917]" italics="true" pageId="6" pageNumber="290">Kynotus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
It is similar to
|
||
<taxonomicName authority="Keller, 1887" authorityName="Keller" authorityYear="1887" box="[181,453,923,949]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="darwini">
|
||
<emphasis box="[181,303,925,949]" italics="true" pageId="6" pageNumber="290">K. darwini</emphasis>
|
||
<bibRefCitation author="KELLER, C." box="[312,453,923,949]" pageId="6" pageNumber="290" refId="ref4206" refString="KELLER, C. 1887. Reisebilder aus Ostafrika und Madagaskar. Leipzig: C. F. Winter Press." type="book" year="1887">Keller, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName authority="Michaelsen, 1891" authorityName="Michaelsen" authorityYear="1891" box="[496,944,923,949]" class="Liliopsida" family="Strelitziaceae" genus="Ravenala" kingdom="Plantae" order="Zingiberales" pageId="6" pageNumber="290" phylum="Tracheophyta" rank="species" species="madagascariensis">
|
||
<emphasis box="[496,732,925,949]" italics="true" pageId="6" pageNumber="290">K. madagascariensis</emphasis>
|
||
Michaelsen, 1891
|
||
</taxonomicName>
|
||
) but differs from the latter in its larger size, the position of the genital setae (
|
||
<quantity box="[879,984,955,981]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="6" pageNumber="290" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
|
||
<taxonomicName authorityName="Keller" authorityYear="1887" box="[993,1080,957,981]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="darwini">
|
||
<emphasis box="[993,1080,957,981]" italics="true" pageId="6" pageNumber="290">darwini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="6" pageNumber="290" unit="in" value="15.0" valueMax="16.0" valueMin="14.0">14– 16 in</quantity>
|
||
<taxonomicName authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[252,361,990,1014]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="giganteus">
|
||
<emphasis box="[252,361,990,1014]" italics="true" pageId="6" pageNumber="290">giganteus</emphasis>
|
||
</taxonomicName>
|
||
) and in the spermathecae (which are intracoelomic and sac-shaped in
|
||
<taxonomicName authorityName="Keller" authorityYear="1887" box="[214,337,1022,1046]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="darwini">
|
||
<emphasis box="[214,337,1022,1046]" italics="true" pageId="6" pageNumber="290">K. darwini</emphasis>
|
||
</taxonomicName>
|
||
, intramuscular and egg-shaped in
|
||
<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[741,887,1022,1046]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="giganteus" status="sp. n.">
|
||
<emphasis box="[741,887,1022,1046]" italics="true" pageId="6" pageNumber="290">K. giganteus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[897,962,1020,1046]" pageId="6" pageNumber="290" rank="species">sp. n.</taxonomicNameLabel>
|
||
). Regarding its spermathecal apparatus the new species is similar to
|
||
<taxonomicName authority="Michaelsen, 1931" authorityName="Michaelsen" authorityYear="1931" box="[795,1137,1052,1078]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="friderici">
|
||
<emphasis box="[795,922,1054,1078]" italics="true" pageId="6" pageNumber="290">K. friderici</emphasis>
|
||
Michaelsen, 1931
|
||
</taxonomicName>
|
||
; however, this species possesses only two spermathecal lines (14/15, 15/16) with 2–2 spermathecae on each side. A further difference can be found in the genital setae, which are claw-shaped in
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1931" box="[474,598,1151,1175]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="friderici">
|
||
<emphasis box="[474,598,1151,1175]" italics="true" pageId="6" pageNumber="290">K. friderici</emphasis>
|
||
</taxonomicName>
|
||
and spoon-shaped in
|
||
<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[842,984,1151,1175]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="giganteus" status="sp. n.">
|
||
<emphasis box="[842,984,1151,1175]" italics="true" pageId="6" pageNumber="290">K. giganteus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[991,1051,1149,1175]" pageId="6" pageNumber="290" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="6.[181,1142,891,1304]" pageId="6" pageNumber="290">
|
||
It is remarkable that this giant worm possesses very small spermathecae completely embedded in the body wall, similar to several giant Ecuadorian earthworms such as
|
||
<taxonomicName authority="Zicsi, 1990" authorityName="Zicsi" authorityYear="1990" box="[181,557,1245,1272]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="ischuros">
|
||
<emphasis box="[181,424,1248,1272]" italics="true" pageId="6" pageNumber="290">Martiodrilus ischuros</emphasis>
|
||
Zicsi, 1990
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Rosa, 1895)" baseAuthorityName="Rosa" baseAuthorityYear="1895" box="[566,947,1245,1272]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="crassus">
|
||
<emphasis box="[566,799,1248,1272]" italics="true" pageId="6" pageNumber="290">Martiodrilus crassus</emphasis>
|
||
(Rosa, 1895)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="James, 1990" authorityName="James" authorityYear="1990" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="olivaceous">
|
||
<emphasis italics="true" pageId="6" pageNumber="290">Martiodrilus olivaceous</emphasis>
|
||
James, 1990
|
||
</taxonomicName>
|
||
(each
|
||
<emphasis box="[521,547,1280,1304]" italics="true" pageId="6" pageNumber="290">ca</emphasis>
|
||
<quantity box="[555,662,1278,1304]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="290" unit="mm" value="1000.0">1000 mm</quantity>
|
||
long).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |