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<document id="C01D93B0E5DE26024AB628D35D32B5C3" ID-DOI="10.1111/zoj.12305" ID-ISSN="0024-4082" ID-Zenodo-Dep="4721113" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619443797950" checkinUser="carolina" docAuthor="Bourguignon, Thomas, Scheffrahn, Rudolf H., Nagy, Zoltán Tamás, Sonet, Gontran, Host, Benoît &amp; Roisin, Yves" docDate="2016" docId="03B687D2F5225C0F50B1855EA7DB7B15" docLanguage="en" docName="ZoolJLinnSoc.176.15-35.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 176 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12305" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Grigiotermes Mathews 1977" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="FF8FFFAAF5215C0B53398404A52E7855" masterDocTitle="Towards a revision of the Neotropical soldierless termites (Isoptera: Termitidae): redescription of the genus Grigiotermes Mathews and description of five new genera" masterLastPageNumber="35" masterPageNumber="15" pageNumber="18" updateTime="1698910514992" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="376784CCDA1863413C6A77C3422AC5E9">Towards a revision of the Neotropical soldierless termites (Isoptera: Termitidae): redescription of the genus Grigiotermes Mathews and description of five new genera</mods:title>
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<mods:namePart id="FDF01C7D034918017AE241B82E3558F4">Bourguignon, Thomas</mods:namePart>
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<mods:namePart id="ABC6FCB005FB4DABB6DE79347DA2E274">Scheffrahn, Rudolf H.</mods:namePart>
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<mods:namePart id="C0B2A558A8592457FEA11C5810FC2947">Nagy, Zoltán Tamás</mods:namePart>
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<mods:namePart id="C6DC401127084C143D482CE664593262">Sonet, Gontran</mods:namePart>
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<mods:namePart id="D1557CDF7FC32472D7F35721A98D8785">Host, Benoît</mods:namePart>
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<mods:namePart id="07A3F7EC10EF0D1B27F1B1B0876ABEE6">Roisin, Yves</mods:namePart>
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<mods:title id="BE45A8530E3D675D51EA167FFDF5C3ED">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="D65554353D782CD67FB173331AF699E6">2016</mods:date>
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<mods:number id="9C45D7D94A3FC8C4FA0794A6EFA460DD">2016-01-31</mods:number>
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<mods:number id="0B4763222EDC13F9F44F727451667FD0">176</mods:number>
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<mods:number id="4DBA23B4D501DEA8551E7B43CF95898C">1</mods:number>
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<mods:start id="517BEC7BCF63CE35E2DD767C39F8E297">15</mods:start>
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<treatment id="03B687D2F5225C0F50B1855EA7DB7B15" LSID="urn:lsid:plazi:treatment:03B687D2F5225C0F50B1855EA7DB7B15" httpUri="http://treatment.plazi.org/id/03B687D2F5225C0F50B1855EA7DB7B15" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
<subSubSection id="C305654FF5225C0850B1855EA07D7925" box="[904,1363,346,368]" pageId="3" pageNumber="18" type="nomenclature">
<paragraph id="8BA036C4F5225C0850B1855EA07D7925" blockId="3.[826,1441,346,493]" box="[904,1363,346,368]" pageId="3" pageNumber="18">
<heading id="D0E881A8F5225C0850B1855EA07D7925" box="[904,1363,346,368]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="18" reason="2">
<treatmentCitationGroup id="AB0F11EAF5225C0850B1855EA07D7925" box="[904,1363,346,368]" pageId="3" pageNumber="18">
<treatmentCitation id="0ABE10D5F5225C0850B1855EA07D7925" author="Mathews AGA" box="[904,1363,346,368]" page="96" pageId="3" pageNumber="18" year="1977">
<taxonomicName id="4C1F4D47F5225C0850B1855EA07D7925" ID-CoL="8NS6F" authority="MATHEWS, 1977: 94 - 96" authorityName="Mathews" authorityPageNumber="94 - 96" authorityYear="1977" box="[904,1363,346,368]" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAD6F5225C0850B1855EA112793B" bold="true" box="[904,1084,346,367]" italics="true" pageId="3" pageNumber="18">GRIGIOTERMES</emphasis>
<bibRefCitation id="EF8E4B35F5225C08577D855EA07D7925" author="Mathews AGA" box="[1092,1363,346,368]" pageId="3" pageNumber="18" refId="ref12466" refString="Mathews AGA. 1977. Studies on termites from the Mato Grosso State, Brazil. Rio de Janeiro: Academia Brasileira de Ciencias." type="book" year="1977">MATHEWS, 1977: 94–96</bibRefCitation>
</taxonomicName>
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</heading>
</paragraph>
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<subSubSection id="C305654FF5225C0F50038579A4057AA9" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="8BA036C4F5225C0850038579A65679B8" blockId="3.[826,1441,346,493]" pageId="3" pageNumber="18">
<emphasis id="B96BEAD6F5225C0850038579A1D379C4" box="[826,1277,380,402]" italics="true" pageId="3" pageNumber="18">
<typeStatus id="54A48866F5225C0850038579A64179C7" box="[826,879,381,402]" pageId="3" pageNumber="18">Type</typeStatus>
species, by original designation:
</emphasis>
<taxonomicName id="4C1F4D47F5225C0856338579A17379E5" authority="Mathews, 1977" authorityName="Mathews" authorityYear="1977" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="metoecus">
<emphasis id="B96BEAD6F5225C0856338579A68E79E4" italics="true" pageId="3" pageNumber="18">Grigiotermes metoecus</emphasis>
<bibRefCitation id="EF8E4B35F5225C085093859FA17379E5" author="Mathews AGA" box="[938,1117,411,433]" pageId="3" pageNumber="18" refId="ref12466" refString="Mathews AGA. 1977. Studies on termites from the Mato Grosso State, Brazil. Rio de Janeiro: Academia Brasileira de Ciencias." type="book" year="1977">Mathews, 1977</bibRefCitation>
</taxonomicName>
, herein considered a junior synonym of
<taxonomicName id="4C1F4D47F5225C0850FF85BEA65A79B8" authority="Snyder &amp; Emerson, 1949" authorityName="Snyder &amp; Emerson" authorityYear="1949" class="Insecta" family="Termitidae" genus="Anoplotermes" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="hageni">
<emphasis id="B96BEAD6F5225C0850FF85BEA198799A" box="[966,1206,442,463]" italics="true" pageId="3" pageNumber="18">Anoplotermes hageni</emphasis>
Snyder &amp; Emerson, 1949
</taxonomicName>
.
</paragraph>
<paragraph id="8BA036C4F5225C0850038612A6957A7E" blockId="3.[826,1442,534,923]" box="[826,955,534,555]" pageId="3" pageNumber="18">
<emphasis id="B96BEAD6F5225C0850038612A6957A7E" box="[826,955,534,555]" italics="true" pageId="3" pageNumber="18">Description</emphasis>
</paragraph>
<paragraph id="8BA036C4F5225C0850038630A08C7BCF" blockId="3.[826,1442,534,923]" pageId="3" pageNumber="18">
<emphasis id="B96BEAD6F5225C0850038630A6BF7A1C" box="[826,913,564,585]" italics="true" pageId="3" pageNumber="18">Worker:</emphasis>
Large, head width over
<quantity id="4CE79B21F5225C0857A78630A1D87A1F" box="[1182,1270,564,586]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="18" unit="mm" value="0.9">0.9 mm</quantity>
. Abdomen with subconical distal half and elongated digestive tract. Head pale yellow, rounded, fully covered with various numbers of tiny setae and between 50 and 100 long bristles. Fontanelle small, almost indistinct. Fore tibia long and slender. Digestive tract as in
<figureCitation id="13242A41F5225C0857B086C9A1D07AB7" box="[1161,1278,717,739]" captionStart="Figure 8" captionStartId="14.[144,223,1445,1464]" captionTargetBox="[161,1413,203,1380]" captionTargetId="figure-157@14.[144,1424,195,1416]" captionTargetPageId="14" captionText="Figure 8. Worker digestive tracts. A, Grigiotermes hageni comb. nov.; B, Amplucrutermes inflatus sp. nov.; C, Humutermes krishnai sp. nov.; D, Hydrecotermes arienesho sp. nov.; E, Patawatermes nigripunctatus comb. nov.; F, Rubeotermes jheringi comb. nov.. From left to right: dorsal, right, ventral, and left views." figureDoi="http://doi.org/10.5281/zenodo.4721145" httpUri="https://zenodo.org/record/4721145/files/figure.png" pageId="3" pageNumber="18">Figure 8A</figureCitation>
. Enteric valve inserted into a short extension of the paunch, with welldeveloped armature composed of six spiny plates. Anterior portion of enteric valve consists of six ovoid pads composed of about 100 scale-like lobes. Mesenteric junction forming a small ball overlapping the proctodeum, slightly elongated to form a short, distinct mixed segment.
