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<document id="9DF29FC3D8F59500062757FDEFB7E8C2" ID-CLB-Dataset="21135" ID-DOI="10.1080/00222933.2015.1021875" ID-GBIF-Dataset="b8546f1e-9186-421c-a535-5342fd7fe0d4" ID-ISSN="1464-5262" ID-Zenodo-Dep="4000011" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598369669509" checkinUser="carolina" docAuthor="Aspe, Nonillon M. &amp; James, Samuel W." docDate="2015" docId="03A8E367FFAFFFAAFE0A58DAFE28FB8E" docLanguage="en" docName="JNATHIST.49.37-38.2233-2255.pdf" docOrigin="Journal of Natural History 49 (37)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Polypheretima mindanaoensis Aspe &amp; James 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="2243" masterDocId="FF919B1FFFAAFFA0FF905C5EFFD5FFDC" masterDocTitle="New Polypheretima and Pithemera (Oligochaeta: Megascolecidae) species from the Mt. Malindang Range, Mindanao Island, Philippines" masterLastPageNumber="2255" masterPageNumber="2233" pageNumber="2238" updateTime="1698845390463" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="D6DB11216D77CE99812F3A6C2F328467" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D644C31EBA14E0756D01B6B117E2D14D">
<mods:title id="1F135DC295242C781906A6A7923F8C4A">New Polypheretima and Pithemera (Oligochaeta: Megascolecidae) species from the Mt. Malindang Range, Mindanao Island, Philippines</mods:title>
</mods:titleInfo>
<mods:name id="DEA56FFB5F060715229EACE1542B015F" type="personal">
<mods:role id="3567FC839022C8B19F5796B33B3FD1F0">
<mods:roleTerm id="AB83A5186A1FB531E81AAC9B5ADA656D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0C375CB932C0C5511E7B63F8B431F72E">Aspe, Nonillon M.</mods:namePart>
<mods:affiliation id="E99A1FF2D85D7D97C0949F255BD6185F">Department of Natural History Sciences, Graduate School of Science, Hokkaido University, Sapporo, Japan;</mods:affiliation>
</mods:name>
<mods:name id="B9D55080410DE6EC03176B580571A940" type="personal">
<mods:role id="D62AF7BECECC6D38347DF5F88BA9C11B">
<mods:roleTerm id="A33761AED44EE0861F8145330F6BB8AF">Author</mods:roleTerm>
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<mods:namePart id="D1613894CF4415913E531A10B71C8EFF">James, Samuel W.</mods:namePart>
<mods:affiliation id="F35787693E3EAABE931B444BEB5B2CAF">Department of Biology, College of Liberal Arts and Sciences, University</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9FFCCFAFD17646DD25D79AA72C5F9E6F">text</mods:typeOfResource>
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<mods:title id="1F2DBDA0E3D551788E53D2E2B82E4DAC">Journal of Natural History</mods:title>
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<mods:part id="FD2BA52603ADF2493F9D49802A344FDD">
<mods:date id="F75E1E31C04BDD8B1363D67405817E4C">2015</mods:date>
<mods:detail id="D35FF6435272A2E166D4C5617AD178B2" type="pubDate">
<mods:number id="06F9D31F3C65A696C4E2657820B6322F">2015-03-31</mods:number>
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<mods:detail id="29DE63BAA0D89C116137622F1EBCA1EB" type="volume">
<mods:number id="9098B6523737FD2B99FADD5CB031BDF8">49</mods:number>
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<mods:number id="CBFF75F7465067188E00D814C4513B30">37</mods:number>
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<mods:classification id="1F7595D2EE1EB5CDFAEC7E0832E92C10">journal article</mods:classification>
<mods:identifier id="5682A903B74DA187193AAE4CBA7ECD66" type="CLB-Dataset">21135</mods:identifier>
<mods:identifier id="8C19DAA2DD049BF9472B7B66493DD5E5" type="DOI">10.1080/00222933.2015.1021875</mods:identifier>
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<treatment id="03A8E367FFAFFFAAFE0A58DAFE28FB8E" ID-DOI="http://doi.org/10.5281/zenodo.4329076" ID-GBIF-Taxon="167307926" ID-Zenodo-Dep="4329076" LSID="urn:lsid:plazi:treatment:03A8E367FFAFFFAAFE0A58DAFE28FB8E" httpUri="http://treatment.plazi.org/id/03A8E367FFAFFFAAFE0A58DAFE28FB8E" lastPageId="10" lastPageNumber="2243" pageId="5" pageNumber="2238">
<subSubSection id="C31B01FAFFAFFFA5FE0A58DAFC93FB40" box="[410,838,1156,1180]" pageId="5" pageNumber="2238" type="nomenclature">
<paragraph id="8BBE5271FFAFFFA5FE0A58DAFC93FB40" blockId="5.[410,838,1156,1213]" box="[410,838,1156,1180]" pageId="5" pageNumber="2238">
<heading id="D0F6E51DFFAFFFA5FE0A58DAFC93FB40" box="[410,838,1156,1180]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="2238" reason="2">
<taxonomicName id="4C0129F2FFAFFFA5FE0A58DAFD36FB40" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[410,739,1156,1180]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="2238" phylum="Annelida" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="B9758E63FFAFFFA5FE0A58DAFD36FB40" bold="true" box="[410,739,1156,1180]" italics="true" pageId="5" pageNumber="2238">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFAFFFA5FD7C58DBFC93FB40" box="[748,838,1157,1180]" pageId="5" pageNumber="2238" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B01FAFFAFFFA5FE6958FBFD32FB61" box="[505,743,1189,1213]" pageId="5" pageNumber="2238" type="description">
<paragraph id="8BBE5271FFAFFFA5FE6958FBFD32FB61" blockId="5.[410,838,1156,1213]" box="[505,743,1189,1213]" pageId="5" pageNumber="2238">
(
<figureCitation id="133A4EF4FFAFFFA5FD9158FBFD54FB61" box="[513,641,1189,1213]" captionStart="Figure 2" captionStartId="6.[212,279,862,884]" captionTargetBox="[212,1172,169,830]" captionTargetId="figure@6.[212,1173,168,830]" captionTargetPageId="6" captionText="Figure 2. Schematic drawings of the external anterio-ventral area of (A) Polypheretima mindanaoensis sp. nov., (B) Pithemera malindangensis sp. nov., (C) Pithemera duminagati sp. nov. and (D) Pithemera donvictorianoi sp. nov. Abbreviations: fp, female pore; cl, clitellum; mp, male pore; gm, genital markings. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4000015" httpUri="https://zenodo.org/record/4000015/files/figure.png" pageId="5" pageNumber="2238">Figures 2A</figureCitation>
and
<figureCitation id="133A4EF4FFAFFFA5FD2C58FBFD0BFB61" box="[700,734,1189,1213]" captionStart="Figure 3" captionStartId="7.[144,211,1576,1598]" captionTargetBox="[190,1058,168,1544]" captionTargetId="figure@7.[190,1058,168,1544]" captionTargetPageId="7" captionText="Figure 3. Schematic drawings of the internal morphology of (A) Polypheretima mindanaoensis sp. nov., (B) Pithemera malindangensis sp. nov., (C) Pithemera duminagati sp. nov. and (D) Pithemera donvictorianoi sp. nov. Abbreviations: s, spermatheca; n, dense tufts of nephridia; h, heart; p, prostate gland; c, caecum. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4000017" httpUri="https://zenodo.org/record/4000017/files/figure.png" pageId="5" pageNumber="2238">3A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B01FAFFAFFFA5FF0058A3FF3CFAA0" pageId="5" pageNumber="2238" type="diagnosis">
<paragraph id="8BBE5271FFAFFFA5FF0058A3FF29FAC8" blockId="5.[144,1104,1277,1404]" box="[144,252,1277,1300]" pageId="5" pageNumber="2238">
<heading id="D0F6E51DFFAFFFA5FF0058A3FF29FAC8" box="[144,252,1277,1300]" fontSize="10" level="3" pageId="5" pageNumber="2238" reason="5">
<emphasis id="B9758E63FFAFFFA5FF0058A3FF29FAC8" box="[144,252,1277,1300]" italics="true" pageId="5" pageNumber="2238">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE5271FFAFFFA5FF00597AFF3CFAA0" blockId="5.[144,1104,1277,1404]" pageId="5" pageNumber="2238">
Body white; stout, adult length
<quantity id="4CF9FF94FFAFFFA5FD92597AFD5EFAE0" box="[514,651,1316,1340]" metricMagnitude="-1" metricUnit="m" metricValue="1.04" metricValueMax="1.18" metricValueMin="0.9" pageId="5" pageNumber="2238" unit="mm" value="104.0" valueMax="118.0" valueMin="90.0">90118 mm</quantity>
; paired, sublateral genital markings on each of xix to xxvi; intestinal origin in xv; small spermathecae present in batteries or lacking.
</paragraph>
</subSubSection>
<subSubSection id="C31B01FAFFAFFFA5FF0059E2FDA8FA20" box="[144,637,1468,1532]" pageId="5" pageNumber="2238" type="etymology">
<paragraph id="8BBE5271FFAFFFA5FF0059E2FEDCFA08" blockId="5.[144,637,1468,1532]" box="[144,265,1468,1492]" pageId="5" pageNumber="2238">
<heading id="D0F6E51DFFAFFFA5FF0059E2FEDCFA08" box="[144,265,1468,1492]" fontSize="10" level="3" pageId="5" pageNumber="2238" reason="5">
<emphasis id="B9758E63FFAFFFA5FF0059E2FEDCFA08" box="[144,265,1468,1492]" italics="true" pageId="5" pageNumber="2238">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE5271FFAFFFA5FF0059BAFDA8FA20" blockId="5.[144,637,1468,1532]" box="[144,637,1508,1532]" pageId="5" pageNumber="2238">The species is named for Mindanao Island.</paragraph>
</subSubSection>
<subSubSection id="C31B01FAFFAFFFA6FF005A62FBF0FBAC" lastPageId="6" lastPageNumber="2239" pageId="5" pageNumber="2238" type="materials_examined">
<paragraph id="8BBE5271FFAFFFA5FF005A62FEB6F988" blockId="5.[144,1104,1596,1692]" box="[144,355,1596,1620]" pageId="5" pageNumber="2238">
<heading id="D0F6E51DFFAFFFA5FF005A62FEB6F988" box="[144,355,1596,1620]" fontSize="10" level="3" pageId="5" pageNumber="2238" reason="5">
<emphasis id="B9758E63FFAFFFA5FF005A62FEB6F988" box="[144,355,1596,1620]" italics="true" pageId="5" pageNumber="2238">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE5271FFAFFFA6FF005A3DFBF0FBAC" blockId="5.[144,1104,1596,1692]" lastBlockId="6.[212,1171,1049,1136]" lastPageId="6" lastPageNumber="2239" pageId="5" pageNumber="2238">
<materialsCitation id="3B69582CFFAFFFA6FF005A3DFD71FB8D" ID-GBIF-Occurrence="2848941304" collectingDate="2003-10-09" collectingDateMax="2003-10-15" collectingDateMin="2003-10-09" collectorName="Nonillon Aspe &amp; Nolan Aspe &amp; J. Adeva" country="Philippines" county="Mindanao Island" elevation="1357" lastPageId="6" lastPageNumber="2239" latitude="8.2986" location="Barangay Lake Duminagat" longLatPrecision="7" longitude="123.6169" municipality="Mt. Malindang Range" pageId="5" pageNumber="2238" specimenCount="1" stateProvince="Misamis Occidental" typeStatus="holotype">
<typeStatus id="54BAECD3FFAFFFA5FF005A3DFF2BF9A7" box="[144,254,1635,1659]" pageId="5" pageNumber="2238" type="holotype">Holotype</typeStatus>
: adult (NMA 4527),
<location id="8EDE04AAFFAFFFA5FD955A3AFC9FF9A7" LSID="urn:lsid:plazi:treatment:03A8E367FFAFFFAAFE0A58DAFE28FB8E:8EDE04AAFFAFFFA5FD955A3AFC9FF9A7" box="[517,842,1635,1660]" country="Philippines" county="Mindanao Island" latitude="8.2986" longLatPrecision="7" longitude="123.6169" municipality="Mt. Malindang Range" name="Barangay Lake Duminagat" pageId="5" pageNumber="2238" stateProvince="Misamis Occidental">
<location id="8EDE04AAFFAFFFA5FD955A3AFD6CF9A7" LSID="urn:lsid:plazi:treatment:03A8E367FFAFFFAAFE0A58DAFE28FB8E:8EDE04AAFFAFFFA5FD955A3AFD6CF9A7" box="[517,697,1635,1660]" country="Philippines" county="Mindanao Island" latitude="8.2986" longLatPrecision="7" longitude="123.6169" municipality="Mt. Malindang Range" name="Barangay Lake" pageId="5" pageNumber="2238" stateProvince="Misamis Occidental">
Barangay
<location id="8EDE04AAFFAFFFA5FD105A3DFD6CF9A7" LSID="urn:lsid:plazi:treatment:03A8E367FFAFFFAAFE0A58DAFE28FB8E:8EDE04AAFFAFFFA5FD105A3DFD6CF9A7" box="[640,697,1635,1659]" country="Philippines" county="Mindanao Island" latitude="8.2986" longLatPrecision="7" longitude="123.6169" municipality="Mt. Malindang Range" name="Lake Duminagat" pageId="5" pageNumber="2238" stateProvince="Misamis Occidental">Lake</location>
</location>
Duminagat
</location>
, municipality of
<location id="8EDE04AAFFAFFFA5FB8D5A3AFEDEF947" LSID="urn:lsid:plazi:treatment:03A8E367FFAFFFAAFE0A58DAFE28FB8E:8EDE04AAFFAFFFA5FB8D5A3AFEDEF947" country="Philippines" county="Mindanao Island" latitude="8.2986" longLatPrecision="7" longitude="123.6169" municipality="Mt. Malindang Range" name="Don Victoriano" pageId="5" pageNumber="2238" stateProvince="Misamis Occidental">Don Victoriano</location>
,
<collectingRegion id="49C59C93FFAFFFA5FEB45ADDFD58F947" box="[292,653,1667,1691]" country="Philippines" name="Misamis Occidental" pageId="5" pageNumber="2238">Misamis Occidental Province</collectingRegion>
,
<collectingMunicipality id="6BDAC80BFFAFFFA5FD365ADAFC15F940" box="[678,960,1667,1692]" pageId="5" pageNumber="2238">Mt. Malindang Range</collectingMunicipality>
(
<geoCoordinate id="EE3534B6FFAFFFA5FC4D5ADDFB9EF947" box="[989,1099,1667,1691]" degrees="8.2986" direction="north" orientation="latitude" pageId="5" pageNumber="2238" precision="5" value="8.2986">8.2986°N</geoCoordinate>
,
<geoCoordinate id="EE3534B6FFACFFA6FF445847FE8DFBED" box="[212,344,1049,1073]" degrees="123.6169" direction="east" orientation="longitude" pageId="6" pageNumber="2239" precision="5" value="123.6169">123.6169°E</geoCoordinate>
),
<quantity id="4CF9FF94FFACFFA6FEFC5847FE6AFBEC" box="[364,447,1049,1073]" metricMagnitude="3" metricUnit="m" metricValue="1.3569999999999998" pageId="6" pageNumber="2239" unit="m" value="1357.0">
<elevation id="002CB542FFACFFA6FEFC5847FE6AFBEC" box="[364,447,1049,1073]" metricMagnitude="3" metricUnit="m" metricValue="1.3569999999999998" pageId="6" pageNumber="2239" unit="m" value="1357.0">1357 m</elevation>
</quantity>
a.s.l,
<collectingCounty id="62DF2AFDFFACFFA6FD935847FD1CFBED" box="[515,713,1049,1073]" pageId="6" pageNumber="2239">Mindanao Island</collectingCounty>
,
<collectingCountry id="F31612E1FFACFFA6FD455847FC81FBED" box="[725,852,1049,1073]" name="Philippines" pageId="6" pageNumber="2239">Philippines</collectingCountry>
; coll.
<collectorName id="26F437A7FFACFFA6FC085847FB95FBED" box="[920,1088,1049,1073]" pageId="6" pageNumber="2239">Nonillon Aspe</collectorName>
,
<collectorName id="26F437A7FFACFFA6FBDC5847FED9FB8D" pageId="6" pageNumber="2239">Nolan Aspe</collectorName>
and
<collectorName id="26F437A7FFACFFA6FEDC5867FE60FB8D" box="[332,437,1081,1105]" pageId="6" pageNumber="2239">J. Adeva</collectorName>
,
<date id="FFBF74B1FFACFFA6FE555867FD71FB8D" box="[453,676,1081,1105]" pageId="6" pageNumber="2239" value="2003-10-09" valueMax="2003-10-15" valueMin="2003-10-09">
<collectingDate id="EFFB8D59FFACFFA6FE555867FD71FB8D" box="[453,676,1081,1105]" pageId="6" pageNumber="2239" value="2003-10-09" valueMax="2003-10-15" valueMin="2003-10-09">915 October 2003</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B69582CFFACFFA6FD225867FBF4FBAC" ID-GBIF-Occurrence="2848941306" collectingDate="2003-10-09" collectingDateMax="2003-10-15" collectingDateMin="2003-10-09" collectionCode="ZRC" collectorName="Nonillon Aspe &amp; Nolan Aspe &amp; J. Adeva" country="Philippines" county="Mindanao Island" elevation="1357" latitude="8.2986" location="Barangay Lake Duminagat" longLatPrecision="7" longitude="123.6169" municipality="Mt. Malindang Range" pageId="6" pageNumber="2239" specimenCount="1" stateProvince="Misamis Occidental" typeStatus="paratype">
<typeStatus id="54BAECD3FFACFFA6FD225867FCF1FB8D" box="[690,804,1081,1105]" pageId="6" pageNumber="2239" type="paratype">Paratypes</typeStatus>
: one adult, one juvenile (NMA 4545); two adults (
<collectionCode id="ED10CAB4FFACFFA6FE395806FE30FBAC" box="[425,485,1112,1136]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="2239">ZRC</collectionCode>
.ANN.0013); same collection data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF7E02F9FFACFFA6FF445F00FD33FC10" ID-DOI="http://doi.org/10.5281/zenodo.4000015" ID-Zenodo-Dep="4000015" httpUri="https://zenodo.org/record/4000015/files/figure.png" pageId="6" pageNumber="2239" startId="6.[212,279,862,884]" targetBox="[212,1172,169,830]" targetPageId="6">
<paragraph id="8BBE5271FFACFFA6FF445F00FD33FC10" blockId="6.[212,1172,862,972]" pageId="6" pageNumber="2239">
Figure 2. Schematic drawings of the external anterio-ventral area of (A)
<taxonomicName id="4C0129F2FFACFFA6FC535F01FEE6FC4D" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="2239" phylum="Annelida" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="B9758E63FFACFFA6FC535F01FEE6FC4D" italics="true" pageId="6" pageNumber="2239">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFACFFA6FEAB5F22FE5AFC4D" box="[315,399,892,913]" pageId="6" pageNumber="2239" rank="species">sp. nov.</taxonomicNameLabel>
, (B)
<taxonomicName id="4C0129F2FFACFFA6FE545F22FD1CFC4D" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[452,713,892,913]" class="Clitellata" family="Megascolecidae" genus="Pithemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="2239" phylum="Annelida" rank="species" species="malindangensis" status="sp. nov.">
<emphasis id="B9758E63FFACFFA6FE545F22FD1CFC4D" box="[452,713,892,913]" italics="true" pageId="6" pageNumber="2239">Pithemera malindangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFACFFA6FD425F22FCF3FC4D" box="[722,806,892,913]" pageId="6" pageNumber="2239" rank="species">sp. nov.</taxonomicNameLabel>
, (C)
<taxonomicName id="4C0129F2FFACFFA6FCCC5F22FBEFFC4D" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[860,1082,892,913]" class="Clitellata" family="Megascolecidae" genus="Pithemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="2239" phylum="Annelida" rank="species" species="duminagati" status="sp. nov.">
<emphasis id="B9758E63FFACFFA6FCCC5F22FBEFFC4D" box="[860,1082,892,913]" italics="true" pageId="6" pageNumber="2239">Pithemera duminagati</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFACFFA6FBD25F22FB41FC4D" box="[1090,1172,892,913]" pageId="6" pageNumber="2239" rank="species">sp. nov.</taxonomicNameLabel>
and (D)
<taxonomicName id="4C0129F2FFACFFA6FEA15FC7FDFBFC72" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[305,558,921,942]" class="Clitellata" family="Megascolecidae" genus="Pithemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="2239" phylum="Annelida" rank="species" species="donvictorianoi" status="sp. nov.">
<emphasis id="B9758E63FFACFFA6FEA15FC7FDFBFC72" box="[305,558,921,942]" italics="true" pageId="6" pageNumber="2239">Pithemera donvictorianoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFACFFA6FDA85FC7FD5EFC72" box="[568,651,921,942]" pageId="6" pageNumber="2239" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: fp, female pore; cl, clitellum; mp, male pore; gm, genital markings. Scale bars: 5 mm.
</paragraph>
</caption>
<subSubSection id="C31B01FAFFACFFA8FF4458E0FCF5FD22" lastPageId="8" lastPageNumber="2241" pageId="6" pageNumber="2239" type="description">
<paragraph id="8BBE5271FFACFFA6FF4458E0FE86FB09" blockId="6.[212,1172,1214,1691]" box="[212,339,1214,1237]" pageId="6" pageNumber="2239">
<heading id="D0F6E51DFFACFFA6FF4458E0FE86FB09" box="[212,339,1214,1237]" fontSize="10" level="3" pageId="6" pageNumber="2239" reason="5">
<emphasis id="B9758E63FFACFFA6FF4458E0FE86FB09" box="[212,339,1214,1237]" italics="true" pageId="6" pageNumber="2239">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE5271FFACFFA6FF4458BAFD96FA20" blockId="6.[212,1172,1214,1691]" pageId="6" pageNumber="2239">
White, clitellum pinkish-grey. Body stout, adult length
<quantity id="4CF9FF94FFACFFA6FCD258BAFC1DFB20" box="[834,968,1252,1277]" metricMagnitude="-1" metricUnit="m" metricValue="1.04" metricValueMax="1.18" metricValueMin="0.9" pageId="6" pageNumber="2239" unit="mm" value="104.0" valueMax="118.0" valueMin="90.0">90118 mm</quantity>
; diameter
<quantity id="4CF9FF94FFACFFA6FBAE58BAFB41FB20" box="[1086,1172,1252,1276]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="2239" unit="mm" value="5.1">5.1 mm</quantity>
at x,
<quantity id="4CF9FF94FFACFFA6FE86595BFE8EFAC0" box="[278,347,1285,1308]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="2239" unit="mm" value="7.0">7 mm</quantity>
at xx; 140141 segments (n = four adults); body cylindrical in crosssection, tail narrowing gradually to sharp point. First dorsal pore at 12/13; spermathecal pores lacking or inconspicuous; female pore single in xiv, male pores on paired low papillae on xviii, 0.23 circumference apart, 10 setae between openings. Clitellum annular, from xiv to xvi. Setae pointed posteriorly, unevenly distributed around segmental equators; 4153 setae on vii, 4446 setae on xx; dorsal and ventral setal gaps lacking. Genital markings widely paired on xix to xxv and/or xxvi, between the sixth and seventh setal lines.
</paragraph>
<paragraph id="8BBE5271FFACFFA8FF6E5A5DFCD9FEDD" blockId="6.[212,1172,1214,1691]" lastBlockId="8.[212,1173,168,766]" lastPageId="8" lastPageNumber="2241" pageId="6" pageNumber="2239">Septa 4/5/6/7/8 muscular, 8/9 absent, 9/10 present around dorsal vessel and hearts; 10/11 to 13/14 thickly muscular. Dense tufts of nephridia on anterior faces of 5/6 and 6/7; nephridia of intestinal segments located at septumbody wall junction, mainly on body wall near anterior and posterior faces of septa. Oesophageal gizzard large, extending from viiix; oesophagus with lamellar inner surface extend from xixiii; intestinal origin in xv, caeca lacking; typhlosole a simple fold of about one sixth lumen diameter originating in xvi; intestinal wall without longitudinal blood vessels but with two pairs of vertical vessels per segment.</paragraph>
<caption id="DF7E02F9FFADFFA7FF005A76FD6CF949" ID-DOI="http://doi.org/10.5281/zenodo.4000017" ID-Zenodo-Dep="4000017" httpUri="https://zenodo.org/record/4000017/files/figure.png" pageId="7" pageNumber="2240" startId="7.[144,211,1576,1598]" targetBox="[190,1058,168,1544]" targetPageId="7">
<paragraph id="8BBE5271FFADFFA7FF005A76FD6CF949" blockId="7.[144,1104,1576,1686]" pageId="7" pageNumber="2240">
Figure 3. Schematic drawings of the internal morphology of (A)
<taxonomicName id="4C0129F2FFADFFA7FCB85A77FB9AF9E2" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[808,1103,1577,1598]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="2240" phylum="Annelida" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="B9758E63FFADFFA7FCB85A77FB9AF9E2" box="[808,1103,1577,1598]" italics="true" pageId="7" pageNumber="2240">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFADFFA7FF005A18FF30F987" box="[144,229,1606,1627]" pageId="7" pageNumber="2240" rank="species">sp. nov.</taxonomicNameLabel>
, (B)
<taxonomicName id="4C0129F2FFADFFA7FE8B5A18FDF4F987" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[283,545,1606,1627]" class="Clitellata" family="Megascolecidae" genus="Pithemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="2240" phylum="Annelida" rank="species" species="malindangensis" status="sp. nov.">
<emphasis id="B9758E63FFADFFA7FE8B5A18FDF4F987" box="[283,545,1606,1627]" italics="true" pageId="7" pageNumber="2240">Pithemera malindangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFADFFA7FDBB5A18FD55F987" box="[555,640,1606,1627]" pageId="7" pageNumber="2240" rank="species">sp. nov.</taxonomicNameLabel>
, (C)
<taxonomicName id="4C0129F2FFADFFA7FD285A18FC42F987" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[696,919,1606,1627]" class="Clitellata" family="Megascolecidae" genus="Pithemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="2240" phylum="Annelida" rank="species" species="duminagati" status="sp. nov.">
<emphasis id="B9758E63FFADFFA7FD285A18FC42F987" box="[696,919,1606,1627]" italics="true" pageId="7" pageNumber="2240">Pithemera duminagati</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFADFFA7FC315A18FC21F987" box="[929,1012,1606,1627]" pageId="7" pageNumber="2240" rank="species">sp. nov.</taxonomicNameLabel>
and (D)
<taxonomicName id="4C0129F2FFADFFA7FF005A3DFE5FF9A4" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[144,394,1635,1656]" class="Clitellata" family="Megascolecidae" genus="Pithemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="2240" phylum="Annelida" rank="species" species="donvictorianoi" status="sp. nov.">
<emphasis id="B9758E63FFADFFA7FF005A3DFE5FF9A4" box="[144,394,1635,1656]" italics="true" pageId="7" pageNumber="2240">Pithemera donvictorianoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFADFFA7FE005A3DFE34F9A4" box="[400,481,1635,1656]" pageId="7" pageNumber="2240" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: s, spermatheca; n, dense tufts of nephridia; h, heart; p, prostate gland; c, caecum. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="8BBE5271FFA2FFA8FF6E5D56FEBBFEA3" blockId="8.[212,1173,168,766]" pageId="8" pageNumber="2241">Hearts in x to xii, oesophageal. Commissural vessels in vi, vii and ix, lateral; lacking in viii. Supra-oesophageal vessel extending from x to xii; extra-oesophageal vessels joining ventral oesophageal wall in x, receiving efferent parieto-oesophageal vessels in xiv.</paragraph>
<paragraph id="8BBE5271FFA2FFA8FF6E5DD9FCF5FD22" blockId="8.[212,1173,168,766]" pageId="8" pageNumber="2241">Ovaries and funnels free in xiii, spermathecae lacking in two adults, one adult individual with five spermathecae closely lining up on the left side of vi, another adult individual with three spermathecae closely lining up on the left side and five spermathecae closely lining up on the right in vi and two spermathecae on left and two on right in vii; spermathecae small, ampulla pyriform, spermathecal duct short, slender; diverticulum stalk long and slender, attached ectally to duct, with one kink, terminating in short, sausage-shaped receptacle; no nephridia on spermathecal ducts. Male sexual system holandric, testes and funnels enclosed in paired ventral sacs in x and xi; seminal vesicles in xi and xii; pseudovesicles in xiii; vasa deferentia slender, free from body wall en route to ental end of prostatic ducts; prostates in xvi to xxi, each prostate racemose, trilobed but compact; copulatory bursae lacking.</paragraph>
</subSubSection>
<subSubSection id="C31B01FAFFA2FFAAFF445F0DFE28FB8E" lastPageId="10" lastPageNumber="2243" pageId="8" pageNumber="2241" type="discussion">
<paragraph id="8BBE5271FFA2FFA8FF445F0DFEE3FCB7" blockId="8.[212,1173,851,1649]" box="[212,310,851,875]" pageId="8" pageNumber="2241">
<heading id="D0F6E51DFFA2FFA8FF445F0DFEE3FCB7" box="[212,310,851,875]" fontSize="10" level="3" pageId="8" pageNumber="2241" reason="5">
<emphasis id="B9758E63FFA2FFA8FF445F0DFEE3FCB7" box="[212,310,851,875]" italics="true" pageId="8" pageNumber="2241">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE5271FFA2FFAAFF445F25FE42FE63" blockId="8.[212,1173,851,1649]" lastBlockId="10.[212,1172,168,831]" lastPageId="10" lastPageNumber="2243" pageId="8" pageNumber="2241">
<taxonomicName id="4C0129F2FFA2FFA8FF445F25FDE2FC4F" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[212,567,891,915]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="B9758E63FFA2FFA8FF445F25FDE2FC4F" box="[212,567,891,915]" italics="true" pageId="8" pageNumber="2241">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFA2FFA8FDDB5F22FD62FC4F" box="[587,695,892,915]" pageId="8" pageNumber="2241" rank="species">sp. nov.</taxonomicNameLabel>
, the first member of
<taxonomicName id="4C0129F2FFA2FFA8FC7C5F25FB41FC4F" authorityName="Michaelsen" authorityYear="1934" box="[1004,1172,891,915]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA2FFA8FC7C5F25FB41FC4F" box="[1004,1172,891,915]" italics="true" pageId="8" pageNumber="2241">Polypheretima</emphasis>
</taxonomicName>
reported from Mindanao Island, belongs to the
<taxonomicName id="4C0129F2FFA2FFA8FC9A5FC2FBEEFC6F" authority="Perrier, 1872" authorityName="Perrier" authorityYear="1872" box="[778,1083,923,947]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA2FFA8FC9A5FC2FC49FC6F" box="[778,924,923,947]" italics="true" pageId="8" pageNumber="2241">Po. elongata</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FC345FC5FBEEFC6F" author="Perrier E" box="[932,1083,923,947]" pageId="8" pageNumber="2241" pagination="5 - 198" refId="ref10741" refString="Perrier E. 1872. Recherches pour servir a l' hestoire des Lombriciens terrestres. Nouv Archs Mus Hist Nat Paris. 8: 5 - 198." type="journal article" year="1872">Perrier, 1872</bibRefCitation>
</taxonomicName>
species group of
<bibRefCitation id="EF902F80FFA2FFA8FED95FE5FE26FC0F" author="Easton EG" box="[329,499,955,979]" pageId="8" pageNumber="2241" pagination="1 - 128" refId="ref9694" refString="Easton EG. 1979. A revision of the ' acaecate' earthworms of the Pheretima group (Megascolecidae: Oligochaeta): Archipheretima, Metapheretima, Planapheretima, Pleionogaster and Polypheretima. Bull Brit Mus Nat Hist (Zool). 35: 1 - 128." type="journal article" year="1979">Easton (1979)</bibRefCitation>
, characterised by having a pair of genital markings in xix, successive segments in line with the male pores, paired batteries of up to 28 spermathecae in vi and/or vii, and shallow copulatory bursae with no stalked glands.
<bibRefCitation id="EF902F80FFA2FFA8FEA75845FE37FBEE" author="Easton EG" box="[311,482,1050,1074]" pageId="8" pageNumber="2241" pagination="1 - 128" refId="ref9694" refString="Easton EG. 1979. A revision of the ' acaecate' earthworms of the Pheretima group (Megascolecidae: Oligochaeta): Archipheretima, Metapheretima, Planapheretima, Pleionogaster and Polypheretima. Bull Brit Mus Nat Hist (Zool). 35: 1 - 128." type="journal article" year="1979">Easton (1979)</bibRefCitation>
included five species in the group:
<emphasis id="B9758E63FFA2FFA8FC295845FEFCFB8E" italics="true" pageId="8" pageNumber="2241">
<taxonomicName id="4C0129F2FFA2FFA8FC295845FB80FBEF" authorityName="Perrier" authorityYear="1872" box="[953,1109,1051,1075]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="elongata">Po. elongata</taxonomicName>
;
<taxonomicName id="4C0129F2FFA2FFA8FBFC5845FEFCFB8E" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="everetti">Po. everetti</taxonomicName>
</emphasis>
;
<taxonomicName id="4C0129F2FFA2FFA8FED75865FC4EFB8E" authority="Beddard and Fedarb, 1895" authorityName="Beddard and Fedarb" authorityYear="1895" box="[327,923,1082,1107]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="kinabaluensis">
<emphasis id="B9758E63FFA2FFA8FED75865FDCAFB8F" box="[327,543,1083,1107]" italics="true" pageId="8" pageNumber="2241">Po. kinabaluensis</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FDA55864FC4EFB8E" author="Beddard FE &amp; Fedarb SM" box="[565,923,1082,1106]" pageId="8" pageNumber="2241" pagination="69 - 73" refId="ref9546" refString="Beddard FE, Fedarb SM. 1895. On some Perichaetidae from the Eastern Archipelago collected by Mr. Everett. J Nat Hist Ser 6. 16: 69 - 73." type="journal article" year="1895">Beddard and Fedarb, 1895</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C0129F2FFA2FFA8FC2A5865FE74FBAE" authority="Michaelsen, 1899" authorityName="Michaelsen" authorityYear="1899" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="phacellotheca">
<emphasis id="B9758E63FFA2FFA8FC2A5865FB41FB8F" box="[954,1172,1083,1107]" italics="true" pageId="8" pageNumber="2241">Po. phacellotheca</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FF445804FE74FBAE" author="Michaelsen W" box="[212,417,1114,1138]" pageId="8" pageNumber="2241" pagination="1 - 122" refId="ref10479" refString="Michaelsen W. 1899. Terrikolen von verschiedenen Gebieten der Erde. Jb Hamb Wiss Anst. 16: 1 - 122." type="journal article" year="1899">Michaelsen, 1899</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C0129F2FFA2FFA8FE785805FCEBFBAE" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[488,830,1114,1138]" class="Clitellata" family="Megascolecidae" genus="Perichaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="stelleri">
<emphasis id="B9758E63FFA2FFA8FE785805FDB3FBAE" box="[488,614,1114,1138]" italics="true" pageId="8" pageNumber="2241">Po. stelleri</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FDE15804FCEBFBAE" author="Michaelsen W" box="[625,830,1114,1138]" pageId="8" pageNumber="2241" pagination="209 - 261" refId="ref10457" refString="Michaelsen W. 1892. Terricolen der Berliner Zoologischen Sammlung, II. Arch Naturgesch. 58: 209 - 261." type="journal article" year="1892">Michaelsen, 1892</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C0129F2FFA2FFA8FCC05804FF24FB4E" authorityName="Aspe &amp; James" authorityYear="2015" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="mindanaoensis">
<emphasis id="B9758E63FFA2FFA8FCC05804FF24FB4E" italics="true" pageId="8" pageNumber="2241">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
markedly differs from
<taxonomicName id="4C0129F2FFA2FFA8FD8F5825FD6DFB4E" authorityName="Perrier" authorityYear="1872" box="[543,696,1146,1170]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA2FFA8FD8F5825FD6DFB4E" box="[543,696,1146,1170]" italics="true" pageId="8" pageNumber="2241">Po. elongata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0129F2FFA2FFA8FC925825FC5BFB4E" box="[770,910,1147,1170]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="everetti">
<emphasis id="B9758E63FFA2FFA8FC925825FC5BFB4E" box="[770,910,1147,1170]" italics="true" pageId="8" pageNumber="2241">Po. everetti</emphasis>
</taxonomicName>
in size (355 ×
<quantity id="4CF9FF94FFA2FFA8FBF05824FB41FB4E" box="[1120,1172,1146,1170]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="2241" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C0129F2FFA2FFA8FF4458C5FEBCFB6E" authorityName="Perrier" authorityYear="1872" box="[212,361,1178,1202]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA2FFA8FF4458C5FEBCFB6E" box="[212,361,1178,1202]" italics="true" pageId="8" pageNumber="2241">Po. elongata</emphasis>
</taxonomicName>
; 300 ×
<quantity id="4CF9FF94FFA2FFA8FE4058C4FDDBFB6E" box="[464,526,1178,1202]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="8" pageNumber="2241" unit="in" value="12.0">12 in</quantity>
<taxonomicName id="4C0129F2FFA2FFA8FD8A58C5FD76FB6E" box="[538,675,1179,1202]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="everetti">
<emphasis id="B9758E63FFA2FFA8FD8A58C5FD76FB6E" box="[538,675,1179,1202]" italics="true" pageId="8" pageNumber="2241">Po. everetti</emphasis>
</taxonomicName>
), in the number of segments covered by the genital markings (extending from xix to xxii in
<taxonomicName id="4C0129F2FFA2FFA8FCC158E5FC32FB0E" authorityName="Perrier" authorityYear="1872" box="[849,999,1210,1234]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA2FFA8FCC158E5FC32FB0E" box="[849,999,1210,1234]" italics="true" pageId="8" pageNumber="2241">Po. elongata</emphasis>
</taxonomicName>
; xix to xxi in
<taxonomicName id="4C0129F2FFA2FFA8FF445885FE8EFB2E" box="[212,347,1243,1266]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="everetti">
<emphasis id="B9758E63FFA2FFA8FF445885FE8EFB2E" box="[212,347,1243,1266]" italics="true" pageId="8" pageNumber="2241">Po. everetti</emphasis>
</taxonomicName>
) and in the number of setae on vii (usually
<quantity id="4CF9FF94FFA2FFA8FCE95884FC3BFB2E" box="[889,1006,1242,1266]" metricMagnitude="0" metricUnit="m" metricValue="2.667" metricValueMax="3.3019999999999996" metricValueMin="2.032" pageId="8" pageNumber="2241" unit="in" value="105.0" valueMax="130.0" valueMin="80.0">80130 in</quantity>
<taxonomicName id="4C0129F2FFA2FFA8FC685885FB59FB2E" authorityName="Perrier" authorityYear="1872" box="[1016,1164,1242,1266]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA2FFA8FC685885FB59FB2E" box="[1016,1164,1242,1266]" italics="true" pageId="8" pageNumber="2241">Po. elongata</emphasis>
</taxonomicName>
;
<quantity id="4CF9FF94FFA2FFA8FF4458A7FEF7FACD" box="[212,290,1273,1297]" metricMagnitude="0" metricUnit="m" metricValue="3.3019999999999996" pageId="8" pageNumber="2241" unit="in" value="130.0">130 in</quantity>
<taxonomicName id="4C0129F2FFA2FFA8FEA058A5FE6EFACE" box="[304,443,1275,1298]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="everetti">
<emphasis id="B9758E63FFA2FFA8FEA058A5FE6EFACE" box="[304,443,1275,1298]" italics="true" pageId="8" pageNumber="2241">Po. everetti</emphasis>
</taxonomicName>
) (
<tableCitation id="C68367CAFFA2FFA8FE4C58A7FDEEFACD" box="[476,571,1273,1297]" captionStart="Table 2" captionStartId="9.[143,201,584,606]" captionText="Table 2. Comparison of characters among Polypheretima species in the Philippines. +, present; -, absent." pageId="8" pageNumber="2241">Table 2</tableCitation>
).
<taxonomicName id="4C0129F2FFA2FFA8FDCA58A4FC62FACE" authorityName="Aspe &amp; James" authorityYear="2015" box="[602,951,1274,1298]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="mindanaoensis">
<emphasis id="B9758E63FFA2FFA8FDCA58A4FC62FACE" box="[602,951,1274,1298]" italics="true" pageId="8" pageNumber="2241">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
(white) differs in colour from
<taxonomicName id="4C0129F2FFA2FFA8FEFF5944FE22FAED" box="[367,503,1306,1329]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="everetti">
<emphasis id="B9758E63FFA2FFA8FEFF5944FE22FAED" box="[367,503,1306,1329]" italics="true" pageId="8" pageNumber="2241">Po. everetti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0129F2FFA2FFA8FDAB5944FCDDFAEE" authorityName="Beddard and Fedarb" authorityYear="1895" box="[571,776,1306,1330]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="kinabaluensis">
<emphasis id="B9758E63FFA2FFA8FDAB5944FCDDFAEE" box="[571,776,1306,1330]" italics="true" pageId="8" pageNumber="2241">Po. kinabaluensis</emphasis>
</taxonomicName>
(reddish-purple and red, respectively). Those individuals of
<taxonomicName id="4C0129F2FFA2FFA8FDA15964FC5EFA8E" authorityName="Aspe &amp; James" authorityYear="2015" box="[561,907,1338,1362]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="mindanaoensis">
<emphasis id="B9758E63FFA2FFA8FDA15964FC5EFA8E" box="[561,907,1338,1362]" italics="true" pageId="8" pageNumber="2241">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
that have spermathecae have fewer spermathecae in each battery than
<taxonomicName id="4C0129F2FFA2FFA8FC8B5904FC0BFAAE" authorityName="Beddard and Fedarb" authorityYear="1895" box="[795,990,1370,1394]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="kinabaluensis">
<emphasis id="B9758E63FFA2FFA8FC8B5904FC0BFAAE" box="[795,990,1370,1394]" italics="true" pageId="8" pageNumber="2241">Po. kinabaluensis</emphasis>
</taxonomicName>
(
<quantity id="4CF9FF94FFA2FFA8FC7F5907FB91FAAD" box="[1007,1092,1369,1393]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="3.048" metricValueMin="1.524" pageId="8" pageNumber="2241" unit="in" value="9.0" valueMax="12.0" valueMin="6.0">612 in</quantity>
5/6/7),
<taxonomicName id="4C0129F2FFA2FFA8FF445924FE4EFA4D" authorityName="Michaelsen" authorityYear="1899" box="[212,411,1401,1425]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="phacellotheca">
<emphasis id="B9758E63FFA2FFA8FF445924FE4EFA4D" box="[212,411,1401,1425]" italics="true" pageId="8" pageNumber="2241">Po. phacellotheca</emphasis>
</taxonomicName>
(
<quantity id="4CF9FF94FFA2FFA8FE3F5927FDD2FA4D" box="[431,519,1401,1425]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="8" pageNumber="2241" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">912 in</quantity>
5/6) or
<taxonomicName id="4C0129F2FFA2FFA8FDFD5924FD3DFA4D" authorityName="Michaelsen" authorityYear="1892" box="[621,744,1401,1425]" class="Clitellata" family="Megascolecidae" genus="Perichaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="stelleri">
<emphasis id="B9758E63FFA2FFA8FDFD5924FD3DFA4D" box="[621,744,1401,1425]" italics="true" pageId="8" pageNumber="2241">Po. stelleri</emphasis>
</taxonomicName>
(up to
<quantity id="4CF9FF94FFA2FFA8FCD95927FC50FA4D" box="[841,901,1401,1425]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="8" pageNumber="2241" unit="in" value="28.0">28 in</quantity>
5/6/7), and individuals have fewer setae (4153) on vii than
<taxonomicName id="4C0129F2FFA2FFA8FD0659C4FC8AFA6D" authorityName="Michaelsen" authorityYear="1899" box="[662,863,1433,1457]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="phacellotheca">
<emphasis id="B9758E63FFA2FFA8FD0659C4FC8AFA6D" box="[662,863,1433,1457]" italics="true" pageId="8" pageNumber="2241">Po. phacellotheca</emphasis>
</taxonomicName>
(80) or
<taxonomicName id="4C0129F2FFA2FFA8FC5A59C4FB93FA6D" authorityName="Michaelsen" authorityYear="1892" box="[970,1094,1433,1457]" class="Clitellata" family="Megascolecidae" genus="Perichaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="stelleri">
<emphasis id="B9758E63FFA2FFA8FC5A59C4FB93FA6D" box="[970,1094,1433,1457]" italics="true" pageId="8" pageNumber="2241">Po. stelleri</emphasis>
</taxonomicName>
(130). Finally,
<taxonomicName id="4C0129F2FFA2FFA8FEA859E4FDDEFA0D" authorityName="Aspe &amp; James" authorityYear="2015" box="[312,523,1465,1489]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="mindanaoensis">
<emphasis id="B9758E63FFA2FFA8FEA859E4FDDEFA0D" box="[312,523,1465,1489]" italics="true" pageId="8" pageNumber="2241">Po. mindanaoensis</emphasis>
</taxonomicName>
differs from all congeners in the
<taxonomicName id="4C0129F2FFA2FFA8FC3959E4FBEFFA0D" authorityName="Perrier" authorityYear="1872" box="[937,1082,1465,1489]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA2FFA8FC3959E4FBEFFA0D" box="[937,1082,1465,1489]" italics="true" pageId="8" pageNumber="2241">Po. elongata</emphasis>
</taxonomicName>
species group in lacking copulatory bursae. Congeners reported from the
<collectingCountry id="F31612E1FFA2FFA8FB885986FB41FA2C" box="[1048,1172,1496,1520]" name="Philippines" pageId="8" pageNumber="2241">Philippines</collectingCountry>
include
<taxonomicName id="4C0129F2FFA2FFA8FEAB59A7FCFFF9CC" authority="Hong and James, 2008 a" authorityName="Hong and James" authorityYear="2008" box="[315,810,1528,1553]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="fruticosa">
<emphasis id="B9758E63FFA2FFA8FEAB59A7FE0DF9CD" box="[315,472,1529,1553]" italics="true" pageId="8" pageNumber="2241">Po. fruticosa</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FE7E59A7FCFFF9CC" author="Hong Y &amp; James SW" box="[494,810,1528,1553]" pageId="8" pageNumber="2241" pagination="341 - 354" refId="ref9836" refString="Hong Y, James SW. 2008 a. Nine new species of earthworms (Oligochaeta: Megascolecidae) of the Banaue rice terraces, Philippines. Rev Suisse Zool. 115: 341 - 354." type="journal article" year="2008">Hong and James, 2008a</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0129F2FFA2FFA8FCD759A7FEA6F9EC" authority="Hong and James, 2008 a" authorityName="Hong and James" authorityYear="2008" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="perlucidula">
<emphasis id="B9758E63FFA2FFA8FCD759A7FC29F9CD" box="[839,1020,1529,1553]" italics="true" pageId="8" pageNumber="2241">Po. perlucidula</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FB8359A7FEA6F9EC" author="Hong Y &amp; James SW" pageId="8" pageNumber="2241" pagination="341 - 354" refId="ref9836" refString="Hong Y, James SW. 2008 a. Nine new species of earthworms (Oligochaeta: Megascolecidae) of the Banaue rice terraces, Philippines. Rev Suisse Zool. 115: 341 - 354." type="journal article" year="2008">Hong and James, 2008a</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C0129F2FFA2FFA8FE565A47FC0DF9EC" authority="Hong and James, 2008 a" authorityName="Hong and James" authorityYear="2008" box="[454,984,1560,1585]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="bannaworensis">
<emphasis id="B9758E63FFA2FFA8FE565A47FD49F9ED" box="[454,668,1561,1585]" italics="true" pageId="8" pageNumber="2241">Po. bannaworensis</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FD3D5A47FC0DF9EC" author="Hong Y &amp; James SW" box="[685,984,1560,1585]" pageId="8" pageNumber="2241" pagination="341 - 354" refId="ref9836" refString="Hong Y, James SW. 2008 a. Nine new species of earthworms (Oligochaeta: Megascolecidae) of the Banaue rice terraces, Philippines. Rev Suisse Zool. 115: 341 - 354." type="journal article" year="2008">Hong and James, 2008a</bibRefCitation>
</taxonomicName>
, from Banaue, Luzon Island;
<taxonomicName id="4C0129F2FFA2FFA8FE115A67FCA9F98C" authority="Hong and James, 2011 a" authorityName="Hong and James" authorityYear="2011" box="[385,892,1592,1617]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="pagudpudensis">
<emphasis id="B9758E63FFA2FFA8FE115A67FD87F98C" box="[385,594,1592,1616]" italics="true" pageId="8" pageNumber="2241">Po. pagudpudensis</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FDCF5A67FCA9F98C" author="Hong Y &amp; James SW" box="[607,892,1592,1617]" pageId="8" pageNumber="2241" pagination="19 - 28" refId="ref10026" refString="Hong Y, James SW. 2011 a. New species of Pheretima, Pithemera, and Polypheretima (Clitellata: Megascolecidae) from Kalbaryo, Luzon Island, Philippines. Raffles B Zool. 59: 19 - 28." type="journal article" year="2011">Hong and James, 2011a</bibRefCitation>
</taxonomicName>
from Kalbaryo, Luzon Island;
<taxonomicName id="4C0129F2FFA2FFA8FEA25A07FD42F9AC" authority="Beddard, 1912" authorityName="Beddard" authorityYear="1912" box="[306,663,1624,1648]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2241" phylum="Annelida" rank="species" species="monticola">
<emphasis id="B9758E63FFA2FFA8FEA25A07FE02F9AC" box="[306,471,1624,1648]" italics="true" pageId="8" pageNumber="2241">Po. monticola</emphasis>
<bibRefCitation id="EF902F80FFA2FFA8FE795A06FD42F9AC" author="Beddard FE" box="[489,663,1624,1648]" pageId="8" pageNumber="2241" pagination="79 - 203" refId="ref9517" refString="Beddard FE. 1912. The oligochaeta terricolae of the Philippines. Part 1. The genus Pheretima. Philipp J Sci. 7: 79 - 203." type="journal article" year="1912">Beddard, 1912</bibRefCitation>
</taxonomicName>
from Mt. Pulong (Mt. Pulag),
<collectingRegion id="49C59C93FFA2FFAAFBA85A07FEECFF1C" country="Philippines" lastPageId="10" lastPageNumber="2243" name="Benguet" pageId="8" pageNumber="2241">Benguet Province</collectingRegion>
, and from Luzon Island; and
<taxonomicName id="4C0129F2FFA0FFAAFD075CF7FCCDFF1C" box="[663,792,169,192]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="everetti">
<emphasis id="B9758E63FFA0FFAAFD075CF7FCCDFF1C" box="[663,792,169,192]" italics="true" pageId="10" pageNumber="2243">Po. everetti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0129F2FFA0FFAAFCC55CF7FC36FF1C" authorityName="Perrier" authorityYear="1872" box="[853,995,168,192]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA0FFAAFCC55CF7FC36FF1C" box="[853,995,168,192]" italics="true" pageId="10" pageNumber="2243">Po. elongata</emphasis>
</taxonomicName>
, both collected on Balabac Island,
<collectingRegion id="49C59C93FFA0FFAAFE585C96FDF9FF3C" box="[456,556,200,224]" country="Philippines" name="Palawan" pageId="10" pageNumber="2243">Palawan</collectingRegion>
.
<emphasis id="B9758E63FFA0FFAAFDAF5C96FEA7FEDC" italics="true" pageId="10" pageNumber="2243">
<taxonomicName id="4C0129F2FFA0FFAAFDAF5C96FCB9FF3C" authorityName="Hong and James" authorityYear="2008" box="[575,876,200,224]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="perlucidula">Polypheretima perlucidula</taxonomicName>
,
<taxonomicName id="4C0129F2FFA0FFAAFC115C97FB8FFF3C" authorityName="Hong and James" authorityYear="2008" box="[897,1114,200,224]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="bannaworensis">Po. bannaworensis</taxonomicName>
,
<taxonomicName id="4C0129F2FFA0FFAAFBFD5C97FEA7FEDC" authorityName="Hong and James" authorityYear="2011" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="pagudpudensis">Po. pagudpudensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0129F2FFA0FFAAFE515CB7FDB3FEDC" authorityName="Beddard" authorityYear="1912" box="[449,614,232,256]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="monticola">
<emphasis id="B9758E63FFA0FFAAFE515CB7FDB3FEDC" box="[449,614,232,256]" italics="true" pageId="10" pageNumber="2243">Po. monticola</emphasis>
</taxonomicName>
belong to the
<taxonomicName id="4C0129F2FFA0FFAAFCA55CB7FC62FEDC" baseAuthorityName="Michaelsen" baseAuthorityYear="1924" box="[821,951,232,256]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="bifaria">
<emphasis id="B9758E63FFA0FFAAFCA55CB7FC62FEDC" box="[821,951,232,256]" italics="true" pageId="10" pageNumber="2243">Po. bifaria</emphasis>
</taxonomicName>
species group of
<bibRefCitation id="EF902F80FFA0FFAAFF445D56FEA1FEFC" author="Easton EG" box="[212,372,264,288]" pageId="10" pageNumber="2243" pagination="1 - 128" refId="ref9694" refString="Easton EG. 1979. A revision of the ' acaecate' earthworms of the Pheretima group (Megascolecidae: Oligochaeta): Archipheretima, Metapheretima, Planapheretima, Pleionogaster and Polypheretima. Bull Brit Mus Nat Hist (Zool). 35: 1 - 128." type="journal article" year="1979">Easton (1979)</bibRefCitation>
, characterised by having pairs of spermathecae in 5/68/9 or only in 6/7 or 7/8, while
<taxonomicName id="4C0129F2FFA0FFAAFE095D77FDFCFE9C" authorityName="Hong and James" authorityYear="2008" box="[409,553,296,320]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="fruticosa">
<emphasis id="B9758E63FFA0FFAAFE095D77FDFCFE9C" box="[409,553,296,320]" italics="true" pageId="10" pageNumber="2243">Po. fruticosa</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C0129F2FFA0FFAAFC895D77FB59FE9C" authority="Michaelsen, 1907" authorityName="Michaelsen" authorityYear="1907" box="[793,1164,296,320]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="voeltzkowi">
<emphasis id="B9758E63FFA0FFAAFC895D77FC68FE9C" box="[793,957,296,320]" italics="true" pageId="10" pageNumber="2243">Po. voeltzkowi</emphasis>
<bibRefCitation id="EF902F80FFA0FFAAFC575D76FB59FE9C" author="Michaelsen W" box="[967,1164,296,320]" pageId="10" pageNumber="2243" pagination="41 - 50" refId="ref10502" refString="Michaelsen W. 1907. Oligochaten von Madagaskar, den Comoren und anderen Inseln des westlichen Indischen Ozeans. Reise in Ostafrika in den Jahren 1903 - 1905. Wiss Ergebn Reise Ostafr. 2: 41 - 50." type="journal article" year="1907">Michaelsen, 1907</bibRefCitation>
</taxonomicName>
, characterised by having a pair of spermathecae in only in 5/6. The new species also differs from the other species in the number of setae between male pores, in the genital marking pattern and in the size and segmental position of the prostate glands (
<tableCitation id="C68367CAFFA0FFAAFEBF5DF9FE52FE63" box="[303,391,423,447]" captionStart="Table 2" captionStartId="9.[143,201,584,606]" captionText="Table 2. Comparison of characters among Polypheretima species in the Philippines. +, present; -, absent." pageId="10" pageNumber="2243">Table 2</tableCitation>
).
</paragraph>
<caption id="DF7E02F9FFA3FFA9FF1F5E16FB13FD82" box="[143,1222,584,606]" pageId="9" pageNumber="2242">
<paragraph id="8BBE5271FFA3FFA9FF1F5E16FB13FD82" blockId="9.[143,1222,584,606]" box="[143,1222,584,606]" pageId="9" pageNumber="2242">
Table 2. Comparison of characters among
<taxonomicName id="4C0129F2FFA3FFA9FDDD5E16FD0BFD81" authorityName="Michaelsen" authorityYear="1934" box="[589,734,584,605]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FDDD5E16FD0BFD81" box="[589,734,584,605]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
species in the Philippines. +, present; -, absent.
</paragraph>
</caption>
<paragraph id="8BBE5271FFA3FFA9FF1F5E24F953FAE3" pageId="9" pageNumber="2242">
<table id="F901A0D1FFA3005FFF1F5E24F96EFAE3" box="[143,1723,634,1343]" gridcols="9" gridrows="25" pageId="9" pageNumber="2242">
<tr id="35315033FFA3005FFF1F5E24F96EFD50" box="[143,1723,634,652]" gridrow="0" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5E24FEE7FD50" box="[143,306,634,652]" gridcol="0" gridrow="0" pageId="9" pageNumber="2242">Character</th>
<th id="76E0394FFFA3005FFEC45E24FE3AFD50" box="[340,495,634,652]" gridcol="1" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FEF95E24FE09FD50" authorityName="Michaelsen" authorityYear="1934" box="[361,476,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FEF95E24FE09FD50" box="[361,476,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
<th id="76E0394FFFA3005FFD815E24FD1FFD50" box="[529,714,634,652]" gridcol="2" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FDA15E24FD7CFD50" authorityName="Michaelsen" authorityYear="1934" box="[561,681,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FDA15E24FD7CFD50" box="[561,681,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
<th id="76E0394FFFA3005FFD795E24FC5FFD50" box="[745,906,634,652]" gridcol="3" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FC905E24FCADFD50" authorityName="Michaelsen" authorityYear="1934" box="[768,888,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FC905E24FCADFD50" box="[768,888,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
<th id="76E0394FFFA3005FFC3F5E24FB95FD50" box="[943,1088,634,652]" gridcol="4" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FC2A5E24FBE7FD50" authorityName="Michaelsen" authorityYear="1934" box="[954,1074,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FC2A5E24FBE7FD50" box="[954,1074,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
<th id="76E0394FFFA3005FFBCE5E24FB08FD50" box="[1118,1245,634,652]" gridcol="5" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FBF25E24FB0FFD50" authorityName="Michaelsen" authorityYear="1934" box="[1122,1242,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FBF25E24FB0FFD50" box="[1122,1242,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
<th id="76E0394FFFA3005FFB6E5E24FAAFFD50" box="[1278,1402,634,652]" gridcol="6" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FA915E24FAACFD50" authorityName="Michaelsen" authorityYear="1934" box="[1281,1401,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FA915E24FAACFD50" box="[1281,1401,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
<th id="76E0394FFFA3005FFA0D5E24F9CFFD50" box="[1437,1562,634,652]" gridcol="7" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FA315E24F9CCFD50" authorityName="Michaelsen" authorityYear="1934" box="[1441,1561,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9FA315E24F9CCFD50" box="[1441,1561,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
<th id="76E0394FFFA3005FF9AF5E24F96EFD50" box="[1599,1723,634,652]" gridcol="8" gridrow="0" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9F9D35E24F96EFD50" authorityName="Michaelsen" authorityYear="1934" box="[1603,1723,634,652]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="genus">
<emphasis id="B9758E63FFA3FFA9F9D35E24F96EFD50" box="[1603,1723,634,652]" italics="true" pageId="9" pageNumber="2242">Polypheretima</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35315033FFA3005FFF1F5EC9F96EFD75" box="[143,1723,663,681]" gridrow="1" pageId="9" pageNumber="2242" rowspan-0="1">
<td id="76E0394FFFA3005FFEC45EC9FE3AFD75" box="[340,495,663,681]" gridcol="1" gridrow="1" pageId="9" pageNumber="2242">
<emphasis id="B9758E63FFA3FFA9FEF05EC9FE1CFD75" box="[352,457,663,681]" italics="true" pageId="9" pageNumber="2242">mindanaoesis</emphasis>
sp.
</td>
<td id="76E0394FFFA3005FFD815EC9FD1FFD75" box="[529,714,663,681]" gridcol="2" gridrow="1" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FD815EC9FD1FFD75" authority="Perrier, 1872" authorityName="Perrier" authorityYear="1872" box="[529,714,663,681]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA3FFA9FD815EC9FD82FD75" box="[529,599,663,681]" italics="true" pageId="9" pageNumber="2242">elongata</emphasis>
<bibRefCitation id="EF902F80FFA3FFA9FDCD5EC9FD1FFD75" author="Perrier E" box="[605,714,663,681]" pageId="9" pageNumber="2242" pagination="5 - 198" refId="ref10741" refString="Perrier E. 1872. Recherches pour servir a l' hestoire des Lombriciens terrestres. Nouv Archs Mus Hist Nat Paris. 8: 5 - 198." type="journal article" year="1872">Perrier, 1872</bibRefCitation>
</taxonomicName>
</td>
<td id="76E0394FFFA3005FFD795EC9FC5FFD75" box="[745,906,663,681]" gridcol="3" gridrow="1" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FD675EC9FC57FD75" authority="Beddard" authorityName="Beddard" box="[759,898,663,681]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="species" species="everetti">
<emphasis id="B9758E63FFA3FFA9FD675EC9FCE1FD75" box="[759,820,663,681]" italics="true" pageId="9" pageNumber="2242">everetti</emphasis>
Beddard
</taxonomicName>
</td>
<td id="76E0394FFFA3005FFC3F5EC9FB95FD75" box="[943,1088,663,681]" gridcol="4" gridrow="1" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FC5E5EC9FBCAFD75" authorityName="Beddard" authorityYear="1912" box="[974,1055,663,681]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="species" species="monticola">
<emphasis id="B9758E63FFA3FFA9FC5E5EC9FBCAFD75" box="[974,1055,663,681]" italics="true" pageId="9" pageNumber="2242">monticola</emphasis>
</taxonomicName>
</td>
<td id="76E0394FFFA3005FFBCE5EC9FB08FD75" box="[1118,1245,663,681]" gridcol="5" gridrow="1" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FBCF5EC9FB08FD75" authority="Hong" authorityName="Hong and James" authorityYear="2008" box="[1119,1245,663,681]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="species" species="fruticosa">
<emphasis id="B9758E63FFA3FFA9FBCF5EC9FB72FD75" box="[1119,1191,663,681]" italics="true" pageId="9" pageNumber="2242">fruticosa</emphasis>
Hong
</taxonomicName>
</td>
<td id="76E0394FFFA3005FFB6E5EC9FAAFFD75" box="[1278,1402,663,681]" gridcol="6" gridrow="1" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FA805EC9FABEFD75" authorityName="Hong and James" authorityYear="2008" box="[1296,1387,663,681]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="species" species="perlucidula">
<emphasis id="B9758E63FFA3FFA9FA805EC9FABEFD75" box="[1296,1387,663,681]" italics="true" pageId="9" pageNumber="2242">perlucidula</emphasis>
</taxonomicName>
</td>
<td id="76E0394FFFA3005FFA0D5EC9F9CFFD75" box="[1437,1562,663,681]" gridcol="7" gridrow="1" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9FA325EC9F9CFFD75" authorityName="Hong and James" authorityYear="2008" box="[1442,1562,663,681]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="species" species="bannaworensis">
<emphasis id="B9758E63FFA3FFA9FA325EC9F9CFFD75" box="[1442,1562,663,681]" italics="true" pageId="9" pageNumber="2242">bannaworensis</emphasis>
</taxonomicName>
</td>
<td id="76E0394FFFA3005FF9AF5EC9F96EFD75" box="[1599,1723,663,681]" gridcol="8" gridrow="1" pageId="9" pageNumber="2242">
<taxonomicName id="4C0129F2FFA3FFA9F9D45EC9F96EFD75" authorityName="Hong and James" authorityYear="2011" box="[1604,1723,663,681]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2242" phylum="Annelida" rank="species" species="pagudpudensis">
<emphasis id="B9758E63FFA3FFA9F9D45EC9F96EFD75" box="[1604,1723,663,681]" italics="true" pageId="9" pageNumber="2242">pagudpudensis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="35315033FFA3005FFF1F5EEAF96EFD1A" box="[143,1723,692,710]" gridrow="2" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-2="1">
<td id="76E0394FFFA3005FFEC45EEAFE3AFD1A" box="[340,495,692,710]" gridcol="1" gridrow="2" pageId="9" pageNumber="2242">nov.</td>
<td id="76E0394FFFA3005FFD795EEAFC5FFD1A" box="[745,906,692,710]" gridcol="3" gridrow="2" pageId="9" pageNumber="2242">and Fedarb, 1895</td>
<td id="76E0394FFFA3005FFC3F5EEAFB95FD1A" box="[943,1088,692,710]" gridcol="4" gridrow="2" pageId="9" pageNumber="2242">
<bibRefCitation id="EF902F80FFA3FFA9FC295EEAFBE0FD1A" author="Beddard FE" box="[953,1077,692,710]" pageId="9" pageNumber="2242" pagination="79 - 203" refId="ref9517" refString="Beddard FE. 1912. The oligochaeta terricolae of the Philippines. Part 1. The genus Pheretima. Philipp J Sci. 7: 79 - 203." type="journal article" year="1912">Beddard, 1912</bibRefCitation>
</td>
<td id="76E0394FFFA3005FFBCE5EEAFB08FD1A" box="[1118,1245,692,710]" gridcol="5" gridrow="2" pageId="9" pageNumber="2242">and James,</td>
<td id="76E0394FFFA3005FFB6E5EEAFAAFFD1A" box="[1278,1402,692,710]" gridcol="6" gridrow="2" pageId="9" pageNumber="2242">Hong and</td>
<td id="76E0394FFFA3005FFA0D5EEAF9CFFD1A" box="[1437,1562,692,710]" gridcol="7" gridrow="2" pageId="9" pageNumber="2242">Hong and</td>
<td id="76E0394FFFA3005FF9AF5EEAF96EFD1A" box="[1599,1723,692,710]" gridcol="8" gridrow="2" pageId="9" pageNumber="2242">Hong and</td>
</tr>
<tr id="35315033FFA3005FFF1F5E8FF96EFD3F" box="[143,1723,721,739]" gridrow="3" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="76E0394FFFA3005FFBCE5E8FFB08FD3F" box="[1118,1245,721,739]" gridcol="5" gridrow="3" pageId="9" pageNumber="2242">2008a</td>
<td id="76E0394FFFA3005FFB6E5E8FFAAFFD3F" box="[1278,1402,721,739]" gridcol="6" gridrow="3" pageId="9" pageNumber="2242">James, 2008a</td>
<td id="76E0394FFFA3005FFA0D5E8FF9CFFD3F" box="[1437,1562,721,739]" gridcol="7" gridrow="3" pageId="9" pageNumber="2242">James, 2008a</td>
<td id="76E0394FFFA3005FF9AF5E8FF96EFD3F" box="[1599,1723,721,739]" gridcol="8" gridrow="3" pageId="9" pageNumber="2242">James, 2011a</td>
</tr>
<tr id="35315033FFA3005FFF1F5F5CF96EFCC8" box="[143,1723,770,788]" gridrow="4" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5F5CFEE7FCC8" box="[143,306,770,788]" gridcol="0" gridrow="4" pageId="9" pageNumber="2242">Body pigmentation</th>
<td id="76E0394FFFA3005FFEC45F5CFE3AFCC8" box="[340,495,770,788]" gridcol="1" gridrow="4" pageId="9" pageNumber="2242">White</td>
<td id="76E0394FFFA3005FFD815F5CFD1FFCC8" box="[529,714,770,788]" gridcol="2" gridrow="4" pageId="9" pageNumber="2242">White</td>
<td id="76E0394FFFA3005FFD795F5CFC5FFCC8" box="[745,906,770,788]" gridcol="3" gridrow="4" pageId="9" pageNumber="2242">Reddish purple</td>
<td id="76E0394FFFA3005FFC3F5F5CFB95FCC8" box="[943,1088,770,788]" gridcol="4" gridrow="4" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFBCE5F5CFB08FCC8" box="[1118,1245,770,788]" gridcol="5" gridrow="4" pageId="9" pageNumber="2242">Brown dorsal</td>
<td id="76E0394FFFA3005FFB6E5F5CFAAFFCC8" box="[1278,1402,770,788]" gridcol="6" gridrow="4" pageId="9" pageNumber="2242">White</td>
<td id="76E0394FFFA3005FFA0D5F5CF9CFFCC8" box="[1437,1562,770,788]" gridcol="7" gridrow="4" pageId="9" pageNumber="2242">Brownish</td>
<td id="76E0394FFFA3005FF9AF5F5CF96EFCC8" box="[1599,1723,770,788]" gridcol="8" gridrow="4" pageId="9" pageNumber="2242">White</td>
</tr>
<tr id="35315033FFA3005FFF1F5F41F96EFCED" box="[143,1723,799,817]" gridrow="5" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<td id="76E0394FFFA3005FFA0D5F41F9CFFCED" box="[1437,1562,799,817]" gridcol="7" gridrow="5" pageId="9" pageNumber="2242">dorsal</td>
</tr>
<tr id="35315033FFA3005FFF1F5F67F96EFC97" box="[143,1723,825,843]" gridrow="6" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5F67FEE7FC97" box="[143,306,825,843]" gridcol="0" gridrow="6" pageId="9" pageNumber="2242">Length</th>
<td id="76E0394FFFA3005FFEC45F67FE3AFC97" box="[340,495,825,843]" gridcol="1" gridrow="6" pageId="9" pageNumber="2242">90118</td>
<td id="76E0394FFFA3005FFD815F67FD1FFC97" box="[529,714,825,843]" gridcol="2" gridrow="6" pageId="9" pageNumber="2242">355</td>
<td id="76E0394FFFA3005FFD795F67FC5FFC97" box="[745,906,825,843]" gridcol="3" gridrow="6" pageId="9" pageNumber="2242">300</td>
<td id="76E0394FFFA3005FFC3F5F67FB95FC97" box="[943,1088,825,843]" gridcol="4" gridrow="6" pageId="9" pageNumber="2242">130</td>
<td id="76E0394FFFA3005FFBCE5F67FB08FC97" box="[1118,1245,825,843]" gridcol="5" gridrow="6" pageId="9" pageNumber="2242">3961</td>
<td id="76E0394FFFA3005FFB6E5F67FAAFFC97" box="[1278,1402,825,843]" gridcol="6" gridrow="6" pageId="9" pageNumber="2242">3856</td>
<td id="76E0394FFFA3005FFA0D5F67F9CFFC97" box="[1437,1562,825,843]" gridcol="7" gridrow="6" pageId="9" pageNumber="2242">5174</td>
<td id="76E0394FFFA3005FF9AF5F67F96EFC97" box="[1599,1723,825,843]" gridcol="8" gridrow="6" pageId="9" pageNumber="2242">5663</td>
</tr>
<tr id="35315033FFA3005FFF1F5F0AF96EFCBA" box="[143,1723,852,870]" gridrow="7" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5F0AFEE7FCBA" box="[143,306,852,870]" gridcol="0" gridrow="7" pageId="9" pageNumber="2242">Setae vii; xx</th>
<td id="76E0394FFFA3005FFEC45F0AFE3AFCBA" box="[340,495,852,870]" gridcol="1" gridrow="7" pageId="9" pageNumber="2242">4153; 4446</td>
<td id="76E0394FFFA3005FFD815F0AFD1FFCBA" box="[529,714,852,870]" gridcol="2" gridrow="7" pageId="9" pageNumber="2242">Usually 80 to 130;?</td>
<td id="76E0394FFFA3005FFD795F0AFC5FFCBA" box="[745,906,852,870]" gridcol="3" gridrow="7" pageId="9" pageNumber="2242">Numerous, up to</td>
<td id="76E0394FFFA3005FFC3F5F0AFB95FCBA" box="[943,1088,852,870]" gridcol="4" gridrow="7" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFBCE5F0AFB08FCBA" box="[1118,1245,852,870]" gridcol="5" gridrow="7" pageId="9" pageNumber="2242">44; 41</td>
<td id="76E0394FFFA3005FFB6E5F0AFAAFFCBA" box="[1278,1402,852,870]" gridcol="6" gridrow="7" pageId="9" pageNumber="2242">50; 23</td>
<td id="76E0394FFFA3005FFA0D5F0AF9CFFCBA" box="[1437,1562,852,870]" gridcol="7" gridrow="7" pageId="9" pageNumber="2242">39; 46</td>
<td id="76E0394FFFA3005FF9AF5F0AF96EFCBA" box="[1599,1723,852,870]" gridcol="8" gridrow="7" pageId="9" pageNumber="2242">63; 27</td>
</tr>
<tr id="35315033FFA3005FFF1F5F2FF96EFC5F" box="[143,1723,881,899]" gridrow="8" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76E0394FFFA3005FFD795F2FFC5FFC5F" box="[745,906,881,899]" gridcol="3" gridrow="8" pageId="9" pageNumber="2242">130;?</td>
</tr>
<tr id="35315033FFA3005FFF1F5FD2F96EFC42" box="[143,1723,908,926]" gridrow="9" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5FD2FEE7FC42" box="[143,306,908,926]" gridcol="0" gridrow="9" pageId="9" pageNumber="2242">Setae bet. male</th>
<td id="76E0394FFFA3005FFEC45FD2FE3AFC42" box="[340,495,908,926]" gridcol="1" gridrow="9" pageId="9" pageNumber="2242">10</td>
<td id="76E0394FFFA3005FFD815FD2FD1FFC42" box="[529,714,908,926]" gridcol="2" gridrow="9" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFD795FD2FC5FFC42" box="[745,906,908,926]" gridcol="3" gridrow="9" pageId="9" pageNumber="2242">612</td>
<td id="76E0394FFFA3005FFC3F5FD2FB95FC42" box="[943,1088,908,926]" gridcol="4" gridrow="9" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFBCE5FD2FB08FC42" box="[1118,1245,908,926]" gridcol="5" gridrow="9" pageId="9" pageNumber="2242">9</td>
<td id="76E0394FFFA3005FFB6E5FD2FAAFFC42" box="[1278,1402,908,926]" gridcol="6" gridrow="9" pageId="9" pageNumber="2242">12</td>
<td id="76E0394FFFA3005FFA0D5FD2F9CFFC42" box="[1437,1562,908,926]" gridcol="7" gridrow="9" pageId="9" pageNumber="2242">67</td>
<td id="76E0394FFFA3005FF9AF5FD2F96EFC42" box="[1599,1723,908,926]" gridcol="8" gridrow="9" pageId="9" pageNumber="2242">8</td>
</tr>
<tr id="35315033FFA3005FFF1F5FF7F96EFC67" box="[143,1723,937,955]" gridrow="10" pageId="9" pageNumber="2242" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76E0394FFFA3005FFF1F5FF7FEE7FC67" box="[143,306,937,955]" gridcol="0" gridrow="10" pageId="9" pageNumber="2242">pores</th>
</tr>
<tr id="35315033FFA3005FFF1F5F9DF96EFC0A" box="[143,1723,963,982]" gridrow="11" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5F9DFEE7FC0A" box="[143,306,963,982]" gridcol="0" gridrow="11" pageId="9" pageNumber="2242">Sper. pore spacing</th>
<td id="76E0394FFFA3005FFEC45F9DFE3AFC0A" box="[340,495,963,982]" gridcol="1" gridrow="11" pageId="9" pageNumber="2242"></td>
<td id="76E0394FFFA3005FFD815F9DFD1FFC0A" box="[529,714,963,982]" gridcol="2" gridrow="11" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFD795F9DFC5FFC0A" box="[745,906,963,982]" gridcol="3" gridrow="11" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFC3F5F9DFB95FC0A" box="[943,1088,963,982]" gridcol="4" gridrow="11" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFBCE5F9DFB08FC0A" box="[1118,1245,963,982]" gridcol="5" gridrow="11" pageId="9" pageNumber="2242">0.25</td>
<td id="76E0394FFFA3005FFB6E5F9DFAAFFC0A" box="[1278,1402,963,982]" gridcol="6" gridrow="11" pageId="9" pageNumber="2242">0.28</td>
<td id="76E0394FFFA3005FFA0D5F9DF9CFFC0A" box="[1437,1562,963,982]" gridcol="7" gridrow="11" pageId="9" pageNumber="2242">0.220.25</td>
<td id="76E0394FFFA3005FF9AF5F9DF96EFC0A" box="[1599,1723,963,982]" gridcol="8" gridrow="11" pageId="9" pageNumber="2242">?</td>
</tr>
<tr id="35315033FFA3005FFF1F5F80F96EFC2C" box="[143,1723,990,1008]" gridrow="12" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5F80FEE7FC2C" box="[143,306,990,1008]" gridcol="0" gridrow="12" pageId="9" pageNumber="2242">Male pore spacing</th>
<td id="76E0394FFFA3005FFEC45F80FE3AFC2C" box="[340,495,990,1008]" gridcol="1" gridrow="12" pageId="9" pageNumber="2242">0.23</td>
<td id="76E0394FFFA3005FFD815F80FD1FFC2C" box="[529,714,990,1008]" gridcol="2" gridrow="12" pageId="9" pageNumber="2242">0.25</td>
<td id="76E0394FFFA3005FFD795F80FC5FFC2C" box="[745,906,990,1008]" gridcol="3" gridrow="12" pageId="9" pageNumber="2242">0.25</td>
<td id="76E0394FFFA3005FFC3F5F80FB95FC2C" box="[943,1088,990,1008]" gridcol="4" gridrow="12" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFBCE5F80FB08FC2C" box="[1118,1245,990,1008]" gridcol="5" gridrow="12" pageId="9" pageNumber="2242">0.27</td>
<td id="76E0394FFFA3005FFB6E5F80FAAFFC2C" box="[1278,1402,990,1008]" gridcol="6" gridrow="12" pageId="9" pageNumber="2242">0.32</td>
<td id="76E0394FFFA3005FFA0D5F80F9CFFC2C" box="[1437,1562,990,1008]" gridcol="7" gridrow="12" pageId="9" pageNumber="2242">0.24</td>
<td id="76E0394FFFA3005FF9AF5F80F96EFC2C" box="[1599,1723,990,1008]" gridcol="8" gridrow="12" pageId="9" pageNumber="2242">0.23</td>
</tr>
<tr id="35315033FFA3005FFF1F5FA6F96EFBD7" box="[143,1723,1016,1035]" gridrow="13" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5FA6FEE7FBD7" box="[143,306,1016,1035]" gridcol="0" gridrow="13" pageId="9" pageNumber="2242">Location of</th>
<td id="76E0394FFFA3005FFEC45FA6FE3AFBD7" box="[340,495,1016,1035]" gridcol="1" gridrow="13" pageId="9" pageNumber="2242">0 or up to 5</td>
<td id="76E0394FFFA3005FFD815FA6FD1FFBD7" box="[529,714,1016,1035]" gridcol="2" gridrow="13" pageId="9" pageNumber="2242">Rarely more than 3</td>
<td id="76E0394FFFA3005FFD795FA6FC5FFBD7" box="[745,906,1016,1035]" gridcol="3" gridrow="13" pageId="9" pageNumber="2242">612 spermathecae</td>
<td id="76E0394FFFA3005FFC3F5FA6FB95FBD7" box="[943,1088,1016,1035]" gridcol="4" gridrow="13" pageId="9" pageNumber="2242">Paired in 5/68/9</td>
<td id="76E0394FFFA3005FFBCE5FA6FB08FBD7" box="[1118,1245,1016,1035]" gridcol="5" gridrow="13" pageId="9" pageNumber="2242">Paired in 5/6</td>
<td id="76E0394FFFA3005FFB6E5FA6FAAFFBD7" box="[1278,1402,1016,1035]" gridcol="6" gridrow="13" pageId="9" pageNumber="2242">Paired in 5/6/7</td>
<td id="76E0394FFFA3005FFA0D5FA6F9CFFBD7" box="[1437,1562,1016,1035]" gridcol="7" gridrow="13" pageId="9" pageNumber="2242">Paired in 5/</td>
<td id="76E0394FFFA3005FF9AF5FA6F96EFBD7" box="[1599,1723,1016,1035]" gridcol="8" gridrow="13" pageId="9" pageNumber="2242">Paired in 5/6/7</td>
</tr>
<tr id="35315033FFA3005FFF1F5848F96EFBF4" box="[143,1723,1046,1064]" gridrow="14" pageId="9" pageNumber="2242" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="76E0394FFFA3005FFF1F5848FEE7FBF4" box="[143,306,1046,1064]" gridcol="0" gridrow="14" pageId="9" pageNumber="2242">spermathecae</th>
<td id="76E0394FFFA3005FFEC45848FE3AFBF4" box="[340,495,1046,1064]" gridcol="1" gridrow="14" pageId="9" pageNumber="2242">spermathecae</td>
<td id="76E0394FFFA3005FFD815848FD1FFBF4" box="[529,714,1046,1064]" gridcol="2" gridrow="14" pageId="9" pageNumber="2242">spermathecae in</td>
<td id="76E0394FFFA3005FFD795848FC5FFBF4" box="[745,906,1046,1064]" gridcol="3" gridrow="14" pageId="9" pageNumber="2242">in each battery</td>
<td id="76E0394FFFA3005FFA0D5848F9CFFBF4" box="[1437,1562,1046,1064]" gridcol="7" gridrow="14" pageId="9" pageNumber="2242">68/9</td>
</tr>
<tr id="35315033FFA3005FFF1F586DF96EFB99" box="[143,1723,1075,1093]" gridrow="15" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76E0394FFFA3005FFEC4586DFE3AFB99" box="[340,495,1075,1093]" gridcol="1" gridrow="15" pageId="9" pageNumber="2242">in each battery</td>
<td id="76E0394FFFA3005FFD81586DFD1FFB99" box="[529,714,1075,1093]" gridcol="2" gridrow="15" pageId="9" pageNumber="2242">each battery in 5/6/7</td>
<td id="76E0394FFFA3005FFD79586DFC5FFB99" box="[745,906,1075,1093]" gridcol="3" gridrow="15" pageId="9" pageNumber="2242">in 5/6/7</td>
</tr>
<tr id="35315033FFA3005FFF1F580EF96EFBBE" box="[143,1723,1104,1122]" gridrow="16" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76E0394FFFA3005FFEC4580EFE3AFBBE" box="[340,495,1104,1122]" gridcol="1" gridrow="16" pageId="9" pageNumber="2242">in 5/6, or 5/6/7</td>
</tr>
<tr id="35315033FFA3005FFF1F5835F96EFBA1" box="[143,1723,1131,1149]" gridrow="17" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5835FEE7FBA1" box="[143,306,1131,1149]" gridcol="0" gridrow="17" pageId="9" pageNumber="2242">Genital markings</th>
<td id="76E0394FFFA3005FFEC45835FE3AFBA1" box="[340,495,1131,1149]" gridcol="1" gridrow="17" pageId="9" pageNumber="2242">Paired in xixxxvi</td>
<td id="76E0394FFFA3005FFD815835FD1FFBA1" box="[529,714,1131,1149]" gridcol="2" gridrow="17" pageId="9" pageNumber="2242">Paired in ixxxii</td>
<td id="76E0394FFFA3005FFD795835FC5FFBA1" box="[745,906,1131,1149]" gridcol="3" gridrow="17" pageId="9" pageNumber="2242">Paired in xixxxi</td>
<td id="76E0394FFFA3005FFC3F5835FB95FBA1" box="[943,1088,1131,1149]" gridcol="4" gridrow="17" pageId="9" pageNumber="2242">Paired in ix, xvii,</td>
<td id="76E0394FFFA3005FFBCE5835FB08FBA1" box="[1118,1245,1131,1149]" gridcol="5" gridrow="17" pageId="9" pageNumber="2242">Paired in xviii</td>
<td id="76E0394FFFA3005FFB6E5835FAAFFBA1" box="[1278,1402,1131,1149]" gridcol="6" gridrow="17" pageId="9" pageNumber="2242"></td>
<td id="76E0394FFFA3005FFA0D5835F9CFFBA1" box="[1437,1562,1131,1149]" gridcol="7" gridrow="17" pageId="9" pageNumber="2242"></td>
<td id="76E0394FFFA3005FF9AF5835F96EFBA1" box="[1599,1723,1131,1149]" gridcol="8" gridrow="17" pageId="9" pageNumber="2242">Paired in xvii,</td>
</tr>
<tr id="35315033FFA3005FFF1F58D6F96EFB46" box="[143,1723,1160,1178]" gridrow="18" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-6="1" rowspan-7="1">
<td id="76E0394FFFA3005FFC3F58D6FB95FB46" box="[943,1088,1160,1178]" gridcol="4" gridrow="18" pageId="9" pageNumber="2242">xixxx</td>
<td id="76E0394FFFA3005FFBCE58D6FB08FB46" box="[1118,1245,1160,1178]" gridcol="5" gridrow="18" pageId="9" pageNumber="2242">or xix</td>
<td id="76E0394FFFA3005FF9AF58D6F96EFB46" box="[1599,1723,1160,1178]" gridcol="8" gridrow="18" pageId="9" pageNumber="2242">xix, xx</td>
</tr>
<tr id="35315033FFA3005FFF1F58FDF96EFB69" box="[143,1723,1187,1205]" gridrow="19" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F58FDFEE7FB69" box="[143,306,1187,1205]" gridcol="0" gridrow="19" pageId="9" pageNumber="2242">Septa in 5/613/14</th>
<td id="76E0394FFFA3005FFEC458FDFE3AFB69" box="[340,495,1187,1205]" gridcol="1" gridrow="19" pageId="9" pageNumber="2242">+ in 4/5; in 8/9</td>
<td id="76E0394FFFA3005FFD8158FDFD1FFB69" box="[529,714,1187,1205]" gridcol="2" gridrow="19" pageId="9" pageNumber="2242">- in 9/10</td>
<td id="76E0394FFFA3005FFD7958FDFC5FFB69" box="[745,906,1187,1205]" gridcol="3" gridrow="19" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFC3F58FDFB95FB69" box="[943,1088,1187,1205]" gridcol="4" gridrow="19" pageId="9" pageNumber="2242">- in 5/68/9 and</td>
<td id="76E0394FFFA3005FFBCE58FDFB08FB69" box="[1118,1245,1187,1205]" gridcol="5" gridrow="19" pageId="9" pageNumber="2242">- in 8/9</td>
<td id="76E0394FFFA3005FFB6E58FDFAAFFB69" box="[1278,1402,1187,1205]" gridcol="6" gridrow="19" pageId="9" pageNumber="2242">+</td>
<td id="76E0394FFFA3005FFA0D58FDF9CFFB69" box="[1437,1562,1187,1205]" gridcol="7" gridrow="19" pageId="9" pageNumber="2242">+</td>
<td id="76E0394FFFA3005FF9AF58FDF96EFB69" box="[1599,1723,1187,1205]" gridcol="8" gridrow="19" pageId="9" pageNumber="2242">+</td>
</tr>
<tr id="35315033FFA3005FFF1F589EF96EFB0E" box="[143,1723,1216,1234]" gridrow="20" pageId="9" pageNumber="2242" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76E0394FFFA3005FFC3F589EFB95FB0E" box="[943,1088,1216,1234]" gridcol="4" gridrow="20" pageId="9" pageNumber="2242">13/14</td>
</tr>
<tr id="35315033FFA3005FFF1F5884F96EFB30" box="[143,1723,1242,1260]" gridrow="21" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5884FEE7FB30" box="[143,306,1242,1260]" gridcol="0" gridrow="21" pageId="9" pageNumber="2242">Origin of gizzard</th>
<td id="76E0394FFFA3005FFEC45884FE3AFB30" box="[340,495,1242,1260]" gridcol="1" gridrow="21" pageId="9" pageNumber="2242">viii</td>
<td id="76E0394FFFA3005FFD815884FD1FFB30" box="[529,714,1242,1260]" gridcol="2" gridrow="21" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFD795884FC5FFB30" box="[745,906,1242,1260]" gridcol="3" gridrow="21" pageId="9" pageNumber="2242">viii</td>
<td id="76E0394FFFA3005FFC3F5884FB95FB30" box="[943,1088,1242,1260]" gridcol="4" gridrow="21" pageId="9" pageNumber="2242">viii</td>
<td id="76E0394FFFA3005FFBCE5884FB08FB30" box="[1118,1245,1242,1260]" gridcol="5" gridrow="21" pageId="9" pageNumber="2242">viii</td>
<td id="76E0394FFFA3005FFB6E5884FAAFFB30" box="[1278,1402,1242,1260]" gridcol="6" gridrow="21" pageId="9" pageNumber="2242">viii</td>
<td id="76E0394FFFA3005FFA0D5884F9CFFB30" box="[1437,1562,1242,1260]" gridcol="7" gridrow="21" pageId="9" pageNumber="2242">viii</td>
<td id="76E0394FFFA3005FF9AF5884F96EFB30" box="[1599,1723,1242,1260]" gridcol="8" gridrow="21" pageId="9" pageNumber="2242">viii</td>
</tr>
<tr id="35315033FFA3005FFF1F58ABF96EFADB" box="[143,1723,1269,1287]" gridrow="22" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F58ABFEE7FADB" box="[143,306,1269,1287]" gridcol="0" gridrow="22" pageId="9" pageNumber="2242">Origin of</th>
<td id="76E0394FFFA3005FFEC458ABFE3AFADB" box="[340,495,1269,1287]" gridcol="1" gridrow="22" pageId="9" pageNumber="2242">xvi</td>
<td id="76E0394FFFA3005FFD8158ABFD1FFADB" box="[529,714,1269,1287]" gridcol="2" gridrow="22" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFD7958ABFC5FFADB" box="[745,906,1269,1287]" gridcol="3" gridrow="22" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFC3F58ABFB95FADB" box="[943,1088,1269,1287]" gridcol="4" gridrow="22" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFBCE58ABFB08FADB" box="[1118,1245,1269,1287]" gridcol="5" gridrow="22" pageId="9" pageNumber="2242">xx</td>
<td id="76E0394FFFA3005FFB6E58ABFAAFFADB" box="[1278,1402,1269,1287]" gridcol="6" gridrow="22" pageId="9" pageNumber="2242">xxii</td>
<td id="76E0394FFFA3005FFA0D58ABF9CFFADB" box="[1437,1562,1269,1287]" gridcol="7" gridrow="22" pageId="9" pageNumber="2242">xxii</td>
<td id="76E0394FFFA3005FF9AF58ABF96EFADB" box="[1599,1723,1269,1287]" gridcol="8" gridrow="22" pageId="9" pageNumber="2242">xxii</td>
</tr>
<tr id="35315033FFA3005FFF1F594CF96EFAF8" box="[143,1723,1298,1316]" gridrow="23" pageId="9" pageNumber="2242" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76E0394FFFA3005FFF1F594CFEE7FAF8" box="[143,306,1298,1316]" gridcol="0" gridrow="23" pageId="9" pageNumber="2242">typhlosole</th>
</tr>
<tr id="35315033FFA3005FFF1F5973F96EFAE3" box="[143,1723,1325,1343]" gridrow="24" pageId="9" pageNumber="2242">
<th id="76E0394FFFA3005FFF1F5973FEE7FAE3" box="[143,306,1325,1343]" gridcol="0" gridrow="24" pageId="9" pageNumber="2242">Prostate glands</th>
<td id="76E0394FFFA3005FFEC45973FE3AFAE3" box="[340,495,1325,1343]" gridcol="1" gridrow="24" pageId="9" pageNumber="2242">xvixxi</td>
<td id="76E0394FFFA3005FFD815973FD1FFAE3" box="[529,714,1325,1343]" gridcol="2" gridrow="24" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFD795973FC5FFAE3" box="[745,906,1325,1343]" gridcol="3" gridrow="24" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFC3F5973FB95FAE3" box="[943,1088,1325,1343]" gridcol="4" gridrow="24" pageId="9" pageNumber="2242">?</td>
<td id="76E0394FFFA3005FFBCE5973FB08FAE3" box="[1118,1245,1325,1343]" gridcol="5" gridrow="24" pageId="9" pageNumber="2242">xviiixxii</td>
<td id="76E0394FFFA3005FFB6E5973FAAFFAE3" box="[1278,1402,1325,1343]" gridcol="6" gridrow="24" pageId="9" pageNumber="2242">xviixx</td>
<td id="76E0394FFFA3005FFA0D5973F9CFFAE3" box="[1437,1562,1325,1343]" gridcol="7" gridrow="24" pageId="9" pageNumber="2242">xvixx</td>
<td id="76E0394FFFA3005FF9AF5973F96EFAE3" box="[1599,1723,1325,1343]" gridcol="8" gridrow="24" pageId="9" pageNumber="2242">xviixxii</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBE5271FFA0FFAAFF6E5D99FCECFD02" blockId="10.[212,1172,168,831]" pageId="10" pageNumber="2243">
Spermathecae function to receive and store sperm released by the male pores of the partner during copulation; loss of spermathecae usually means a loss of male function. Loss of spermathecae, reduction of testes and lack of spermatozoal iridescence in the sperm funnels in earthworms are indicative of reproduction by parthenogenesis (
<bibRefCitation id="EF902F80FFA0FFAAFD9D5E19FD4DFD83" author="Gates GE" box="[525,664,583,607]" pageId="10" pageNumber="2243" pagination="1 - 327" refId="ref9766" refString="Gates GE. 1972. Burmese earthworms: an introduction to the systematics and biology of megadrile oligochaetes with special reference to Southeast Asia. Trans Am Phil Soc. 62: 1 - 327." type="journal article" year="1972">Gates 1972</bibRefCitation>
), in which the egg develops into a new individual without being fertilised by sperm. In the case of
<taxonomicName id="4C0129F2FFA0FFAAFC375E36FBE9FDA3" authorityName="Perrier" authorityYear="1872" box="[935,1084,615,639]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="elongata">
<emphasis id="B9758E63FFA0FFAAFC375E36FBE9FDA3" box="[935,1084,615,639]" italics="true" pageId="10" pageNumber="2243">Po. elongata</emphasis>
</taxonomicName>
, which has been introduced into many parts of the world (see
<tableCitation id="C68367CAFFA0FFAAFCE65ED8FC07FD42" box="[886,978,646,670]" captionStart="Table 4" captionStartId="18.[212,270,1020,1042]" captionText="Table 4. Diversity of Polypheretima and Pithemera species around the world. Symbols indicate species with broad ranges: *Polypheretima elongata; +Polypheretima everetti; @Polypheretima taprobanae; #Pithemera bicincta. These data are from Tsai et al. (2000); Blakemore (2003, 2007); James (2004b); James et al. (2004); Hong and James (2008a, 2011a); Nguyen et al. (2014, 2015)." pageId="10" pageNumber="2243">Table 4</tableCitation>
),
<bibRefCitation id="EF902F80FFA0FFAAFC7E5ED9FB41FD42" author="Easton EG" box="[1006,1172,646,670]" pageId="10" pageNumber="2243" pagination="1 - 128" refId="ref9694" refString="Easton EG. 1979. A revision of the ' acaecate' earthworms of the Pheretima group (Megascolecidae: Oligochaeta): Archipheretima, Metapheretima, Planapheretima, Pleionogaster and Polypheretima. Bull Brit Mus Nat Hist (Zool). 35: 1 - 128." type="journal article" year="1979">Easton (1979)</bibRefCitation>
observed that athecate individuals are especially numerous in introduced populations, presumably reproducing parthenogenetically.
</paragraph>
<paragraph id="8BBE5271FFA0FFAAFF6E5EB9FD80FCE2" blockId="10.[212,1172,168,831]" pageId="10" pageNumber="2243">
In one specimen of
<taxonomicName id="4C0129F2FFA0FFAAFE4D5EB9FD7EFD22" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[477,683,742,766]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="B9758E63FFA0FFAAFE4D5EB9FD7EFD22" box="[477,683,742,766]" italics="true" pageId="10" pageNumber="2243">Po. mindanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFA0FFAAFD235EB9FCDAFD22" box="[691,783,743,766]" pageId="10" pageNumber="2243" rank="species">sp. nov.</taxonomicNameLabel>
, the ventral nerve cord has a solid dark core, perhaps related to parasitism. We observed several nematodes near an empty nerve cord swelling in xvii.
</paragraph>
<paragraph id="8BBE5271FFA0FFAAFF445FCDFE84FC77" blockId="10.[212,1172,915,1106]" box="[212,337,915,939]" pageId="10" pageNumber="2243">
<heading id="D0F6E51DFFA0FFAAFF445FCDFE84FC77" box="[212,337,915,939]" fontSize="10" level="3" pageId="10" pageNumber="2243" reason="5">
<emphasis id="B9758E63FFA0FFAAFF445FCDFE84FC77" box="[212,337,915,939]" italics="true" pageId="10" pageNumber="2243">Occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE5271FFA0FFAAFF445FE5FE28FB8E" blockId="10.[212,1172,915,1106]" pageId="10" pageNumber="2243">
<taxonomicName id="4C0129F2FFA0FFAAFF445FE5FDF0FC0F" authority="Aspe &amp; James, 2015" authorityName="Aspe &amp; James" authorityYear="2015" box="[212,549,955,979]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="2243" phylum="Annelida" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="B9758E63FFA0FFAAFF445FE5FDF0FC0F" box="[212,549,955,979]" italics="true" pageId="10" pageNumber="2243">Polypheretima mindanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2463318FFA0FFAAFDBF5FE5FD59FC0E" box="[559,652,955,978]" pageId="10" pageNumber="2243" rank="species">sp. nov.</taxonomicNameLabel>
was found at two of five collecting sites. In all, eight individuals were collected in disturbed forests in Barangays Lake Duminagat and Sibucal at elevations of
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a.s.l. The soil it inhabited was covered with thick leaf litter and roots, bryophytes and lichens. We did not observe it on rotten logs.
</paragraph>
</subSubSection>
</treatment>
</document>