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<document id="60E0BA1595E4BCC4C96479A127146B58" ID-DOI="10.11646/zootaxa.4088.2.3" ID-GBIF-Dataset="89bf91a3-7c89-4d23-ae74-caffb9164795" ID-ISSN="1175-5326" ID-Zenodo-Dep="267005" ID-ZooBank="431D8F02-C8D3-4E97-8FAC-04558E5BA60C" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1457520271182" checkinUser="donat" docAuthor="Han, H. L., Pan, Z. H. &amp; Kononenko, V. S." docDate="2016" docId="03DC8796C058FF9E3C998595FB66E9C2" docLanguage="en" docName="15567-5097-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atrovirensis sacculatus Han, Pan &amp; Kononenko, 2016, sp. n." docType="treatment" docVersion="10" lastPageNumber="216" masterDocId="FFE5FFEEC056FF913C0E8337FFDEED2A" masterDocTitle="A review of the genus Atrovirensis Kononenko, 2001 with description of four new species from China (Lepidoptera, Noctuidae: Xyleninae, Apameini)" masterLastPageNumber="220" masterPageNumber="201" pageNumber="215" updateTime="1698177376185" updateUser="plazi">
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<mods:title id="95A64BFA6D4C552117560D31354462BD">A review of the genus Atrovirensis Kononenko, 2001 with description of four new species from China (Lepidoptera, Noctuidae: Xyleninae, Apameini)</mods:title>
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<mods:namePart id="ECE2821D4085B893C36DCCA26A9D1023">Han, H. L.</mods:namePart>
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<mods:namePart id="501438B6EA7D33A6E8B7D1B00D3C37AD">Pan, Z. H.</mods:namePart>
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<mods:namePart id="0D277B2BCA0D5DA0A140E9DE32F315E3">Kononenko, V. S.</mods:namePart>
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<mods:date id="1759D2211732B884B8AC11BA57C7FAD5">2016</mods:date>
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<treatment id="03DC8796C058FF9E3C998595FB66E9C2" ID-DOI="http://doi.org/10.5281/zenodo.6084435" ID-GBIF-Taxon="117196261" ID-Zenodo-Dep="6084435" LSID="urn:lsid:plazi:treatment:03DC8796C058FF9E3C998595FB66E9C2" httpUri="http://treatment.plazi.org/id/03DC8796C058FF9E3C998595FB66E9C2" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
<subSubSection id="C36F650BC058FF9F3C998595FEA4EBF4" pageId="14" pageNumber="215" type="nomenclature">
<paragraph id="8BCA3680C058FF9F3C998595FE27EB97" blockId="14.[151,505,1698,1758]" box="[151,505,1698,1725]" pageId="14" pageNumber="215">
<heading id="D08281ECC058FF9F3C998595FE27EB97" bold="true" box="[151,505,1698,1725]" fontSize="11" level="1" pageId="14" pageNumber="215" reason="1">
<emphasis id="B901EA92C058FF9F3C998595FE27EB97" bold="true" box="[151,505,1698,1725]" pageId="14" pageNumber="215">
<taxonomicName id="4C754D03C058FF9F3C998595FE6EEB96" box="[151,432,1698,1724]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="sacculatus" status="sp. nov.">
<emphasis id="B901EA92C058FF9F3C998595FE6EEB96" bold="true" box="[151,432,1698,1724]" italics="true" pageId="14" pageNumber="215">Atrovirensis sacculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257E9C058FF9F3DB98594FE27EB97" box="[439,505,1699,1725]" pageId="14" pageNumber="215" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA3680C058FF9F3C9985F2FEA4EBF4" blockId="14.[151,505,1698,1758]" box="[151,378,1733,1758]" pageId="14" pageNumber="215">
(
<figureCitation id="134E2A05C058FF9F3C9185F2FEC1EBF4" box="[159,287,1733,1758]" captionStart="FIGURES 1 15" captionStartId="3.[151,263,1738,1759]" captionTargetBox="[190,1394,195,1718]" captionTargetId="figure@3.[190,1395,195,1719]" captionTargetPageId="3" captionText="FIGURES 1 15. Atrovirensis spp., adults. 1. A. atrovirens, Nepal (after Gyulai et al. 2013); 2. A. annamita, HT, Vietnam (after Gyulai et al. 2013); 3. A. parannamita sp. n., male HT, China, Aut. Reg. Xizang, genit. prep. HHL- 2930 - 1; 4. A. parannamita sp. n., female PT, China, Aut. Reg. Xizang, genit. prep. HHL- 2906 - 2; 5. A. pedibus, HT, China, Prov. Hubei (after Gyulai et al. 2013); 6. A. sp. cf. A. parannamita, male, China, Aut. Reg. Xizang, genit. prep. HHL- 2907 - 1; 7. A. sinica, HT, China, Prov. Guangdong, genit. prep. ZFMK 1624; 8. A. owadai, Prov. Guangdong, HT, China (after Gyulai et al. 2013); 9. A. taiwani, HT, Taiwan (after Gyulai et al. 2013); 10. A. furcatus sp. n., male HT, China, Prov. Guangdong, genit. prep. HHL- 2921 - 1; 11. A. furcatus sp. n., female PT, China, Prov. Guangdong, genit. prep. HHL- 2919 - 2; 12. A. sp. cf. furcatus, male, China, Prov. Guangdong, genit. prep. HHL- 2918 - 1; 13. A. sp. cf. furcatus, female, China, Prov. Guangdong, genit. prep. HHL- 2909 - 2; 14. A. sacculatus sp. n., male HT, China, Aut. Reg. Xizang, genit. prep. HHL- 2910 - 1; 15. A. sacculatus sp. n., male PT, China, Aut. Reg. Xizang, genit. prep. HHL- 2908 - 1." httpUri="https://zenodo.org/record/267006/files/figure.png" pageId="14" pageNumber="215">Figs 14, 15</figureCitation>
,
<figureCitation id="134E2A05C058FF9F3D2585F1FE97EBF4" box="[299,329,1734,1758]" captionStart="FIGURES 30 32" captionStartId="7.[151,263,1840,1861]" captionTargetBox="[151,1436,193,1812]" captionTargetId="figure@7.[151,1436,193,1818]" captionTargetPageId="7" captionText="FIGURES 30 32. Atrovirensis spp., male genitalia. 30. A. furcatus sp. n., HT, China, Prov. Guangdong, genit. prep. HHL- 2921 - 1; 31. A. sp. cf. furcatus, China, Prov. Guangdong, genit. prep. HHL- 2918 - 1; 32. A. sacculatus sp. n., HT, China, Aut. Reg. Xizang, genit. prep. HHL- 2910 - 1." httpUri="https://zenodo.org/record/267010/files/figure.png" pageId="14" pageNumber="215">32</figureCitation>
,
<figureCitation id="134E2A05C058FF9F3D5B85F2FEACEBF4" box="[341,370,1733,1758]" captionStart="FIGURES 45 47" captionStartId="16.[151,263,1832,1853]" captionTargetBox="[151,1436,564,1810]" captionTargetId="figure@16.[151,1436,555,1810]" captionTargetPageId="16" captionText="FIGURES 45 47. Atrovirensis spp., female genitalia, habitats. 45. A. sacculatus sp. n., PT, China, Aut. Reg. Xizang, genit. prep. HHL- 2947 - 2; 46. A. yoshimotoi, PT, China, Prov. Guangdong, NSMT-SW- 021 (after Gyulai et al., 2013); 47. A. miraculosus sp. n., PT, China, Prov. Fujian, genit. prep. HHL- 2914 - 2; 48. A. euplexina, China, Prov. Sichuan, genit. prep. GB 12189; 49. Montane monsoon evergreen forest at elevation 2200 m—the habitats of Atrovirensis spp. in Linzhi, Hanmi- Motuo, Aut. Reg. Xizang, China. 50. Montane evergreen mixed forest at elevation 1500 m the habitat of Atrovirensis spp. in Mt. Nanling (Nanling Nat. Park.), Prov. Guangdong, China." httpUri="https://zenodo.org/record/267013/files/figure.png" pageId="14" pageNumber="215">45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F650BC058FF9F3C998439FB72EA44" pageId="14" pageNumber="215" type="materials_examined">
<paragraph id="8BCA3680C058FF9F3C998439FB72EA44" blockId="14.[151,1436,1805,2010]" pageId="14" pageNumber="215">
<emphasis id="B901EA92C058FF9F3C998439FEE0EA0C" bold="true" box="[151,318,1805,1830]" pageId="14" pageNumber="215">Type material</emphasis>
.
<emphasis id="B901EA92C058FF9F3D428439FE1CEA0F" bold="true" box="[332,450,1806,1829]" pageId="14" pageNumber="215">Holotype:</emphasis>
male, China, Aut. Reg. Xizang, Linzhi, Hanmi, 2200 m, 2028.vii.2013, Z.H. Pan leg., genitalia prep. HHL-2910-1, Coll. NEFU, Harbin.
<emphasis id="B901EA92C058FF9F3EDF8405FC8CEA63" bold="true" box="[721,850,1842,1865]" pageId="14" pageNumber="215">Paratypes:</emphasis>
2 males, 1 female, same locality, data and collector, genitalia prep. males, HHL-2908-1, HHL-2931-1, female HHL-2947-2. Coll. NEFU, Harbin.
</paragraph>
</subSubSection>
<subSubSection id="C36F650BC058FF9E3CC9844EFE69ECA2" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215" type="diagnosis">
<paragraph id="8BCA3680C058FF9E3CC9844EFE69ECA2" blockId="14.[151,1436,1805,2010]" lastBlockId="15.[151,1436,151,1256]" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
<emphasis id="B901EA92C058FF9F3CC9844EFE9FEAB8" bold="true" box="[199,321,1913,1938]" pageId="14" pageNumber="215">Diagnosis.</emphasis>
Wingspan 3235 mm.
<taxonomicName id="4C754D03C058FF9F3E4D844DFD06EAB8" box="[579,728,1913,1938]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="sacculatus" status="sp. nov.">
<emphasis id="B901EA92C058FF9F3E4D844DFD06EAB8" box="[579,728,1913,1938]" italics="true" pageId="14" pageNumber="215">A. sacculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257E9C058FF9F3ED0844DFCC3EABB" box="[734,797,1914,1937]" pageId="14" pageNumber="215" rank="species">
<emphasis id="B901EA92C058FF9F3ED0844DFCC3EABB" bold="true" box="[734,797,1914,1937]" pageId="14" pageNumber="215">sp. n.</emphasis>
</taxonomicNameLabel>
is the third new species of
<taxonomicName id="4C754D03C058FF9F3844844DFB53EABB" box="[1098,1165,1914,1937]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B901EA92C058FF9F3844844DFB53EABB" box="[1098,1165,1914,1937]" italics="true" pageId="14" pageNumber="215">sinica</emphasis>
</taxonomicName>
species-group. It differs externally from its congeners by smaller size (wingspan 3235 mm), bright colouration forewing, resembling
<taxonomicName id="4C754D03C058FF9F398A84A9FEF6EAF3" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="parannamita">
<emphasis id="B901EA92C058FF9F398A84A9FEF6EAF3" italics="true" pageId="14" pageNumber="215">A. parannamita</emphasis>
</taxonomicName>
. Wing pattern formed by golden-green elements on dark-brownish background and especially expressed golden-green subterminal field; hindwing yellowish-grey brown, darker than in
<taxonomicName id="4C754D03C059FF9E38BD83AFFAB8ED85" box="[1203,1382,152,176]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="parannamita">
<emphasis id="B901EA92C059FF9E38BD83AFFAB8ED85" box="[1203,1382,152,176]" italics="true" pageId="15" pageNumber="216">A. parannamita</emphasis>
</taxonomicName>
, but paler than in other members of
<taxonomicName id="4C754D03C059FF9E3E07838AFD92EDFE" box="[521,588,189,212]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B901EA92C059FF9E3E07838AFD92EDFE" box="[521,588,189,212]" italics="true" pageId="15" pageNumber="216">sinica</emphasis>
</taxonomicName>
species-group. By male genitalia structure it is close to
<taxonomicName id="4C754D03C059FF9E38D1838BFA8AEDFF" box="[1247,1364,188,213]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="owadai">
<emphasis id="B901EA92C059FF9E38D1838BFA8AEDFF" box="[1247,1364,188,213]" italics="true" pageId="15" pageNumber="216">A. owadai</emphasis>
</taxonomicName>
, from which it differs by narrower and more apically hooked uncus, broader juxta, more rounded cucullus, shorter neck of cucullus and angled lateral extension of sacculus (rounded in
<taxonomicName id="4C754D03C059FF9E3F878233FC20EC37" box="[905,1022,260,285]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="owadai">
<emphasis id="B901EA92C059FF9E3F878233FC20EC37" box="[905,1022,260,285]" italics="true" pageId="15" pageNumber="216">A. owadai</emphasis>
</taxonomicName>
and in all species of
<taxonomicName id="4C754D03C059FF9E38F18232FA9CEC36" box="[1279,1346,261,284]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B901EA92C059FF9E38F18232FA9CEC36" box="[1279,1346,261,284]" italics="true" pageId="15" pageNumber="216">sinica</emphasis>
</taxonomicName>
-species group). Aedeagus with rather short diverticula, medial diverticulum armed by short cutter-like cornutus, different in shape from other species of the group. Female differs by well-formed quadrangular antrum and very short sclerotised ductus bursae.
</paragraph>
</subSubSection>
<subSubSection id="C36F650BC059FF9E3CC982A3FE61E956" pageId="15" pageNumber="216" type="description">
<paragraph id="8BCA3680C059FF9E3CC982A3FB2FEE3F" blockId="15.[151,1436,151,1256]" pageId="15" pageNumber="216">
<emphasis id="B901EA92C059FF9E3CC982A3FE77EC87" bold="true" box="[199,425,404,429]" pageId="15" pageNumber="216">Description. Adult</emphasis>
(
<figureCitation id="134E2A05C059FF9E3DB182A3FD95EC87" box="[447,587,404,429]" captionStart="FIGURES 1 15" captionStartId="3.[151,263,1738,1759]" captionTargetBox="[190,1394,195,1718]" captionTargetId="figure@3.[190,1395,195,1719]" captionTargetPageId="3" captionText="FIGURES 1 15. Atrovirensis spp., adults. 1. A. atrovirens, Nepal (after Gyulai et al. 2013); 2. A. annamita, HT, Vietnam (after Gyulai et al. 2013); 3. A. parannamita sp. n., male HT, China, Aut. Reg. Xizang, genit. prep. HHL- 2930 - 1; 4. A. parannamita sp. n., female PT, China, Aut. Reg. Xizang, genit. prep. HHL- 2906 - 2; 5. A. pedibus, HT, China, Prov. Hubei (after Gyulai et al. 2013); 6. A. sp. cf. A. parannamita, male, China, Aut. Reg. Xizang, genit. prep. HHL- 2907 - 1; 7. A. sinica, HT, China, Prov. Guangdong, genit. prep. ZFMK 1624; 8. A. owadai, Prov. Guangdong, HT, China (after Gyulai et al. 2013); 9. A. taiwani, HT, Taiwan (after Gyulai et al. 2013); 10. A. furcatus sp. n., male HT, China, Prov. Guangdong, genit. prep. HHL- 2921 - 1; 11. A. furcatus sp. n., female PT, China, Prov. Guangdong, genit. prep. HHL- 2919 - 2; 12. A. sp. cf. furcatus, male, China, Prov. Guangdong, genit. prep. HHL- 2918 - 1; 13. A. sp. cf. furcatus, female, China, Prov. Guangdong, genit. prep. HHL- 2909 - 2; 14. A. sacculatus sp. n., male HT, China, Aut. Reg. Xizang, genit. prep. HHL- 2910 - 1; 15. A. sacculatus sp. n., male PT, China, Aut. Reg. Xizang, genit. prep. HHL- 2908 - 1." httpUri="https://zenodo.org/record/267006/files/figure.png" pageId="15" pageNumber="216">Figs 14, 15</figureCitation>
). Head and thorax covered with yellowish-green hair-like scales, palps blackish-brown. Ground colour of forewing blackish-brown with yellowish-green dusting; main elements of wing pattern yellowish-green; costal field yellowish-green dusting and with distinct yellowish-green and blackish marks; basal field yellowish-green, basal line distinct; subbasal field filled with yellowish-green, subbasal line slightly waved; medial field blackish-brown with yellow-greenish suffusion, claviform traceable; orbicular and reniform stigmata filled with yellowish-green, rather small, orbicular rounded, both stigmata encircled by dark line; veins suffused with grey; postmedial line waved; subterminal field bright, yellowish-green, with ash-grey veins and light brownish outer suffusion; subterminal line dentate, extended in M1-M2 veins area, pale yellowish-green; terminal field blackish-brown with grey suffusion; terminal line waved, as row black semilunar streaks; cilia yellowishbrown, with black terminal spots between veins. Hindwing yellowish-grey, paler in outer part, with hardly traceable discal spot and in some specimens with diffused terminal band; cilia pale, yellowish-grey.
</paragraph>
<paragraph id="8BCA3680C059FF9E3CC98017FDC5EEC6" blockId="15.[151,1436,151,1256]" pageId="15" pageNumber="216">
<emphasis id="B901EA92C059FF9E3CC98017FEA6EE12" bold="true" box="[199,376,799,824]" pageId="15" pageNumber="216">Male genitalia.</emphasis>
(
<figureCitation id="134E2A05C059FF9E3D898028FE07EE12" box="[391,473,799,824]" captionStart="FIGURES 30 32" captionStartId="7.[151,263,1840,1861]" captionTargetBox="[151,1436,193,1812]" captionTargetId="figure@7.[151,1436,193,1818]" captionTargetPageId="7" captionText="FIGURES 30 32. Atrovirensis spp., male genitalia. 30. A. furcatus sp. n., HT, China, Prov. Guangdong, genit. prep. HHL- 2921 - 1; 31. A. sp. cf. furcatus, China, Prov. Guangdong, genit. prep. HHL- 2918 - 1; 32. A. sacculatus sp. n., HT, China, Aut. Reg. Xizang, genit. prep. HHL- 2910 - 1." httpUri="https://zenodo.org/record/267010/files/figure.png" pageId="15" pageNumber="216">Fig. 32</figureCitation>
). Uncus rather short, thinner than in related species, slightly hooked apically; tegumen with broad peniculus, juxta plate-like, broad; valva narrower than in
<taxonomicName id="4C754D03C059FF9E3FAC8073FBCAEE77" box="[930,1044,836,861]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="owadai">
<emphasis id="B901EA92C059FF9E3FAC8073FBCAEE77" box="[930,1044,836,861]" italics="true" pageId="15" pageNumber="216">A. owadai</emphasis>
</taxonomicName>
, with shorter neck of cucullus and angled lateral extension of sacculus; harpe straight, rather short, smaller than in related species; cucullus larger and more rounded than
<emphasis id="B901EA92C059FF9E3D7E80BAFE23EE8F" box="[368,509,908,933]" italics="true" pageId="15" pageNumber="216">
in
<taxonomicName id="4C754D03C059FF9E3D8280BBFE23EE8F" box="[396,509,908,933]" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="owadai">A. owadai</taxonomicName>
</emphasis>
. Aedeagus with tubular vesica ended by short ventral diverticulum armed with two small spines and larger medial diverticulum armed with short (about two times shorter than in other species of
<taxonomicName id="4C754D03C059FF9E398A8087FF04EEC6" class="Insecta" family="Noctuidae" genus="Atrovirensis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B901EA92C059FF9E398A8087FF04EEC6" italics="true" pageId="15" pageNumber="216">A. sinica</emphasis>
</taxonomicName>
group), cutter-like cornutus.
</paragraph>
<paragraph id="8BCA3680C059FF9E3CC980CFFE61E956" blockId="15.[151,1436,151,1256]" pageId="15" pageNumber="216">
<emphasis id="B901EA92C059FF9E3CC980CFFE43E93A" bold="true" box="[199,413,1015,1040]" pageId="15" pageNumber="216">Female genitalia.</emphasis>
(
<figureCitation id="134E2A05C059FF9E3DB980C0FDCAE93A" box="[439,532,1015,1040]" captionStart="FIGURES 45 47" captionStartId="16.[151,263,1832,1853]" captionTargetBox="[151,1436,564,1810]" captionTargetId="figure@16.[151,1436,555,1810]" captionTargetPageId="16" captionText="FIGURES 45 47. Atrovirensis spp., female genitalia, habitats. 45. A. sacculatus sp. n., PT, China, Aut. Reg. Xizang, genit. prep. HHL- 2947 - 2; 46. A. yoshimotoi, PT, China, Prov. Guangdong, NSMT-SW- 021 (after Gyulai et al., 2013); 47. A. miraculosus sp. n., PT, China, Prov. Fujian, genit. prep. HHL- 2914 - 2; 48. A. euplexina, China, Prov. Sichuan, genit. prep. GB 12189; 49. Montane monsoon evergreen forest at elevation 2200 m—the habitats of Atrovirensis spp. in Linzhi, Hanmi- Motuo, Aut. Reg. Xizang, China. 50. Montane evergreen mixed forest at elevation 1500 m the habitat of Atrovirensis spp. in Mt. Nanling (Nanling Nat. Park.), Prov. Guangdong, China." httpUri="https://zenodo.org/record/267013/files/figure.png" pageId="15" pageNumber="216">Fig. 45</figureCitation>
). Ovipositor relatively small, papillae anales constricted apically; antrum quadrangular in shape with antevaginal plate formed by two small scobinate lobes; the ductus bursae very short, sclerotised and bifurcate in joining with corpus bursae; corpus bursae long, narrow, somewhat constricted medially, extended in proximal part.
</paragraph>
</subSubSection>
<subSubSection id="C36F650BC059FF9E3CC987BFFF31E9EF" pageId="15" pageNumber="216" type="etymology">
<paragraph id="8BCA3680C059FF9E3CC987BFFF31E9EF" blockId="15.[151,1436,151,1256]" pageId="15" pageNumber="216">
<emphasis id="B901EA92C059FF9E3CC987BFFE90E9B5" bold="true" box="[199,334,1160,1183]" pageId="15" pageNumber="216">Etymology.</emphasis>
The species name indicates for angular shape of sacculus which is the characteristic feature of new species.
</paragraph>
</subSubSection>
<subSubSection id="C36F650BC059FF9E3CC987F8FB66E9C2" box="[199,1208,1231,1256]" pageId="15" pageNumber="216" type="distribution">
<paragraph id="8BCA3680C059FF9E3CC987F8FB66E9C2" blockId="15.[151,1436,151,1256]" box="[199,1208,1231,1256]" pageId="15" pageNumber="216">
<emphasis id="B901EA92C059FF9E3CC987F8FEBEE9C2" bold="true" box="[199,352,1231,1256]" pageId="15" pageNumber="216">Distribution.</emphasis>
China (Aut. Reg. Xizang). The species is known only from its type-locality.
</paragraph>
</subSubSection>
</treatment>
</document>