</paragraph>
<caption id="DF60664CF5225C08539D81BBA01C7D87" ID-DOI="http://doi.org/10.5281/zenodo.4721117" ID-Zenodo-Dep="4721117" box="[164,1330,1471,1491]" httpUri="https://zenodo.org/record/4721117/files/figure.png" pageId="3" pageNumber="18" startId="3.[164,243,1471,1490]" targetBox="[164,1444,1011,1441]" targetPageId="3">
<paragraph id="8BA036C4F5225C08539D81BBA01C7D87" blockId="3.[164,1442,1471,1588]" box="[164,1330,1471,1491]" pageId="3" pageNumber="18">
<emphasis id="B96BEAD6F5225C08539D81BBA4227D87" bold="true" box="[164,268,1471,1490]" pageId="3" pageNumber="18">Figure 1.</emphasis>
Worker.
<emphasis id="B96BEAD6F5225C08524B81BBA7C77D87" bold="true" box="[370,745,1471,1490]" pageId="3" pageNumber="18">
<taxonomicName id="4C1F4D47F5225C08524B81BBA74C7D87" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" baseAuthorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" baseAuthorityYear="2016" box="[370,610,1471,1490]" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="hageni" status="comb. nov.">
<emphasis id="B96BEAD6F5225C08524B81BBA74C7D87" bold="true" box="[370,610,1471,1490]" italics="true" pageId="3" pageNumber="18">Grigiotermes hageni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857ADF5225C08515581BBA7C77D87" box="[620,745,1471,1490]" pageId="3" pageNumber="18" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
A, enteric valve (spread on slide); B, foreleg; C, head.
</paragraph>
</caption>
<paragraph id="8BA036C4F5225C08539D8200A6447E61" blockId="3.[164,1442,1471,1588]" pageId="3" pageNumber="18">
<emphasis id="B96BEAD6F5225C08539D8200A5D37E42" bold="true" box="[164,253,1540,1559]" pageId="3" pageNumber="18">Table 2.</emphasis>
Measurements (in mm) of
<specimenCount id="9D19FD4DF5225C0851198200A7B97E42" box="[544,663,1540,1559]" pageId="3" pageNumber="18" type="worker">24 workers</specimenCount>
, and
<specimenCount id="9D19FD4DF5225C0851E88200A61F7E42" box="[721,817,1540,1559]" pageId="3" pageNumber="18" type="male">five male</specimenCount>
and
<specimenCount id="9D19FD4DF5225C08505E8200A6F47E42" box="[871,986,1540,1559]" pageId="3" pageNumber="18" type="female">five female</specimenCount>
imagos from five colonies of
<emphasis id="B96BEAD6F5225C0856328200A4557E61" bold="true" pageId="3" pageNumber="18">
<taxonomicName id="4C1F4D47F5225C0856328200A5DA7E61" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" baseAuthorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" baseAuthorityYear="2016" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="hageni" status="comb. nov.">
<emphasis id="B96BEAD6F5225C0856328200A5DA7E61" bold="true" italics="true" pageId="3" pageNumber="18">Grigiotermes hageni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857ADF5225C0853C48225A4557E61" box="[253,379,1569,1588]" pageId="3" pageNumber="18" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="4C1F4D47F5225C0852B28225A7597E61" authorityName="Mathews" authorityYear="1977" box="[395,631,1569,1588]" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="metoecus">
<emphasis id="B96BEAD6F5225C0852B28225A7597E61" box="[395,631,1569,1588]" italics="true" pageId="3" pageNumber="18">Grigiotermes metoecus</emphasis>
</taxonomicName>
, from
<bibRefCitation id="EF8E4B35F5225C0851F88225A64D7E61" author="Mathews AGA" box="[705,867,1569,1588]" pageId="3" pageNumber="18" refId="ref12466" refString="Mathews AGA. 1977. Studies on termites from the Mato Grosso State, Brazil. Rio de Janeiro: Academia Brasileira de Ciencias." type="book" year="1977">Mathews, 1977</bibRefCitation>
)
</paragraph>
<paragraph id="8BA036C4F5225C08518A825FA0677F0D" pageId="3" pageNumber="18">
<table id="F91FC464F522A3F4539D825FA08F7F0D" box="[164,1441,1627,1880]" colsContinueIn="12.[144,1421,1627,1880]" gridcols="4" gridrows="8" pageId="3" pageNumber="18">
<tr id="352F3486F522A3F4539D825FA08F7E3B" box="[164,1441,1627,1646]" gridrow="0" pageId="3" pageNumber="18" rowspan-0="1">
<th id="76FE5DFAF522A3F4518A825FA61E7E3B" box="[691,816,1627,1646]" gridcol="1" gridrow="0" pageId="3" pageNumber="18">Worker</th>
<th id="76FE5DFAF522A3F4573D825FA1477E3B" box="[1028,1129,1627,1646]" gridcol="2" gridrow="0" pageId="3" pageNumber="18">Imago ♂</th>
<th id="76FE5DFAF522A3F45605825FA08F7E3B" box="[1340,1441,1627,1646]" gridcol="3" gridrow="0" pageId="3" pageNumber="18">Imago ♀</th>
</tr>
<tr id="352F3486F522A3F4539D8291A08F7EFD" box="[164,1441,1685,1704]" gridrow="1" pageId="3" pageNumber="18">
<th id="76FE5DFAF522A3F4539D8291A4F17EFD" box="[164,479,1685,1704]" gridcol="0" gridrow="1" pageId="3" pageNumber="18">Head length to end of clypeus</th>
<td id="76FE5DFAF522A3F4518A8291A61E7EFD" box="[691,816,1685,1704]" gridcol="1" gridrow="1" pageId="3" pageNumber="18">0.91–1.11</td>
<td id="76FE5DFAF522A3F4573D8291A1477EFD" box="[1028,1129,1685,1704]" gridcol="2" gridrow="1" pageId="3" pageNumber="18">–</td>
<td id="76FE5DFAF522A3F456058291A08F7EFD" box="[1340,1441,1685,1704]" gridcol="3" gridrow="1" pageId="3" pageNumber="18">–</td>
</tr>
<tr id="352F3486F522A3F4539D82B6A08F7E90" box="[164,1441,1714,1733]" gridrow="2" pageId="3" pageNumber="18">
<th id="76FE5DFAF522A3F4539D82B6A4F17E90" box="[164,479,1714,1733]" gridcol="0" gridrow="2" pageId="3" pageNumber="18">Head width</th>
<td id="76FE5DFAF522A3F4518A82B6A61E7E90" box="[691,816,1714,1733]" gridcol="1" gridrow="2" pageId="3" pageNumber="18">0.94–1.29</td>
<td id="76FE5DFAF522A3F4573D82B6A1477E90" box="[1028,1129,1714,1733]" gridcol="2" gridrow="2" pageId="3" pageNumber="18">1.45–1.48</td>
<td id="76FE5DFAF522A3F4560582B6A08F7E90" box="[1340,1441,1714,1733]" gridcol="3" gridrow="2" pageId="3" pageNumber="18">1.50–1.54</td>
</tr>
<tr id="352F3486F522A3F4539D82CBA08F7EB6" box="[164,1441,1743,1763]" gridrow="3" pageId="3" pageNumber="18">
<th id="76FE5DFAF522A3F4539D82CBA4F17EB6" box="[164,479,1743,1763]" gridcol="0" gridrow="3" pageId="3" pageNumber="18">Eye diameter</th>
<td id="76FE5DFAF522A3F4518A82CBA61E7EB6" box="[691,816,1743,1763]" gridcol="1" gridrow="3" pageId="3" pageNumber="18">–</td>
<td id="76FE5DFAF522A3F4573D82CBA1477EB6" box="[1028,1129,1743,1763]" gridcol="2" gridrow="3" pageId="3" pageNumber="18">0.34–0.36</td>
<td id="76FE5DFAF522A3F4560582CBA08F7EB6" box="[1340,1441,1743,1763]" gridcol="3" gridrow="3" pageId="3" pageNumber="18">0.36–0.37</td>
</tr>
<tr id="352F3486F522A3F4539D82E8A08F7F55" box="[164,1441,1772,1792]" gridrow="4" pageId="3" pageNumber="18">
<th id="76FE5DFAF522A3F4539D82E8A4F17F55" box="[164,479,1772,1792]" gridcol="0" gridrow="4" pageId="3" pageNumber="18">Pronotum length</th>
<td id="76FE5DFAF522A3F4518A82E8A61E7F55" box="[691,816,1772,1792]" gridcol="1" gridrow="4" pageId="3" pageNumber="18">–</td>
<td id="76FE5DFAF522A3F4573D82E8A1477F55" box="[1028,1129,1772,1792]" gridcol="2" gridrow="4" pageId="3" pageNumber="18">0.69–0.71</td>
<td id="76FE5DFAF522A3F4560582E8A08F7F55" box="[1340,1441,1772,1792]" gridcol="3" gridrow="4" pageId="3" pageNumber="18">0.74–0.80</td>
</tr>
<tr id="352F3486F522A3F4539D830EA08F7F48" box="[164,1441,1802,1821]" gridrow="5" pageId="3" pageNumber="18">
<th id="76FE5DFAF522A3F4539D830EA4F17F48" box="[164,479,1802,1821]" gridcol="0" gridrow="5" pageId="3" pageNumber="18">Pronotum width</th>
<td id="76FE5DFAF522A3F4518A830EA61E7F48" box="[691,816,1802,1821]" gridcol="1" gridrow="5" pageId="3" pageNumber="18">0.60–0.91</td>
<td id="76FE5DFAF522A3F4573D830EA1477F48" box="[1028,1129,1802,1821]" gridcol="2" gridrow="5" pageId="3" pageNumber="18">1.27–1.30</td>
<td id="76FE5DFAF522A3F45605830EA08F7F48" box="[1340,1441,1802,1821]" gridcol="3" gridrow="5" pageId="3" pageNumber="18">1.36–1.51</td>
</tr>
<tr id="352F3486F522A3F4539D8323A08F7F6F" box="[164,1441,1831,1850]" gridrow="6" pageId="3" pageNumber="18">
<th id="76FE5DFAF522A3F4539D8323A4F17F6F" box="[164,479,1831,1850]" gridcol="0" gridrow="6" pageId="3" pageNumber="18">Hind tibia length</th>
<td id="76FE5DFAF522A3F4518A8323A61E7F6F" box="[691,816,1831,1850]" gridcol="1" gridrow="6" pageId="3" pageNumber="18">0.82–1.13</td>
<td id="76FE5DFAF522A3F4573D8323A1477F6F" box="[1028,1129,1831,1850]" gridcol="2" gridrow="6" pageId="3" pageNumber="18">1.48–1.51</td>
<td id="76FE5DFAF522A3F456058323A08F7F6F" box="[1340,1441,1831,1850]" gridcol="3" gridrow="6" pageId="3" pageNumber="18">1.48–1.54</td>
</tr>
<tr id="352F3486F522A3F4539D8340A08F7F0D" box="[164,1441,1860,1880]" gridrow="7" pageId="3" pageNumber="18">
<th id="76FE5DFAF522A3F4539D8340A4F17F0D" box="[164,479,1860,1880]" gridcol="0" gridrow="7" pageId="3" pageNumber="18">Fore tibia width/length</th>
<td id="76FE5DFAF522A3F4518A8340A61E7F0D" box="[691,816,1860,1880]" gridcol="1" gridrow="7" pageId="3" pageNumber="18">0.244–0.408</td>
<td id="76FE5DFAF522A3F4573D8340A1477F0D" box="[1028,1129,1860,1880]" gridcol="2" gridrow="7" pageId="3" pageNumber="18">–</td>
<td id="76FE5DFAF522A3F456058340A08F7F0D" box="[1340,1441,1860,1880]" gridcol="3" gridrow="7" pageId="3" pageNumber="18">–</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA036C4F5255C0F53A984C1A5FF794D" blockId="4.[144,759,197,494]" pageId="4" pageNumber="19">Foregut with very wide P1 (first protocol segment) through which food bolus is generally visible as a serpentine mass.</paragraph>
<paragraph id="8BA036C4F5255C0F53918525A7C379BB" blockId="4.[144,759,197,494]" pageId="4" pageNumber="19">
<emphasis id="B96BEAD6F5255C0F53918525A5D97963" box="[168,247,289,310]" italics="true" pageId="4" pageNumber="19">Imago:</emphasis>
Sclerotized body parts dark brown, covered by thousands of tiny setae. Head covered by about 100 scattered long bristles. Eyes and ocelli large. Fontanelle fairly small to medium size, situated at the level of eyes. Pronotum elongated, covered by dozens of long bristles. Posterior margins of meso- and metanotum slightly emarginated, with strongly rounded angles.
</paragraph>
<paragraph id="8BA036C4F5255C0F53A98614A42E7A70" blockId="4.[144,759,528,764]" box="[144,256,528,549]" pageId="4" pageNumber="19">
<emphasis id="B96BEAD6F5255C0F53A98614A42E7A70" box="[144,256,528,549]" italics="true" pageId="4" pageNumber="19">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BA036C4F5255C0F53A9862AA4057AA9" blockId="4.[144,759,528,764]" pageId="4" pageNumber="19">
<taxonomicName id="4C1F4D47F5255C0F53A9862AA40C7A16" authorityName="Mathews" authorityYear="1977" box="[144,290,558,579]" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAD6F5255C0F53A9862AA40C7A16" box="[144,290,558,579]" italics="true" pageId="4" pageNumber="19">Grigiotermes</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C1F4D47F5255C0F52FE862BA7467A11" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[455,616,559,580]" class="Insecta" family="Termitidae" genus="Patawatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAD6F5255C0F52FE862BA7467A11" box="[455,616,559,580]" italics="true" pageId="4" pageNumber="19">Patawatermes</emphasis>
</taxonomicName>
from which it can be distinguished by its enteric valve adorned with six spiny sclerotized plates as in
<figureCitation id="13242A41F5255C0F517C866FA7EA7AD5" box="[581,708,619,641]" captionStart="Figure 1" captionStartId="3.[164,243,1471,1490]" captionTargetBox="[164,1444,1011,1441]" captionTargetId="figure-360@3.[164,1444,1011,1441]" captionTargetPageId="3" captionText="Figure 1. Worker. Grigiotermes hageni comb. nov. A, enteric valve (spread on slide); B, foreleg; C, head." figureDoi="http://doi.org/10.5281/zenodo.4721117" httpUri="https://zenodo.org/record/4721117/files/figure.png" pageId="4" pageNumber="19">Figures 1A</figureCitation>
,
<figureCitation id="13242A41F5255C0F51E9866FA7DD7AD5" box="[720,755,619,640]" captionStart="Figure 7" captionStartId="13.[164,243,1171,1190]" captionTargetBox="[323,1283,194,1141]" captionTargetId="figure-210@13.[323,1283,194,1141]" captionTargetPageId="13" captionText="Figure 7. Worker enteric valve armature, depicted by confocal microscopy. A, Grigiotermes hageni comb. nov.; B, Amplucrutermes inflatus sp. nov.; C, Humutermes krishnai sp. nov.; D, Humutermes noiroti sp. nov.; E, Hydrecotermes arienesho sp. nov.; F, Hydrecotermes kawaii sp. nov.; G, Patawatermes turricola comb. nov.; H, Patawatermes nigripunctatus comb. nov.; I, Rubeotermes jheringi comb. nov. (G, I, reproduced with the permission of Host et al., 2014)." figureDoi="http://doi.org/10.5281/zenodo.4721141" httpUri="https://zenodo.org/record/4721141/files/figure.png" pageId="4" pageNumber="19">7A</figureCitation>
.
<taxonomicName id="4C1F4D47F5255C0F53A9868EA40C7ACA" authorityName="Mathews" authorityYear="1977" box="[144,290,650,671]" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAD6F5255C0F53A9868EA40C7ACA" box="[144,290,650,671]" italics="true" pageId="4" pageNumber="19">Grigiotermes</emphasis>
</taxonomicName>
is distinguishable from other genera of South American soldierless termites by its workers with widened P
<quantity id="4CE79B21F5255C0F523386C3A41A7A88" box="[266,308,711,733]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="19" unit="in" value="1.0">1 in</quantity>
which the food bolus has a visible serpentine path.
</paragraph>
</subSubSection>
<subSubSection id="C305654FF5255C0F53A8872EA7DB7B15" box="[145,757,810,832]" pageId="4" pageNumber="19" type="description">
<paragraph id="8BA036C4F5255C0F53A8872EA7DB7B15" blockId="4.[145,757,810,896]" box="[145,757,810,832]" pageId="4" pageNumber="19">
<taxonomicName id="4C1F4D47F5255C0F53A8872EA7DB7B15" authority="HAGENI (SNYDER &amp; EMERSON, 1949)" authorityName="HAGENI (SNYDER &amp; EMERSON" authorityYear="1949" box="[145,757,810,832]" class="Insecta" family="Termitidae" genus="Grigiotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAD6F5255C0F53A8872EA4887B6A" bold="true" box="[145,422,810,831]" italics="true" pageId="4" pageNumber="19">GRIGIOTERMES HAGENI</emphasis>
(SNYDER &amp; EMERSON, 1949)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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