treatments-xml/data/03/DC/87/03DC87A39559F605F1D42CF6FDB4206C.xml
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<document id="7715A3973A3EFD727469CFFB782915D2" ID-DOI="10.5281/zenodo.185020" ID-GBIF-Dataset="30df05c7-bf86-489b-adca-cf70db449d58" ID-ISSN="1175-5326" ID-Zenodo-Dep="185020" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459949045561" checkinUser="plazi" docAuthor="Marin, Ivan" docDate="2009" docId="03DC87A39559F605F1D42CF6FDB4206C" docLanguage="en" docName="zt01971p049.pdf" docOrigin="Zootaxa 1971" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laomenes ceratophthalmus Borradaile 1915" docType="treatment" docVersion="7" lastPageNumber="39" masterDocId="FFE5FFDB9579F623F1432864FFF42206" masterDocTitle="Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity" masterLastPageNumber="49" masterPageNumber="1" pageNumber="33" updateTime="1698230776601" updateUser="plazi">
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<mods:title id="22FC002636F45D02F42B6C98D03F3D5E">Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity</mods:title>
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<mods:namePart id="CC00D1F718A2762A8C8F5567CF220BCF">Marin, Ivan</mods:namePart>
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<mods:date id="F09FF934D10935ACA6055CD54AAD5BED">2009</mods:date>
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<treatment id="03DC87A39559F605F1D42CF6FDB4206C" ID-DOI="http://doi.org/10.5281/zenodo.6213146" ID-GBIF-Taxon="119376430" ID-Zenodo-Dep="6213146" LSID="urn:lsid:plazi:treatment:03DC87A39559F605F1D42CF6FDB4206C" httpUri="http://treatment.plazi.org/id/03DC87A39559F605F1D42CF6FDB4206C" lastPageId="38" lastPageNumber="39" pageId="32" pageNumber="33">
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<paragraph id="8BCA36B59559F603F1D42CF6FD2426AA" blockId="32.[151,720,1170,1231]" box="[151,720,1170,1196]" pageId="32" pageNumber="33">
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<taxonomicName id="4C754D369559F603F1D42CF6FD2426AA" ID-CoL="ebead364-e4a5-4ae4-9774-e9654e54d3ea" authority="Borradaile, 1915" authorityName="Borradaile" authorityYear="1915" box="[151,720,1170,1196]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA79559F603F1D42CF6FD2426AA" bold="true" box="[151,720,1170,1196]" pageId="32" pageNumber="33">
<emphasis id="B901EAA79559F603F1D42CF6FE1126AA" bold="true" box="[151,485,1170,1196]" italics="true" pageId="32" pageNumber="33">Laomenes ceratophthalmus</emphasis>
(
<bibRefCitation id="EFE44B449559F603F0B72CF6FD3C26AA" author="Borradaile" box="[500,712,1170,1196]" pageId="32" pageNumber="48" refString="Borradaile, LA. (1915) Notes on Carides. Annals and Magazine of Natural History, 8, 205 - 213." type="journal article" year="1915">Borradaile, 1915</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCA36B59559F603F1D42CD1FE4C26C9" blockId="32.[151,720,1170,1231]" box="[151,440,1205,1231]" pageId="32" pageNumber="33">
(
<figureCitation id="134E2A309559F603F1DC2CD1FED926C9" box="[159,301,1205,1231]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStart-3="FIGURE 28" captionStart-4="FIGURE 29" captionStartId-0="33.[151,255,2013,2037]" captionStartId-1="34.[151,255,1944,1968]" captionStartId-2="35.[151,255,1336,1360]" captionStartId-3="36.[151,255,1940,1964]" captionStartId-4="37.[151,255,1931,1955]" captionTargetBox-0="[171,1409,1103,2012]" captionTargetBox-1="[196,1398,199,1911]" captionTargetBox-2="[174,1408,204,1289]" captionTargetBox-3="[194,1393,195,1890]" captionTargetBox-4="[186,1407,187,1884]" captionTargetId-0="figure@33.[151,1436,1082,2018]" captionTargetId-1="figure@34.[184,1408,194,1922]" captionTargetId-2="figure@35.[153,1438,194,1307]" captionTargetId-3="figure@36.[181,1405,180,1908]" captionTargetId-4="figure@37.[184,1408,180,1908]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionTargetPageId-3="36" captionTargetPageId-4="37" captionText-0="FIGURE 25. Laomenes ceratophthalmus (Borradaile, 1915). A, ovigerous female, B, male. Scale 1 mm." captionText-1="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." captionText-2="FIGURE 27. Laomenes ceratophthalmus (Borradaile, 1915), juvenile (pcl. 3 mm) (A, B), juvenile (pcl. 6 mm) (C, D), juvenile (pcl. 11 mm) (E). A E, distal part of carapace and rostrum. Scale 1 mm." captionText-3="FIGURE 28. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, mandible. C, same, distal margin of incisor process. D, maxillule. E, maxilla. F, maxilliped I. G, maxilliped II. H, maxilliped III. I, oral appendages and thoratic sternite III. Scale 1 mm; C, I, without scale." captionText-4="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri-0="https://zenodo.org/record/185043/files/figure.png" httpUri-1="https://zenodo.org/record/185044/files/figure.png" httpUri-2="https://zenodo.org/record/185045/files/figure.png" httpUri-3="https://zenodo.org/record/185046/files/figure.png" httpUri-4="https://zenodo.org/record/185047/files/figure.png" pageId="32" pageNumber="33">Figs. 2529</figureCitation>
,
<figureCitation id="134E2A309559F603F07A2CD1FEA326C9" box="[313,343,1205,1231]" captionStart="FIGURE 33" captionStartId="41.[151,255,1584,1608]" captionTargetBox="[151,1436,194,1553]" captionTargetId="figure@41.[151,1436,194,1555]" captionTargetPageId="41" captionText="FIGURE 33. Coloration of Laomenes ceratophthalmus (Borradaile, 1915) (A, C, E) and Laomenes nudirostris (Bruce, 1978) (B, D, F). A D, ovigerous females. E, F, males." httpUri="https://zenodo.org/record/185051/files/figure.png" pageId="32" pageNumber="33">33</figureCitation>
A, C, E)
</paragraph>
</subSubSection>
<subSubSection id="C36F653E9559F603F1D42D64FEE02771" pageId="32" pageNumber="33" type="reference_group">
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<treatmentCitationGroup id="AB65119B9559F603F1D42D64FEE02771" infered="true" pageId="32" pageNumber="33">
<taxonomicName id="4C754D369559F603F1D42D64FD302711" ID-CoL="4a11748e-6dd0-47e5-a909-6d50036fde82" authority="Borradaile 1915: 211" authorityName="Borradaile" authorityPageNumber="211" authorityYear="1915" box="[151,708,1280,1303]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA79559F603F1D42D64FE282711" box="[151,476,1280,1303]" italics="true" pageId="32" pageNumber="33">Periclimenes ceratophthalmus</emphasis>
<treatmentCitation id="0AD410A49559F603F0A02D64FD302711" author="Borradaile" box="[483,708,1280,1303]" page="211" pageId="32" pageNumber="33" year="1915">
<bibRefCitation id="EFE44B449559F603F0A02D64FD672711" author="Borradaile" box="[483,659,1280,1303]" pageId="32" pageNumber="48" refString="Borradaile, LA. (1915) Notes on Carides. Annals and Magazine of Natural History, 8, 205 - 213." type="journal article" year="1915">Borradaile 1915</bibRefCitation>
: 211
</treatmentCitation>
</taxonomicName>
[
<typeStatus id="54CE88179559F603F3922D64FCF62711" box="[721,770,1280,1303]" pageId="32" pageNumber="33">type</typeStatus>
locality:
<collectingCountry id="F36276259559F603F2252D64FC692711" box="[870,925,1280,1303]" name="Malta" pageId="32" pageNumber="33">Malé</collectingCountry>
Atoll,
<collectingCountry id="F36276259559F603F2AB2D64FB6D2711" box="[1000,1177,1280,1303]" name="Maldives" pageId="32" pageNumber="33">Maldive Islands</collectingCountry>
]; 1917: 324, 365, pl. 54; fig. 9. —
<treatmentCitation id="0AD410A49559F603F06D2D44FE042731" author="Holthuis" box="[302,496,1312,1335]" page="56" pageId="32" pageNumber="33" year="1952">
<bibRefCitation id="EFE44B449559F603F06D2D44FE3D2731" author="Holthuis" box="[302,457,1312,1335]" pageId="32" pageNumber="48" refString="Holthuis, L. B. (1952) The Decapoda of the Siboga Expedition. Part XI. The Palaemonidae collected by the Siboga and Snellius Expeditions with remarks on other species. II. Subfamily Pontoniinae. Siboga Expedition Monograph, 39 a, 10, 1 - 252." type="journal article" year="1952">Holthuis 1952</bibRefCitation>
: 56
</treatmentCitation>
, fig. 20. —
<treatmentCitation id="0AD410A49559F603F3332D44FCD62731" author="Bruce" box="[624,802,1312,1335]" page="192" pageId="32" pageNumber="33" year="1974">
<bibRefCitation id="EFE44B449559F603F3332D44FD1B2731" author="Bruce" box="[624,751,1312,1335]" pageId="32" pageNumber="48" refString="Bruce A. J. (1974) A report on a small collection of pontoniinid shrimps from the Island of Farquhar. Crustaceana, 27, 189 - 203." type="journal article" year="1974">Bruce 1974</bibRefCitation>
: 192
</treatmentCitation>
193, fig. 2; 1983: 880, figs. 4AD, 5, 6AC,
<specimenCount id="9D73FD3C9559F603F44D2D44FAC42731" box="[1294,1328,1312,1335]" pageId="32" pageNumber="33" type="generic">7F.</specimenCount>
<emphasis id="B901EAA79559F603F1D42D24FE2E2751" box="[151,474,1344,1367]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="4C754D369559F603F1D42D24FE4F2751" ID-CoL="ebead364-e4a5-4ae4-9774-e9654e54d3ea" box="[151,443,1344,1367]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ceratophthalmus">Laomenes ceratophthalmus</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AD410A49559F603F0A22D24FD0C2751" author="Okuno" box="[481,760,1344,1367]" page="119" pageId="32" pageNumber="33" year="2007">
<bibRefCitation id="EFE44B449559F603F0A22D24FD3C2751" author="Okuno" box="[481,712,1344,1367]" pageId="32" pageNumber="49" refString="Okuno, J. &amp; Fujita, Y. (2007) Resurrection of the genus Laomenes A. H. Clark, 1919 (Decapoda, Caridea, Palaemonidae). Crustaceana, 80 (1), 113 - 124" type="journal article" year="2007">Okuno &amp; Fujita 2007</bibRefCitation>
: 119
</treatmentCitation>
[excluding ovigerous female (CMNHZC 02071) described on
<figureCitation id="134E2A309559F603F1842D04FEFD2771" box="[199,265,1376,1399]" captionStart="FIGURE 2" captionStartId="5.[151,255,1961,1985]" captionTargetBox="[160,1423,201,1877]" captionTargetId="figure@5.[151,1436,194,1902]" captionTargetPageId="5" captionText="FIGURE 2. Laomenes amboinensis (De Man, 1888), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antennule. E, same, distolateral margin of basal antennular segment. F, antenna. Scale 1 mm; E, without scale." httpUri="https://zenodo.org/record/185022/files/figure.png" pageId="32" pageNumber="33">Fig. 2</figureCitation>
].
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</paragraph>
</subSubSection>
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<collectingCountry id="F36276259559F603F0502DC6FE8E27BA" box="[275,378,1442,1468]" name="Vietnam" pageId="32" pageNumber="33">Vietnam</collectingCountry>
. Nhatrang Bay:
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(pcl. 2.5,
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) (
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); 1 ovigerous female (pcl.
<quantity id="4C8D9B509559F603F4592DC6FA8D27BA" box="[1306,1401,1442,1468]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="32" pageNumber="33" unit="mm" value="3.5">3.5 mm</quantity>
),
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(pcl.
<quantity id="4C8D9B509559F603F0512DAEFE9827E2" box="[274,364,1482,1508]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="32" pageNumber="33" unit="mm" value="2.2">2.2 mm</quantity>
) (
<collectionCode id="ED64AE709559F603F0C72DAEFE1227E2" box="[388,486,1482,1508]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="32" pageNumber="33">ZMMU</collectionCode>
), Mun
<collectingCountry id="F36276259559F603F3752DAEFD7527E2" box="[566,641,1482,1508]" name="Iceland" pageId="32" pageNumber="33">Island</collectingCountry>
,
<quantity id="4C8D9B509559F603F3CF2DAEFD0127E2" box="[652,757,1482,1508]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="32" pageNumber="33" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">1520 m</quantity>
, on crinoid
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<emphasis id="B901EAA79559F603F23C2DAFFB4327E2" box="[895,1207,1483,1508]" italics="true" pageId="32" pageNumber="33">Himerometra robustipinna</emphasis>
</taxonomicName>
,
<date id="FFCB10759559F603F5862DAEFAB127E2" box="[1221,1349,1482,1508]" pageId="32" pageNumber="33" value="2006-06-18">18.vi.2006</date>
, coll. I. Marin; 1 ovigerous female (pcl.
<quantity id="4C8D9B509559F603F3552D96FD81240A" box="[534,629,1522,1548]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="32" pageNumber="33" unit="mm" value="3.5">3.5 mm</quantity>
),
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(pcl.
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) (
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), Mun
<collectingCountry id="F36276259559F603F50B2D96FB67240A" box="[1096,1171,1522,1548]" name="Iceland" pageId="32" pageNumber="33">Island</collectingCountry>
,
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,
<date id="FFCB10759559F603F4542D96FA63240A" box="[1303,1431,1522,1548]" pageId="32" pageNumber="33" value="2006-06-26">26.vi.2006</date>
, coll. I. Marin; 1 ovigerous female (pcl.
<quantity id="4C8D9B509559F603F3272E7EFD362432" box="[612,706,1562,1588]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="32" pageNumber="33" unit="mm" value="3.5">3.5 mm</quantity>
),
<specimenCount id="9D73FD3C9559F603F3952E7EFCD02432" box="[726,804,1562,1588]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
(pcl.
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),
<specimenCount id="9D73FD3C9559F603F2992E7EFBEF2432" box="[986,1051,1562,1588]" pageId="32" pageNumber="33" type="generic">1 juv.</specimenCount>
(2.0 mm) (
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),
<collectingCountry id="F36276259559F603F4502E7EFA632432" box="[1299,1431,1562,1588]" name="Iceland" pageId="32" pageNumber="33">Dun Island</collectingCountry>
,
<quantity id="4C8D9B509559F603F1D42E26FEF6245A" box="[151,258,1602,1628]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="32" pageNumber="33" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">1520 m</quantity>
, on crinoid
<taxonomicName id="4C754D369559F603F0CE2E27FD31245A" box="[397,709,1603,1628]" class="Crinoidea" family="Himerometridae" genus="Himerometra" higherTaxonomySource="CoL" kingdom="Animalia" order="Comatulida" pageId="32" pageNumber="33" phylum="Echinodermata" rank="species" species="robustipinna">
<emphasis id="B901EAA79559F603F0CE2E27FD31245A" box="[397,709,1603,1628]" italics="true" pageId="32" pageNumber="33">Himerometra robustipinna</emphasis>
</taxonomicName>
,
<date id="FFCB10759559F603F3962E26FCA1245A" box="[725,853,1602,1628]" pageId="32" pageNumber="33" value="2006-06-20">20.vi.2006</date>
, coll. I. Marin; 1 ovigerous female (pcl.
<quantity id="4C8D9B509559F603F4722E26FA7B245A" box="[1329,1423,1602,1628]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="32" pageNumber="33" unit="mm" value="3.5">3.5 mm</quantity>
),
<specimenCount id="9D73FD3C9559F603F1D42E0EFF122482" box="[151,230,1642,1668]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
(pcl.
<quantity id="4C8D9B509559F603F0682E0EFE7D2482" box="[299,393,1642,1668]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="32" pageNumber="33" unit="mm" value="2.2">2.2 mm</quantity>
), 4 juv (pcl. 1.6, 1.8, 2.0,
<quantity id="4C8D9B509559F603F3F92E0EFCE22482" box="[698,790,1642,1668]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="32" pageNumber="33" unit="mm" value="2.1">2.1 mm</quantity>
) (
<collectionCode id="ED64AE709559F603F26E2E0EFC7B2482" box="[813,911,1642,1668]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="32" pageNumber="33">ZMMU</collectionCode>
),
<collectingCountry id="F36276259559F603F2E02E0EFBDC2482" box="[931,1064,1642,1668]" name="Iceland" pageId="32" pageNumber="33">Dun Island</collectingCountry>
,
<quantity id="4C8D9B509559F603F5772E0EFB802482" box="[1076,1140,1642,1668]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="20.0">20 m</quantity>
, on crinoid
<taxonomicName id="4C754D369559F603F5BC2E0FFED824AA" class="Crinoidea" family="Himerometridae" genus="Himerometra" higherTaxonomySource="CoL" kingdom="Animalia" order="Comatulida" pageId="32" pageNumber="33" phylum="Echinodermata" rank="species" species="robustipinna">
<emphasis id="B901EAA79559F603F5BC2E0FFED824AA" italics="true" pageId="32" pageNumber="33">Himerometra robustipinna</emphasis>
</taxonomicName>
,
<date id="FFCB10759559F603F0792EF6FE4E24AA" box="[314,442,1682,1708]" pageId="32" pageNumber="33" value="2006-06-20">20.vi.2006</date>
, coll. I. Marin.
</paragraph>
</subSubSection>
<subSubSection id="C36F653E9559F602F1862EDEFEB720BC" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BCA36B59559F602F1862EDEFEB720BC" blockId="32.[151,1436,1442,2028]" lastBlockId="33.[151,1437,152,1058]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B901EAA79559F603F1862EDEFEB124D2" bold="true" box="[197,325,1722,1748]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
Carapace swollen, smooth, with antennal and hepatic teeth (
<figureCitation id="134E2A309559F603F55B2EDEFB8424D2" box="[1048,1136,1722,1748]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="32" pageNumber="33">Fig. 26</figureCitation>
A); hepatic tooth triangular, larger than antennal. Rostrum long, slender and straight, tapering distally, with 3 or 4 dorsal teeth in the distal third of dorsal carina, ventral teeth absent. Lateral rostral lamina forming conspicuous supraocular teeth in proximal part (
<figureCitation id="134E2A309559F603F0332F56FE38254A" box="[368,460,1842,1868]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="32" pageNumber="33">Fig. 26</figureCitation>
B). Orbit well developed; inferior orbital margin produced, rounded. Abdominal somites smooth; pleura of abdominal IV somites rounded ventrally (
<figureCitation id="134E2A309559F603F29F2F3EFBC22572" box="[988,1078,1882,1908]" captionStart="FIGURE 25" captionStartId="33.[151,255,2013,2037]" captionTargetBox="[171,1409,1103,2012]" captionTargetId="figure@33.[151,1436,1082,2018]" captionTargetPageId="33" captionText="FIGURE 25. Laomenes ceratophthalmus (Borradaile, 1915). A, ovigerous female, B, male. Scale 1 mm." httpUri="https://zenodo.org/record/185043/files/figure.png" pageId="32" pageNumber="33">Fig. 25</figureCitation>
). Telson about 3 times longer than wide in proximal part (
<figureCitation id="134E2A309559F603F0982FE6FDC6259A" box="[475,562,1922,1948]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="32" pageNumber="33">
Fig.
<date id="FFCB10759559F603F3512FE6FDC6259A" box="[530,562,1922,1948]" pageId="32" pageNumber="33" value="1926-01">26</date>
</figureCitation>
I), narrowing posteriorly, with 2 pairs of small submarginal dorsal spines at 0.55 and 0.75 of telson length. Eyes large (
<figureCitation id="134E2A309559F603F3D22FCEFD1D25C2" box="[657,745,1962,1988]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="32" pageNumber="33">Fig. 26</figureCitation>
C); eyestalk stout, as long as wide, with ovate cornea bearing large blunt apical papilla. Antennule well developed (
<figureCitation id="134E2A309559F603F2092FB6FC5725EA" box="[842,931,2002,2028]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="32" pageNumber="33">Fig. 26</figureCitation>
D); distolateral margin of basal antennular segment with acute distolateral tooth and well marked rounded medial projection (
<figureCitation id="134E2A309558F602F52828FCFB3622B4" box="[1131,1218,152,178]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="33" pageNumber="34">Fig. 26</figureCitation>
E, F), submarginal ventral tooth small, situated near medial border of basal segment. Mandible robust (
<figureCitation id="134E2A309558F602F5D728A4FB1822DC" box="[1172,1260,192,218]" captionStart="FIGURE 28" captionStartId="36.[151,255,1940,1964]" captionTargetBox="[194,1393,195,1890]" captionTargetId="figure@36.[181,1405,180,1908]" captionTargetPageId="36" captionText="FIGURE 28. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, mandible. C, same, distal margin of incisor process. D, maxillule. E, maxilla. F, maxilliped I. G, maxilliped II. H, maxilliped III. I, oral appendages and thoratic sternite III. Scale 1 mm; C, I, without scale." httpUri="https://zenodo.org/record/185046/files/figure.png" pageId="33" pageNumber="34">Fig. 28</figureCitation>
A, B), without palp; incisor process broad, with 7 terminal teeth (
<figureCitation id="134E2A309558F602F3A4288CFCCB2304" box="[743,831,232,258]" captionStart="FIGURE 28" captionStartId="36.[151,255,1940,1964]" captionTargetBox="[194,1393,195,1890]" captionTargetId="figure@36.[181,1405,180,1908]" captionTargetPageId="36" captionText="FIGURE 28. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, mandible. C, same, distal margin of incisor process. D, maxillule. E, maxilla. F, maxilliped I. G, maxilliped II. H, maxilliped III. I, oral appendages and thoratic sternite III. Scale 1 mm; C, I, without scale." httpUri="https://zenodo.org/record/185046/files/figure.png" pageId="33" pageNumber="34">Fig. 28</figureCitation>
C); molar process well developed, robust, with several teeth distally. Maxillule (
<figureCitation id="134E2A309558F602F3402974FDA8232C" box="[515,604,272,298]" captionStart="FIGURE 28" captionStartId="36.[151,255,1940,1964]" captionTargetBox="[194,1393,195,1890]" captionTargetId="figure@36.[181,1405,180,1908]" captionTargetPageId="36" captionText="FIGURE 28. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, mandible. C, same, distal margin of incisor process. D, maxillule. E, maxilla. F, maxilliped I. G, maxilliped II. H, maxilliped III. I, oral appendages and thoratic sternite III. Scale 1 mm; C, I, without scale." httpUri="https://zenodo.org/record/185046/files/figure.png" pageId="33" pageNumber="34">Fig. 28</figureCitation>
D) normal, with well developed bilobed palp; upper lacinia slightly curved, flaring distally, with strong setae distally; lower lacinia robust, flaring distally. Pereiopod I (
<figureCitation id="134E2A309558F602F46A295CFA8A2354" box="[1321,1406,312,338]" captionStart="FIGURE 29" captionStartId="37.[151,255,1931,1955]" captionTargetBox="[186,1407,187,1884]" captionTargetId="figure@37.[184,1408,180,1908]" captionTargetPageId="37" captionText="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="33" pageNumber="34">Fig. 29</figureCitation>
A) slender, carpus without distolateral setae at carpo-propodal articulation; dactylus and fixed finger robust with numerous terminal setae, cutting edge entire, tips not spatulated (
<figureCitation id="134E2A309558F602F2E829ECFBF223A4" box="[939,1030,392,418]" captionStart="FIGURE 29" captionStartId="37.[151,255,1931,1955]" captionTargetBox="[186,1407,187,1884]" captionTargetId="figure@37.[184,1408,180,1908]" captionTargetPageId="37" captionText="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="33" pageNumber="34">Fig. 29</figureCitation>
B). Pereiopod II equal in size and shape, with robust segments (
<figureCitation id="134E2A309558F602F0B229D4FDBC23CC" box="[497,584,432,458]" captionStart="FIGURE 29" captionStartId="37.[151,255,1931,1955]" captionTargetBox="[186,1407,187,1884]" captionTargetId="figure@37.[184,1408,180,1908]" captionTargetPageId="37" captionText="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="33" pageNumber="34">Fig. 29</figureCitation>
C); carpus flared distally, as long as maximal width, with distinct ventrolateral notch (
<figureCitation id="134E2A309558F602F07B29BCFE7A23F4" box="[312,398,472,498]" captionStart="FIGURE 29" captionStartId="37.[151,255,1931,1955]" captionTargetBox="[186,1407,187,1884]" captionTargetId="figure@37.[184,1408,180,1908]" captionTargetPageId="37" captionText="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="33" pageNumber="34">Fig. 29</figureCitation>
D, E); palm cylindrical; fingers slender, with entire cutting edges bearing 7 distinct triangular teeth; with curved tips (
<figureCitation id="134E2A309558F602F34F2A64FD9C201C" box="[524,616,512,538]" captionStart="FIGURE 29" captionStartId="37.[151,255,1931,1955]" captionTargetBox="[186,1407,187,1884]" captionTargetId="figure@37.[184,1408,180,1908]" captionTargetPageId="37" captionText="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="33" pageNumber="34">Fig. 29</figureCitation>
F). Pereiopod III (
<figureCitation id="134E2A309558F602F2082A64FC52201C" box="[843,934,512,538]" captionStart="FIGURE 29" captionStartId="37.[151,255,1931,1955]" captionTargetBox="[186,1407,187,1884]" captionTargetId="figure@37.[184,1408,180,1908]" captionTargetPageId="37" captionText="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="33" pageNumber="34">Fig. 29</figureCitation>
G) slender, segments unarmed; propodus without distoventral spines, with row of long plumose setae distally; dactylus robust (
<figureCitation id="134E2A309558F602F5DE2A4CFB012044" box="[1181,1269,552,578]" captionStart="FIGURE 29" captionStartId="37.[151,255,1931,1955]" captionTargetBox="[186,1407,187,1884]" captionTargetId="figure@37.[184,1408,180,1908]" captionTargetPageId="37" captionText="FIGURE 29. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female (A I), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm." httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="33" pageNumber="34">Fig. 29</figureCitation>
H), as long as maximal width, with long, acute and curved unguis; accessory tooth small, turned down, situates at the middle of dactylus, accompanied with tuft of long slender setae. Uropodal exopod with single mobile spine distolaterally (
<figureCitation id="134E2A309558F602F1902AC4FEDE20BC" box="[211,298,672,698]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="33" pageNumber="34">Fig. 26</figureCitation>
J).
</paragraph>
</subSubSection>
<subSubSection id="C36F653E9558F605F1862AA3FE8C2304" lastPageId="38" lastPageNumber="39" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BCA36B59558F602F1862AA3FF232624" blockId="33.[151,1437,152,1058]" pageId="33" pageNumber="34">
<emphasis id="B901EAA79558F602F1862AA3FECA20E7" bold="true" box="[197,318,711,737]" pageId="33" pageNumber="34">Remarks.</emphasis>
The current specimens are similar to the
<typeStatus id="54CE88179558F602F25C2AACFC7220E4" box="[799,902,712,738]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
described by
<bibRefCitation id="EFE44B449558F602F56B2AACFB0D20E4" author="Borradaile" box="[1064,1273,712,738]" pageId="33" pageNumber="48" refString="Borradaile, LA. (1915) Notes on Carides. Annals and Magazine of Natural History, 8, 205 - 213." type="journal article" year="1915">Borradaile (1915)</bibRefCitation>
and partly redescribed by
<bibRefCitation id="EFE44B449558F602F0702A94FE26210C" author="Bruce" box="[307,466,752,778]" pageId="33" pageNumber="48" refString="Bruce, A. J. (1978) The re-examination of some pontoniine shrimps types first described by L. A. Borradaile (Decapoda, Palaemonidae). Crustaceana, 34 (3), 251 - 268" type="journal article" year="1978">Bruce (1978)</bibRefCitation>
. Despite the
<typeStatus id="54CE88179558F602F32B2A94FD3A210C" box="[616,718,752,778]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
possibly being juvenile, it is possible to recognize such diagnostic features as short eyestalk which is about as long as wide, with moderately produced apical corneal papilla (
<bibRefCitation id="EFE44B449558F602F1B42B24FE4E215C" author="Borradaile" box="[247,442,832,858]" pageId="33" pageNumber="48" refString="Borradaile, L. A. (1917) On the Pontoniinae. The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner. Transactions of the Linnean Society of London, Zoology, 17 (2), 323 - 396." type="journal article" year="1917">Borradaile 1917</bibRefCitation>
,
<figureCitation id="134E2A309558F602F0842B24FDF9215C" box="[455,525,832,858]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="33" pageNumber="34">Fig. 9</figureCitation>
a, b). The rostrum of our specimens bears 3 or 4 dorsal teeth and is toothless ventrally (
<figureCitation id="134E2A309558F602F0522B0CFE9D2184" box="[273,361,872,898]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="33" pageNumber="34">Fig. 26</figureCitation>
A, 27A, C, E) similar to most references to this species (
<bibRefCitation id="EFE44B449558F602F5532B0CFB4B2184" author="Holthuis" box="[1040,1215,872,898]" pageId="33" pageNumber="48" refString="Holthuis, L. B. (1952) The Decapoda of the Siboga Expedition. Part XI. The Palaemonidae collected by the Siboga and Snellius Expeditions with remarks on other species. II. Subfamily Pontoniinae. Siboga Expedition Monograph, 39 a, 10, 1 - 252." type="journal article" year="1952">Holthuis 1952</bibRefCitation>
;
<bibRefCitation id="EFE44B449558F602F58F2B0CFF2121AC" author="Miyake" pageId="33" pageNumber="49" refString="Miyake, S. &amp; Fujino, T. (1968) Pontoniid shrimps from Palau Islands. Journal of the Faculty of Agriculture, Kyushu University, 14 (3), 399 - 431." type="journal article" year="1968">Miyake &amp; Fujino 1968</bibRefCitation>
;
<bibRefCitation id="EFE44B449558F602F1A12BF4FE9921AC" author="Bruce" box="[226,365,912,938]" pageId="33" pageNumber="48" refString="Bruce A. J. (1974) A report on a small collection of pontoniinid shrimps from the Island of Farquhar. Crustaceana, 27, 189 - 203." type="journal article" year="1974">Bruce 1974</bibRefCitation>
,
<bibRefCitation id="EFE44B449558F602F0392BF4FE4D21AC" author="Bruce" box="[378,441,912,938]" pageId="33" pageNumber="48" refString="Bruce A. J. (1983) Expedition Rumphius II (1975). Crustaces parasites, commensaux, etc. IX: Crustaces Decapodes (1 ere partie: Natantia Pontoniinae). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4, 5 A, 871 - 902." type="journal article" year="1983">1983</bibRefCitation>
). The rostrum of the
<typeStatus id="54CE88179558F602F3F72BF4FD1221AC" box="[692,742,912,938]" pageId="33" pageNumber="34">type</typeStatus>
specimen “
<emphasis id="B901EAA79558F602F2302BF4FA7221AF" box="[883,1414,912,937]" italics="true" pageId="33" pageNumber="34">bearing above four teeth, all in its distal half</emphasis>
”; the figure (
<figureCitation id="134E2A309558F602F0542BDCFEAF21D4" box="[279,347,952,978]" captionStart="FIGURE 2" captionStartId="5.[151,255,1961,1985]" captionTargetBox="[160,1423,201,1877]" captionTargetId="figure@5.[151,1436,194,1902]" captionTargetPageId="5" captionText="FIGURE 2. Laomenes amboinensis (De Man, 1888), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antennule. E, same, distolateral margin of basal antennular segment. F, antenna. Scale 1 mm; E, without scale." httpUri="https://zenodo.org/record/185022/files/figure.png" pageId="33" pageNumber="34">Fig. 2</figureCitation>
b in
<bibRefCitation id="EFE44B449558F602F0CD2BDCFDB921D4" author="Borradaile" box="[398,589,952,978]" pageId="33" pageNumber="48" refString="Borradaile, L. A. (1917) On the Pontoniinae. The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner. Transactions of the Linnean Society of London, Zoology, 17 (2), 323 - 396." type="journal article" year="1917">Borradaile 1917</bibRefCitation>
) shows a slender rostrum with the position of the posterior tooth close to the mid-length of the rostrum similarly to one of the juvenile specimens presented in our collection (see
<figureCitation id="134E2A309558F602F42E2B84FF432624" captionStart="FIGURE 27" captionStartId="35.[151,255,1336,1360]" captionTargetBox="[174,1408,204,1289]" captionTargetId="figure@35.[153,1438,194,1307]" captionTargetPageId="35" captionText="FIGURE 27. Laomenes ceratophthalmus (Borradaile, 1915), juvenile (pcl. 3 mm) (A, B), juvenile (pcl. 6 mm) (C, D), juvenile (pcl. 11 mm) (E). A E, distal part of carapace and rostrum. Scale 1 mm." httpUri="https://zenodo.org/record/185045/files/figure.png" pageId="33" pageNumber="34">Fig. 27</figureCitation>
E).
</paragraph>
<caption id="DF0A663D9558F602F1D42FB9FB0C25F2" box="[151,1272,2013,2037]" httpUri="https://zenodo.org/record/185043/files/figure.png" pageId="33" pageNumber="34" targetBox="[171,1409,1103,2012]" targetPageId="33">
<paragraph id="8BCA36B59558F602F1D42FB9FB0C25F2" blockId="33.[151,1272,2013,2037]" box="[151,1272,2013,2037]" pageId="33" pageNumber="34">
<emphasis id="B901EAA79558F602F1D42FB9FED225F3" bold="true" box="[151,294,2013,2037]" pageId="33" pageNumber="34">FIGURE 25.</emphasis>
<taxonomicName id="4C754D369558F602F06E2FBAFCD625F2" authority="(Borradaile, 1915)" authorityName="Borradaile" authorityYear="1915" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[301,802,2013,2037]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA79558F602F06E2FBAFDA725F3" box="[301,595,2014,2037]" italics="true" pageId="33" pageNumber="34">Laomenes ceratophthalmus</emphasis>
(Borradaile, 1915)
</taxonomicName>
. A, ovigerous female, B, male. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF0A663D955BF601F1D42FFCFCDA25F3" httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="34" pageNumber="35" targetBox="[196,1398,199,1911]" targetPageId="34">
<paragraph id="8BCA36B5955BF601F1D42FFCFCDA25F3" blockId="34.[151,1436,1944,2037]" pageId="34" pageNumber="35">
<emphasis id="B901EAA7955BF601F1D42FFCFEDD25B6" bold="true" box="[151,297,1944,1968]" pageId="34" pageNumber="35">FIGURE 26.</emphasis>
<taxonomicName id="4C754D36955BF601F0712FFDFCD825A9" authority="(Borradaile, 1915)" authorityName="Borradaile" authorityYear="1915" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[306,812,1944,1968]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955BF601F0712FFDFDAE25B6" box="[306,602,1945,1968]" italics="true" pageId="34" pageNumber="35">Laomenes ceratophthalmus</emphasis>
(Borradaile, 1915)
</taxonomicName>
, ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF0A663D955AF600F1D42D5CFC012774" httpUri="https://zenodo.org/record/185045/files/figure.png" pageId="35" pageNumber="36" targetBox="[174,1408,204,1289]" targetPageId="35">
<paragraph id="8BCA36B5955AF600F1D42D5CFC012774" blockId="35.[151,1436,1336,1394]" pageId="35" pageNumber="36">
<emphasis id="B901EAA7955AF600F1D42D5CFEDC2756" bold="true" box="[151,296,1336,1360]" pageId="35" pageNumber="36">FIGURE 27.</emphasis>
<taxonomicName id="4C754D36955AF600F06C2D5CFCD32749" authority="(Borradaile, 1915)" authorityName="Borradaile" authorityYear="1915" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[303,807,1336,1359]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F06C2D5CFDA22749" box="[303,598,1336,1359]" italics="true" pageId="35" pageNumber="36">Laomenes ceratophthalmus</emphasis>
(Borradaile, 1915)
</taxonomicName>
, juvenile (pcl. 3 mm) (A, B), juvenile (pcl. 6 mm) (C, D), juvenile (pcl. 11 mm) (E). AE, distal part of carapace and rostrum. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BCA36B5955AF600F1862DACFCA825EC" blockId="35.[151,1436,1480,2026]" pageId="35" pageNumber="36">
<collectingCountry id="F3627625955AF600F1862DACFF1127E4" box="[197,229,1480,1506]" name="American Samoa" pageId="35" pageNumber="36">As</collectingCountry>
indicated above,
<taxonomicName id="4C754D36955AF600F0FD2DACFCF027E7" box="[446,772,1480,1505]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F0FD2DACFCF027E7" box="[446,772,1480,1505]" italics="true" pageId="35" pageNumber="36">Laomenes ceratophthalmus</emphasis>
</taxonomicName>
was previously misidentified with
<taxonomicName id="4C754D36955AF600F5EC2DACFA9A27E7" box="[1199,1390,1480,1505]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="B901EAA7955AF600F5EC2DACFA9A27E7" box="[1199,1390,1480,1505]" italics="true" pageId="35" pageNumber="36">Laomenes tigris</emphasis>
</taxonomicName>
<emphasis id="B901EAA7955AF600F43A2DACFF3E240C" bold="true" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A23257DC955AF600F43A2DACFF3E240C" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Both species share a rostrum with dorsal teeth situated in its distal part and ventrally toothless. Nevertheless, these species can be clearly distinguished by the form of the eyes (eyestalk of
<taxonomicName id="4C754D36955AF600F53D2E7CFA902437" box="[1150,1380,1560,1585]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F53D2E7CFA902437" box="[1150,1380,1560,1585]" italics="true" pageId="35" pageNumber="36">L. ceratophthalmus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE44B44955AF600F1D42E24FED5245C" author="Bruce" box="[151,289,1600,1626]" pageId="35" pageNumber="48" refString="Bruce, A. J. (1978) The re-examination of some pontoniine shrimps types first described by L. A. Borradaile (Decapoda, Palaemonidae). Crustaceana, 34 (3), 251 - 268" type="journal article" year="1978">Bruce 1978</bibRefCitation>
,
<figureCitation id="134E2A30955AF600F06F2E24FE84245C" box="[300,368,1600,1626]" captionStart="FIGURE 2" captionStartId="5.[151,255,1961,1985]" captionTargetBox="[160,1423,201,1877]" captionTargetId="figure@5.[151,1436,194,1902]" captionTargetPageId="5" captionText="FIGURE 2. Laomenes amboinensis (De Man, 1888), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antennule. E, same, distolateral margin of basal antennular segment. F, antenna. Scale 1 mm; E, without scale." httpUri="https://zenodo.org/record/185022/files/figure.png" pageId="35" pageNumber="36">Fig. 2</figureCitation>
A; this paper,
<figureCitation id="134E2A30955AF600F3502E24FD93245C" box="[531,615,1600,1626]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="35" pageNumber="36">Fig. 26</figureCitation>
C) is significantly shorter than in
<taxonomicName id="4C754D36955AF600F2A82E24FBB3245F" box="[1003,1095,1600,1625]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="B901EAA7955AF600F2A82E24FBB3245F" box="[1003,1095,1600,1625]" italics="true" pageId="35" pageNumber="36">L. tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC955AF600F50E2E24FB5E245C" box="[1101,1194,1600,1626]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B901EAA7955AF600F50E2E24FB5E245C" bold="true" box="[1101,1194,1600,1626]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
<bibRefCitation id="EFE44B44955AF600F5F22E24FE632484" author="Kemp" pageId="35" pageNumber="49" refString="Kemp, S. (1925) Notes on Crustacea Decapoda in the Indian Museum, XVII: on various Caridea. Records of the Indian Museum, 27, 249 - 343." type="journal article" year="1925" yearSuffix="a">
(Kemp 1925, Fig. a; this paper,
<figureCitation id="134E2A30955AF600F05B2E0CFE992484" box="[280,365,1640,1666]" captionStart="FIGURE 20" captionStartId="26.[151,255,1663,1687]" captionTargetBox="[166,1427,161,1612]" captionTargetId="figure@26.[153,1438,147,1659]" captionTargetPageId="26" captionText="FIGURE 20. Laomenes tigris sp. nov., ovigerous female (A E), male (F, G). A, B, F, G, distal part of carapace and rostrum. C, distal part of carapace. D, eyes. Scale 1 mm." httpUri="https://zenodo.org/record/185038/files/figure.png" pageId="35" pageNumber="36">Fig. 20</figureCitation>
D))
</bibRefCitation>
, by the presence of dorsal teeth posterior the mid-length of rostrum in
<taxonomicName id="4C754D36955AF600F59A2E0CFAC22487" box="[1241,1334,1640,1665]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="B901EAA7955AF600F59A2E0CFAC22487" box="[1241,1334,1640,1665]" italics="true" pageId="35" pageNumber="36">L. tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC955AF600F47D2E0CFA682484" box="[1342,1436,1640,1666]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B901EAA7955AF600F47D2E0CFA682484" bold="true" box="[1342,1436,1640,1666]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
(vs. all dorsal teeth situated anterior to mid-length of rostrum in
<taxonomicName id="4C754D36955AF600F2C62EF4FB9D24AF" box="[901,1129,1680,1705]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F2C62EF4FB9D24AF" box="[901,1129,1680,1705]" italics="true" pageId="35" pageNumber="36">L. ceratophthalmus</emphasis>
</taxonomicName>
). The propodus of pereiopod III in
<taxonomicName id="4C754D36955AF600F04F2EDCFE9D24D7" box="[268,361,1720,1745]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="B901EAA7955AF600F04F2EDCFE9D24D7" box="[268,361,1720,1745]" italics="true" pageId="35" pageNumber="36">L. tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC955AF600F0332EDCFE3924D4" box="[368,461,1720,1746]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B901EAA7955AF600F0332EDCFE3924D4" bold="true" box="[368,461,1720,1746]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
possesses about a dozen rows of simple setae along the distoventral margin (
<bibRefCitation id="EFE44B44955AF600F4112EDCFEA324FC" author="Kemp" pageId="35" pageNumber="49" refString="Kemp, S. (1925) Notes on Crustacea Decapoda in the Indian Museum, XVII: on various Caridea. Records of the Indian Museum, 27, 249 - 343." type="journal article" year="1925">Kemp 1925; this paper</bibRefCitation>
) while only single distal row setae present in
<taxonomicName id="4C754D36955AF600F2362E84FBAE24FF" box="[885,1114,1760,1785]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F2362E84FBAE24FF" box="[885,1114,1760,1785]" italics="true" pageId="35" pageNumber="36">L. ceratophthalmus</emphasis>
</taxonomicName>
. The distolateral margin of the uropodal exopod in mature specimens of
<taxonomicName id="4C754D36955AF600F3E92F6CFC7B2527" box="[682,911,1800,1825]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F3E92F6CFC7B2527" box="[682,911,1800,1825]" italics="true" pageId="35" pageNumber="36">L. ceratophthalmus</emphasis>
</taxonomicName>
shows only a mobile spine, whilst the distolateral margin of the exopod in
<taxonomicName id="4C754D36955AF600F3492F54FD93254F" box="[522,615,1840,1865]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="B901EAA7955AF600F3492F54FD93254F" box="[522,615,1840,1865]" italics="true" pageId="35" pageNumber="36">L. tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC955AF600F32C2F54FD39254C" box="[623,717,1840,1866]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B901EAA7955AF600F32C2F54FD39254C" bold="true" box="[623,717,1840,1866]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
bears a well-marked fixed tooth and a mobile spine. Further, a tiny fixed tooth on distolateral margin of exopod of uropod can be found in juvenile specimens of
<taxonomicName id="4C754D36955AF600F40F2F3CFEC3259F" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F40F2F3CFEC3259F" italics="true" pageId="35" pageNumber="36">L. ceratophthalmus</emphasis>
</taxonomicName>
. It is most likely that the specimen described by Okuno &amp; Fujita (2006, fig. 2) belongs to
<taxonomicName id="4C754D36955AF600F1D42FCCFEA125C7" box="[151,341,1960,1985]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="B901EAA7955AF600F1D42FCCFEA125C7" box="[151,341,1960,1985]" italics="true" pageId="35" pageNumber="36">Laomenes tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC955AF600F0232FCCFE3425C4" box="[352,448,1960,1986]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B901EAA7955AF600F0232FCCFE3425C4" bold="true" box="[352,448,1960,1986]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
as the eyestalk is significantly longer that in true
<taxonomicName id="4C754D36955AF600F5662FCCFAF825C7" box="[1061,1292,1960,1985]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955AF600F5662FCCFAF825C7" box="[1061,1292,1960,1985]" italics="true" pageId="35" pageNumber="36">L. ceratophthalmus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE44B44955AF600F45D2FCCFF2025EC" author="Borradaile" pageId="35" pageNumber="48" refString="Borradaile, L. A. (1917) On the Pontoniinae. The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner. Transactions of the Linnean Society of London, Zoology, 17 (2), 323 - 396." type="journal article" year="1917">Borradaile 1917</bibRefCitation>
,
<figureCitation id="134E2A30955AF600F1A32FB4FED225EC" box="[224,294,2000,2026]" captionStart="FIGURE 2" captionStartId="5.[151,255,1961,1985]" captionTargetBox="[160,1423,201,1877]" captionTargetId="figure@5.[151,1436,194,1902]" captionTargetPageId="5" captionText="FIGURE 2. Laomenes amboinensis (De Man, 1888), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antennule. E, same, distolateral margin of basal antennular segment. F, antenna. Scale 1 mm; E, without scale." httpUri="https://zenodo.org/record/185022/files/figure.png" pageId="35" pageNumber="36">Fig. 2</figureCitation>
a;
<bibRefCitation id="EFE44B44955AF600F0022FB4FE3F25EC" author="Bruce" box="[321,459,2000,2026]" pageId="35" pageNumber="48" refString="Bruce, A. J. (1978) The re-examination of some pontoniine shrimps types first described by L. A. Borradaile (Decapoda, Palaemonidae). Crustaceana, 34 (3), 251 - 268" type="journal article" year="1978">Bruce 1978</bibRefCitation>
, fig. 2A; present paper,
<figureCitation id="134E2A30955AF600F3A72FB4FCCE25EC" box="[740,826,2000,2026]" captionStart="FIGURE 26" captionStartId="34.[151,255,1944,1968]" captionTargetBox="[196,1398,199,1911]" captionTargetId="figure@34.[184,1408,194,1922]" captionTargetPageId="34" captionText="FIGURE 26. Laomenes ceratophthalmus (Borradaile, 1915), ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, antenna. E, F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, telson. J, distolateral margin of uropodal exopod. Scale 1 mm." httpUri="https://zenodo.org/record/185044/files/figure.png" pageId="35" pageNumber="36">Fig. 26</figureCitation>
C).
</paragraph>
<caption id="DF0A663D955DF607F1D42FF0FD3625F7" httpUri="https://zenodo.org/record/185046/files/figure.png" pageId="36" pageNumber="37" targetBox="[194,1393,195,1890]" targetPageId="36">
<paragraph id="8BCA36B5955DF607F1D42FF0FD3625F7" blockId="36.[151,1436,1940,2033]" pageId="36" pageNumber="37">
<emphasis id="B901EAA7955DF607F1D42FF0FED325AA" bold="true" box="[151,295,1940,1964]" pageId="36" pageNumber="37">FIGURE 28.</emphasis>
<taxonomicName id="4C754D36955DF607F06C2FF1FCD125AD" authority="(Borradaile, 1915)" authorityName="Borradaile" authorityYear="1915" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[303,805,1940,1964]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955DF607F06C2FF1FDA125AA" box="[303,597,1941,1964]" italics="true" pageId="36" pageNumber="37">Laomenes ceratophthalmus</emphasis>
(Borradaile, 1915)
</taxonomicName>
, ovigerous female. A, B, mandible. C, same, distal margin of incisor process. D, maxillule. E, maxilla. F, maxilliped I. G, maxilliped II. H, maxilliped III. I, oral appendages and thoratic sternite III. Scale 1 mm; C, I, without scale.
</paragraph>
</caption>
<caption id="DF0A663D955CF606F1D42FEFFCAB25E1" httpUri="https://zenodo.org/record/185047/files/figure.png" pageId="37" pageNumber="38" targetBox="[186,1407,187,1884]" targetPageId="37">
<paragraph id="8BCA36B5955CF606F1D42FEFFCAB25E1" blockId="37.[151,1436,1931,2023]" pageId="37" pageNumber="38">
<emphasis id="B901EAA7955CF606F1D42FEFFED225A5" bold="true" box="[151,294,1931,1955]" pageId="37" pageNumber="38">FIGURE 29.</emphasis>
<taxonomicName id="4C754D36955CF606F06E2FEFFCD525A4" authority="(Borradaile, 1915)" authorityName="Borradaile" authorityYear="1915" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[301,801,1931,1954]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B901EAA7955CF606F06E2FEFFDA725A4" box="[301,595,1931,1954]" italics="true" pageId="37" pageNumber="38">Laomenes ceratophthalmus</emphasis>
(Borradaile, 1915)
</taxonomicName>
, ovigerous female (AI), male (J). A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. G, pereiopod III. H, same, distal margin of propodus and dactylus. I, endopod of pleopod I. J, appendix masculina. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BCA36B5955FF605F18628F3FE8C2304" blockId="38.[151,1437,151,618]" pageId="38" pageNumber="39">
<emphasis id="B901EAA7955FF605F18628F3FEA022B7" bold="true" box="[197,340,151,177]" pageId="38" pageNumber="39">Coloration.</emphasis>
General coloration of the body and appendages is red (
<figureCitation id="134E2A30955FF605F2A328FCFBC122B4" box="[992,1077,152,178]" captionStart="FIGURE 33" captionStartId="41.[151,255,1584,1608]" captionTargetBox="[151,1436,194,1553]" captionTargetId="figure@41.[151,1436,194,1555]" captionTargetPageId="41" captionText="FIGURE 33. Coloration of Laomenes ceratophthalmus (Borradaile, 1915) (A, C, E) and Laomenes nudirostris (Bruce, 1978) (B, D, F). A D, ovigerous females. E, F, males." httpUri="https://zenodo.org/record/185051/files/figure.png" pageId="38" pageNumber="39">Fig. 33</figureCitation>
A, C, E); carapace with 2 dark red longitudinal bands dorsally, with 3 additional dark red longitudinal bands continuing to abdomen dorsolaterally and laterally.
</paragraph>
</subSubSection>
<subSubSection id="C36F653E955FF605F186296BFB9B23F7" pageId="38" pageNumber="39" type="description">
<paragraph id="8BCA36B5955FF605F186296BFB9B23F7" blockId="38.[151,1437,151,618]" pageId="38" pageNumber="39">
<emphasis id="B901EAA7955FF605F186296BFEF3232F" bold="true" box="[197,263,271,297]" pageId="38" pageNumber="39">Host.</emphasis>
All specimens were collected exclusively from himerometrid crinoids
<taxonomicName id="4C754D36955FF605F5232974FE9C2354" authority="Carpenter, 1881" authorityName="Carpenter" authorityYear="1881" class="Crinoidea" family="Himerometridae" genus="Himerometra" higherTaxonomySource="CoL" kingdom="Animalia" order="Comatulida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="species" species="robustipinna">
<emphasis id="B901EAA7955FF605F5232974FA68232F" box="[1120,1436,272,297]" italics="true" pageId="38" pageNumber="39">Himerometra robustipinna</emphasis>
(Carpenter, 1881)
</taxonomicName>
(Crinoidea:
<taxonomicName id="4C754D36955FF605F0BC295CFD352354" box="[511,705,312,338]" class="Crinoidea" family="Himerometridae" kingdom="Animalia" order="Comatulida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="family">Himerometridae</taxonomicName>
) (
<figureCitation id="134E2A30955FF605F398295CFCC72354" box="[731,819,312,338]" captionStart="FIGURE 34" captionStartId="42.[151,255,1893,1917]" captionTargetBox="[151,1436,182,1846]" captionTargetId="figure@42.[151,1436,182,1846]" captionTargetPageId="42" captionText="FIGURE 34. Crinoid hosts of the representatives of the genus Laomenes A. H. Clark. A, Himerometra robustipinna (Carpenter, 1881) (Himerometridae). B, Amphimetra tesselata (Müller, 1841) (Himerometridae). C, Oxycomanthus benetti (Müller, 1841) (Comasteridae). D, F, Comantheria (Comanthus) briareus (Bell, 1882) (Comasteridae). E, Zygometra punctata A. H. Clark, 1912 (Zygometridae)." httpUri="https://zenodo.org/record/185052/files/figure.png" pageId="38" pageNumber="39">Fig. 34</figureCitation>
A). Other species of crinoids elsewhere observed as hosts of this species are
<taxonomicName id="4C754D36955FF605F0872904FBAB237C" authority="A. H. Clark, 1912" authorityName="A. H. Clark" authorityYear="1912" box="[452,1119,352,378]" class="Crinoidea" family="Mariametridae" genus="Dichrometra" kingdom="Animalia" order="Comatulida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="species" species="afra">
<emphasis id="B901EAA7955FF605F0872904FD62237F" box="[452,662,352,377]" italics="true" pageId="38" pageNumber="39">Dichrometra afra</emphasis>
A. H. Clark, 1912 (Himerometridae)
</taxonomicName>
,
<taxonomicName id="4C754D36955FF605F52E2904FEAA23A4" authority="Hartlaub, 1890" authorityName="Hartlaub" authorityYear="1890" class="Crinoidea" family="Mariametridae" genus="Lamprometra" kingdom="Animalia" order="Comatulida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="species" species="klunzingeri">
<emphasis id="B901EAA7955FF605F52E2904FA68237F" box="[1133,1436,352,377]" italics="true" pageId="38" pageNumber="39">Lamprometra klunzingeri</emphasis>
(Hartlaub, 1890)
</taxonomicName>
,
<taxonomicName id="4C754D36955FF605F02929ECFCE323A4" authority="Smith, 1876" authorityName="Smith" authorityYear="1876" box="[362,791,392,418]" class="Crinoidea" family="Mariametridae" genus="Stephanometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="species" species="indica">
<emphasis id="B901EAA7955FF605F02929ECFD9823A7" box="[362,620,392,417]" italics="true" pageId="38" pageNumber="39">Stephanometra indica</emphasis>
(Smith, 1876)
</taxonomicName>
and
<taxonomicName id="4C754D36955FF605F21129ECFA9123A4" authority="Carpenter, 1881" authorityName="Carpenter" authorityYear="1881" box="[850,1381,392,418]" class="Crinoidea" family="Mariametridae" genus="Stephanometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="species" species="spicata">
<emphasis id="B901EAA7955FF605F21129ECFC3123A7" box="[850,965,392,417]" italics="true" pageId="38" pageNumber="39">S. spicata</emphasis>
(Carpenter, 1881) (Mariametridae)
</taxonomicName>
(see
<bibRefCitation id="EFE44B44955FF605F1D429D4FEDF23CC" author="Bruce" box="[151,299,432,458]" pageId="38" pageNumber="48" refString="Bruce, A. J. (1981) Pontoniine shrimps of Heron Island. Atoll Research Bulletin, 245, 1 - 33." type="journal article" year="1981">Bruce, 1981</bibRefCitation>
,
<bibRefCitation id="EFE44B44955FF605F07A29D4FE8223CC" author="Bruce" box="[313,374,432,458]" pageId="38" pageNumber="48" refString="Bruce, A. J. (1982) The shrimps associated with Indo-Pacific echinoderms, with the description of a new species in the genus Periclimenes Costa (Crustacea, Pontoniinae). Australian Museum Memoir, 16, 192 - 216." type="journal article" year="1982">1982</bibRefCitation>
,
<bibRefCitation id="EFE44B44955FF605F0C629D4FE3723CC" author="Bruce" box="[389,451,432,458]" pageId="38" pageNumber="48" refString="Bruce A. J. (1983) Expedition Rumphius II (1975). Crustaces parasites, commensaux, etc. IX: Crustaces Decapodes (1 ere partie: Natantia Pontoniinae). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4, 5 A, 871 - 902." type="journal article" year="1983">1983</bibRefCitation>
), but it is possible that these records refer to similar undescribed species. Equally, records from the family
<taxonomicName id="4C754D36955FF605F0F729BCFD9423F4" box="[436,608,472,498]" class="Crinoidea" family="Mariametridae" kingdom="Animalia" order="Comatulida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="family">Mariametridae</taxonomicName>
most likely refer to
<taxonomicName id="4C754D36955FF605F20D29BCFBFF23F7" box="[846,1035,472,497]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="B901EAA7955FF605F20D29BCFBFF23F7" box="[846,1035,472,497]" italics="true" pageId="38" pageNumber="39">Laomenes tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC955FF605F55129B3FB9B23F7" box="[1042,1135,471,497]" pageId="38" pageNumber="39" rank="species">
<emphasis id="B901EAA7955FF605F55129B3FB9B23F7" bold="true" box="[1042,1135,471,497]" pageId="38" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C36F653E955FF605F186299BFDB4206C" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BCA36B5955FF605F186299BFDB4206C" blockId="38.[151,1437,151,618]" pageId="38" pageNumber="39">
<emphasis id="B901EAA7955FF605F186299BFE92201F" bold="true" box="[197,358,511,537]" pageId="38" pageNumber="39">Distribution.</emphasis>
Widely distributed in Indo-West Pacific. Known from
<collectingCountry id="F3627625955FF605F54D2A64FBAA201C" box="[1038,1118,512,538]" name="Kenya" pageId="38" pageNumber="39">Kenya</collectingCountry>
,
<collectingCountry id="F3627625955FF605F52D2A64FB2C201C" box="[1134,1240,512,538]" name="Tanzania" pageId="38" pageNumber="39">Zanzibar</collectingCountry>
, the
<collectingCountry id="F3627625955FF605F45B2A64FA6C201C" box="[1304,1432,512,538]" name="Seychelles" pageId="38" pageNumber="39">Seychelles</collectingCountry>
,
<collectingCountry id="F3627625955FF605F1D42A4CFEAF2044" box="[151,347,552,578]" name="Maldives" pageId="38" pageNumber="39">Maldive Islands</collectingCountry>
,
<collectingCountry id="F3627625955FF605F0282A4CFE142044" box="[363,480,552,578]" name="Indonesia" pageId="38" pageNumber="39">Indonesia</collectingCountry>
,
<collectingCountry id="F3627625955FF605F0B22A4CFDAC2044" box="[497,600,552,578]" name="Vietnam" pageId="38" pageNumber="39">Vietnam</collectingCountry>
, the Great Barrier Reef of
<collectingCountry id="F3627625955FF605F2EF2A4CFBEF2044" box="[940,1051,552,578]" name="Australia" pageId="38" pageNumber="39">Australia</collectingCountry>
,
<collectingCountry id="F3627625955FF605F5682A4CFAEF2044" box="[1067,1307,552,578]" name="Papua New Guinea" pageId="38" pageNumber="39">Papua New Guinea</collectingCountry>
, the
<collectingCountry id="F3627625955FF605F4182A4CFA682044" box="[1371,1436,552,578]" name="Palau" pageId="38" pageNumber="39">Palau</collectingCountry>
Islands,
<collectingCountry id="F3627625955FF605F1BB2A34FE49206C" box="[248,445,592,618]" name="Solomon Islands" pageId="38" pageNumber="39">Solomon Islands</collectingCountry>
and
<collectingCountry id="F3627625955FF605F0B62A34FDC8206C" box="[501,572,592,618]" name="Japan" pageId="38" pageNumber="39">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>