215 lines
20 KiB
XML
215 lines
20 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3790301" ID-GBIF-Dataset="884a65c3-a7f0-46bb-8018-8f930c0e73d0" ID-GBIF-Taxon="163750690" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790301" approvalRequired="18" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="17" checkinTime="1587985477403" checkinUser="plazi" docAuthor="Hlavac, Peter & Jaszay, Tomas" docDate="2009" docId="03A8043AF51DFFF4FF026EDEE844728C" docLanguage="en" docName="ZK_article_2108.pdf" docOrigin="ZooKeys 29 (29)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@3c251258" docStyleName="zookeys.2008.journal_article" docTitle="Zyras Stephens 1835" docType="treatment" docVersion="4" lastPageId="4" lastPageNumber="53" masterDocId="FF917C42F51EFFF0FF8E6E33EC177404" masterDocTitle="A revision of the genus Zyras (Zyras) Stephens, 1835 (Coleoptera, Staphylinidae, Aleocharinae). I. Current classification status and the redefinition of the genus" masterLastPageNumber="71" masterPageNumber="49" pageId="3" pageNumber="52" updateTime="1643566306252" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the genus Zyras (Zyras) Stephens, 1835 (Coleoptera, Staphylinidae, Aleocharinae). I. Current classification status and the redefinition of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hlavac, Peter</mods:namePart>
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<mods:affiliation>Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Department of Forest Protection and Entomology, Praha, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jaszay, Tomas</mods:namePart>
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<mods:affiliation>Museum of Šariš, Natural History Department, Bardejov ,, Slovakia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-12-11</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>29</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>29</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>49</mods:start>
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<mods:end>71</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.29.218</mods:identifier>
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<mods:identifier type="GBIF-Dataset">884a65c3-a7f0-46bb-8018-8f930c0e73d0</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576574</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790301" ID-GBIF-Taxon="163750690" ID-Zenodo-Dep="3790301" LSID="urn:lsid:plazi:treatment:03A8043AF51DFFF4FF026EDEE844728C" httpUri="http://treatment.plazi.org/id/03A8043AF51DFFF4FF026EDEE844728C" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
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<subSubSection box="[140,479,236,263]" pageId="3" pageNumber="52" type="nomenclature">
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<paragraph blockId="3.[140,479,236,263]" box="[140,479,236,263]" pageId="3" pageNumber="52">
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<heading box="[140,479,236,263]" fontSize="11" level="2" pageId="3" pageNumber="52" reason="3">
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<emphasis bold="true" box="[140,479,236,263]" pageId="3" pageNumber="52">
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Genus
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<taxonomicName authority="Stephens, 1835" authorityName="Stephens" authorityYear="1835" box="[223,479,236,263]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[223,288,237,263]" italics="true" pageId="3" pageNumber="52">Zyras</emphasis>
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<bibRefCitation author="Stephens JF" box="[296,479,236,263]" pageId="3" pageNumber="52" refId="ref10677" refString="Stephens JF (1835) Illustrations of British entomology, or a synopsis of indigenous insects, containing their generic and specific distinctions, with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, Vol. 5. Baldwin & Cradock, London. pp. 369 - 477." type="book" year="1835">Stephens, 1835</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="52" type="reference_group">
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<paragraph blockId="3.[140,1108,307,651]" pageId="3" pageNumber="52">
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<treatmentCitationGroup pageId="3" pageNumber="52">
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<treatmentCitation author="Stephens JF" box="[140,453,307,334]" page="430" pageId="3" pageNumber="52" year="1835">
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<taxonomicName authority="Stephens, 1835: 430" authorityName="Stephens" authorityPageNumber="430" authorityYear="1835" box="[140,453,307,334]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,196,308,333]" italics="true" pageId="3" pageNumber="52">Zyras</emphasis>
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<bibRefCitation author="Stephens JF" box="[208,453,307,334]" pageId="3" pageNumber="52" refId="ref10677" refString="Stephens JF (1835) Illustrations of British entomology, or a synopsis of indigenous insects, containing their generic and specific distinctions, with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, Vol. 5. Baldwin & Cradock, London. pp. 369 - 477." type="book" year="1835">Stephens, 1835: 430</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus box="[467,522,308,334]" pageId="3" pageNumber="52">Type</typeStatus>
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species:
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<taxonomicName authority="(Stephens)" baseAuthorityName="Stephens" baseAuthorityYear="1832" box="[629,931,307,334]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="haworthi">
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<emphasis box="[629,797,307,333]" italics="true" pageId="3" pageNumber="52">Zyras haworthi</emphasis>
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(Stephens)
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</taxonomicName>
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, established by decision of the International Commission on Zoological Nomenclature (ICZN 1961, Opinion 599)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="3.[140,1108,307,651]" pageId="3" pageNumber="52">
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<treatmentCitationGroup pageId="3" pageNumber="52">
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<taxonomicName authority="Stephens" authorityName="Stephens" authorityYear="1835" box="[140,308,413,439]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,196,414,439]" italics="true" pageId="3" pageNumber="52">Zyras</emphasis>
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Stephens
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</taxonomicName>
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: Cameron, 1939: 497 (key to subgenera and species of
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<collectingCountry box="[924,986,413,439]" name="India" pageId="3" pageNumber="52">India</collectingCountry>
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,
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<collectingCountry box="[997,1108,414,440]" name="Myanmar" pageId="3" pageNumber="52">Myanmar</collectingCountry>
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and
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<collectingCountry box="[237,351,448,475]" name="Sri Lanka" pageId="3" pageNumber="52">Sri Lanka</collectingCountry>
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);
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<treatmentCitation author="Kistner DH" box="[373,584,448,474]" page="143" pageId="3" pageNumber="52" year="1972">
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<bibRefCitation author="Kistner DH" box="[373,584,448,474]" pageId="3" pageNumber="52" pagination="141 - 165" refId="ref8737" refString="Kistner DH (1972) Studies of Japanese myrmecophiles. Part I: The genera Pella and Falagria (Coleoptera, Staphylinidae). In: Entomological Essays to Commemorate the Retirement of Professor K. Yasumatsu, Hokurynken, Tokio 141 - 165." type="book chapter" year="1972">Kistner 1972: 143</bibRefCitation>
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</treatmentCitation>
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(redescription);
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<treatmentCitation author="Scheerpeltz O" box="[776,1017,448,474]" page="39" pageId="3" pageNumber="52" year="1974">
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<bibRefCitation author="Scheerpeltz O" box="[776,1017,448,474]" pageId="3" pageNumber="52" pagination="1 - 49" refId="ref10510" refString="Scheerpeltz O (1974) Coleoptera aus Nordostafrika, Staphylinidae. Acta Entomologica Fennica 31: 1 - 49." type="journal article" year="1974">Scheerpeltz 1974: 39</bibRefCitation>
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</treatmentCitation>
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(key to African subgenera);
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<treatmentCitation author="Seevers CH" box="[410,613,483,510]" page="153" pageId="3" pageNumber="52" year="1978">
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<bibRefCitation author="Seevers CH" box="[410,613,483,510]" pageId="3" pageNumber="52" refId="ref10481" refString="Seevers CH (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology 71: 275 pp." type="book" year="1978">Seevers 1978: 153</bibRefCitation>
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</treatmentCitation>
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(key to Nearctic species);
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<bibRefCitation author="Dvorak M" box="[903,1054,483,510]" pageId="3" pageNumber="52" pagination="93 - 98" refId="ref8408" refString="Dvorak M (1980) Ceskoslovenske druhy rodu Zyras Stephens (Col. Staphylinidae). Entomologicke Problemy 16: 93 - 98." type="journal article" year="1980">Dvořák 1980</bibRefCitation>
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(key to species of Czech and
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<collectingCountry box="[468,655,519,545]" name="Slovakia" pageId="3" pageNumber="52">Slovak Republic</collectingCountry>
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, including
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<emphasis italics="true" pageId="3" pageNumber="52">
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<taxonomicName authorityName="Stephens" authorityYear="1835" box="[787,843,519,545]" class="Insecta" family="Staphylinidae" genus="Pella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Pella</taxonomicName>
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,
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<taxonomicName authorityName="Tottenham" authorityYear="1939" box="[857,920,519,545]" class="Insecta" family="Staphylinidae" genus="Lepla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Lepla</taxonomicName>
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,
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<taxonomicName authorityName="Reitter" authorityYear="1909" box="[934,1104,519,545]" class="Insecta" family="Staphylinidae" genus="Pellochromonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Pellochromonia</taxonomicName>
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,
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<taxonomicName authorityName="Stephens" authorityYear="1835" box="[185,246,555,580]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Zyras</taxonomicName>
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,
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<taxonomicName box="[256,376,555,580]" class="Insecta" family="Staphylinidae" genus="Myrmoecia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Myrmoecia</taxonomicName>
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</emphasis>
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);
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<treatmentCitation author="Dvorak M" box="[400,608,554,580]" page="200" pageId="3" pageNumber="52" year="1984">
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<bibRefCitation author="Dvorak M" box="[400,608,554,580]" pageId="3" pageNumber="52" pagination="190 - 203" refId="ref8462" refString="Dvorak M (1984) Zur Kenntnis einiger myrmekophiler Staphylinidae Coleoptera). Acta Entomologica Bohemoslovaca 81: 190 - 203." type="journal article" year="1984">Dvořák 1984: 200</bibRefCitation>
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</treatmentCitation>
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(key to Palaearctic species including
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<collectingCountry box="[1020,1094,554,581]" name="China" pageId="3" pageNumber="52">China</collectingCountry>
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);
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<bibRefCitation author="Hastir P & Gaspar Ch" box="[185,459,589,615]" pageId="3" pageNumber="52" pagination="17 - 26" refId="ref8579" refString="Hastir P, Gaspar Ch (2001) La myrmecophilie chez les Staphylinidae: la cohabitation de Zyras haworthi Stephens et de Formica sanguinea Latreille. Notes faunistiques de Gembloux, 43: 17 - 26." type="journal article" year="2001">Hastir and Gaspar 2001</bibRefCitation>
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(biology);
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<treatmentCitation author="Klimaszewski J & Pelletier G & Maruyama M & Hlavac P" box="[583,922,589,616]" page="714" pageId="3" pageNumber="52" year="2005">
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<bibRefCitation author="Klimaszewski J & Pelletier G & Maruyama M & Hlavac P" box="[583,922,589,616]" pageId="3" pageNumber="52" pagination="703 - 733" refId="ref8973" refString="Klimaszewski J, Pelletier G, Maruyama M, Hlavac P (2005) Canadian species of the Zyras group of genera and review of the types from America north of Mexico. Revue Suisse de Zoologie 112 (3): 703 - 733." type="journal article" year="2005">Klimaszewski et al. 2005: 714</bibRefCitation>
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</treatmentCitation>
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(restricted redefinition, key to species of America north of
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<collectingCountry box="[677,767,624,651]" name="Mexico" pageId="3" pageNumber="52">Mexico</collectingCountry>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="52" type="diagnosis">
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<paragraph blockId="3.[140,1108,694,1672]" pageId="3" pageNumber="52">
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<emphasis bold="true" box="[140,263,694,720]" pageId="3" pageNumber="52">
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<emphasis box="[140,258,694,720]" inLineHeading="true" pageId="3" pageNumber="52" reason="1">Diagnosis</emphasis>
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.
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</emphasis>
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<taxonomicName authority="(Zyras)" box="[269,404,695,721]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="subGenus" subGenus="Zyras">
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<emphasis box="[269,324,695,720]" italics="true" pageId="3" pageNumber="52">Zyras</emphasis>
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(
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<emphasis box="[340,394,695,720]" italics="true" pageId="3" pageNumber="52">Zyras</emphasis>
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)
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</taxonomicName>
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can be distinguished from the other genera of
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<taxonomicName authorityName="Fleming" authorityYear="1821" box="[913,1056,695,721]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
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by a combination of the following characters: 1) whole body shiny, unicoloured or often with different colouration for head, pronotum, elytra and abdomen, sometimes covered with long setae; 2) all antennomeres petiolate, antennomere III about as long as pedicel; 3) neck absent; 4) pronotum usually with well-defined median antebasal fovea; 5) abdomen simple, not physogastric, parallel-sided; 6) simplified aedeagus, median lobe with enlarged basal capsule and narrowly conical apical lobe; 7) spermatheca sclerotized, very small with narrowly elongate capsule and with extremely long and highly coiled spermathecal duct.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52" type="description">
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<paragraph blockId="3.[140,1108,694,1672]" pageId="3" pageNumber="52">
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<emphasis bold="true" box="[185,335,976,1002]" pageId="3" pageNumber="52">
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<emphasis box="[185,330,976,1002]" inLineHeading="true" pageId="3" pageNumber="52" reason="1">Description</emphasis>
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.
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</emphasis>
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<emphasis box="[342,396,976,1002]" italics="true" pageId="3" pageNumber="52">Body</emphasis>
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(Fig. 1) slender, subparallel-sided. Body length highly variable, ranging from
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<quantity box="[297,398,1012,1038]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="3" pageNumber="52" unit="mm" value="6.0" valueMax="9.0" valueMin="3.0">3–9 mm</quantity>
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, whole body always shiny, unicoloured, or often with different colouration for head, pronotum, elytra and abdomen.
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</paragraph>
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<paragraph blockId="3.[140,1108,694,1672]" pageId="3" pageNumber="52">
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||
<emphasis box="[185,244,1082,1108]" italics="true" pageId="3" pageNumber="52">Head</emphasis>
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(Figs 2, 3) slightly wider than long, posterior margin slightly covered by anterior edge of pronotum, neck absent; temples long, round, about as long as diameter of eyes or longer; occipital suture present, not visible dorsally, ventrally reaching hypostoma, hypostoma narrow; surface with erect or appressed setae. Gula long, evenly divergent from anterior to posterior; submentum fused to gula, broadly expanded anteriorly. Eyes large, oval in lateral view, prominent.
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</paragraph>
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||
<paragraph blockId="3.[140,1108,694,1672]" pageId="3" pageNumber="52">
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||
<emphasis box="[185,288,1293,1319]" italics="true" pageId="3" pageNumber="52">Antennae</emphasis>
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(Fig. 4) with all antennomeres petiolate, antennomere III about as long as II, when bent backwards slightly exceeding base of pronotum.
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</paragraph>
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<paragraph blockId="3.[140,1108,694,1672]" pageId="3" pageNumber="52">
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||
<emphasis box="[185,274,1363,1389]" italics="true" pageId="3" pageNumber="52">Labrum</emphasis>
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(Fig. 5) much wider than long, with shallow median excavation; surface covered with numerous pseudopores and about ten real pores, except on posterior and lateral areas; antero-lateral areas each with 4–6 macrosetae of different length.
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||
</paragraph>
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||
<paragraph blockId="3.[140,1108,694,1672]" pageId="3" pageNumber="52">
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||
<emphasis box="[185,299,1469,1495]" italics="true" pageId="3" pageNumber="52">Mandibles</emphasis>
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(Figs 6–9) almost symmetrical, lacking teeth, with 3–4 small setae present dorsally, mesal areas of dorsal and ventral surfaces covered with numerous pseudopores, prostheca with inner margin pubescent.
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</paragraph>
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<paragraph blockId="3.[140,1108,694,1672]" pageId="3" pageNumber="52">
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||
<emphasis box="[185,266,1575,1601]" italics="true" pageId="3" pageNumber="52">Maxilla</emphasis>
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(Fig. 10) is generalized in shape like for other
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<taxonomicName authorityName="Fleming" authorityYear="1821" box="[754,896,1575,1601]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
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||
, with elongate galea and shortened lacinia, palpomere I minuscule, palpomere II slightly curved and gently short- er that palpomere III, terminal palpomere about three times as short as III, pointed apically.
|
||
</paragraph>
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||
<paragraph blockId="4.[140,1108,1201,1255]" pageId="4" pageNumber="53">
|
||
<emphasis bold="true" box="[140,240,1201,1223]" pageId="4" pageNumber="53">Figure I.</emphasis>
|
||
<taxonomicName authority="(Stephens)" baseAuthorityName="Stephens" baseAuthorityYear="1832" box="[247,562,1201,1223]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="haworthi" subGenus="Zyras">
|
||
<emphasis box="[247,454,1201,1223]" italics="true" pageId="4" pageNumber="53">Zyras (Zyras) haworthi</emphasis>
|
||
(Stephens)
|
||
</taxonomicName>
|
||
, dorsal view (approximative length of photograped specimen is
|
||
<quantity box="[208,283,1233,1254]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="4" pageNumber="53" unit="mm" value="7.1">7.1 mm</quantity>
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||
)
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||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,1293,1672]" pageId="4" pageNumber="53">
|
||
<emphasis box="[185,279,1294,1319]" italics="true" pageId="4" pageNumber="53">
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||
<collectingRegion box="[185,279,1294,1319]" country="Myanmar" name="Chin" pageId="4" pageNumber="53">Mentum</collectingRegion>
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||
</emphasis>
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||
(Fig. 11) trapeziform, posterior and anterior margins truncate, with setae, surface covered with pseudopores bearing minute setae, and about 6–8 pores with longer setae present on antero-lateral areas.
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,1293,1672]" pageId="4" pageNumber="53">
|
||
<emphasis box="[185,272,1399,1425]" italics="true" pageId="4" pageNumber="53">Labium</emphasis>
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||
(Fig. 12) with prementum with two real pores and one setal pore mesolaterally, with about 30–40 pseudopores behind medial setae, apodeme with large, truncate process, lateral lobes of apodeme gently curved, assimetrical, bifurcate apicaly. Ligula bilobed, each lobe long, pointed apically, with fine apical seta. Labial palpus with first segment long, more than twice as long as second and clearly thicker, third segment slightly longer than second and about twice as slender as second.
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||
</paragraph>
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<paragraph blockId="4.[140,1108,1293,1672]" pageId="4" pageNumber="53">
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||
<emphasis box="[185,293,1611,1636]" italics="true" pageId="4" pageNumber="53">Pronotum</emphasis>
|
||
slightly wider than long (Fig. 1), anterior margin straight, posterior margin evenly rounded, posterior corners small but well-defined, smooth or entirely or
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |