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<document ID-DOI="10.5281/zenodo.272951" ID-GBIF-Dataset="e2c88f4c-3ac2-45f9-95e4-99b75561a081" ID-ISSN="1175-5326" ID-Zenodo-Dep="272951" ID-ZooBank="6D68A019-6F63-4AA4-A8B3-92D351F1F69B" checkinTime="1484409208030" checkinUser="plazi" docAuthor="Van, Rob W. M." docDate="2017" docId="03A8001077E2FF1AFF14A6749084F8D0" docLanguage="en" docName="zootaxa.4217.1.1.pdf.imf" docOrigin="Zootaxa 1" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5ce3343f" docStyleName="zootaxa.2013.monograph" docTitle="Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu 2003" docType="treatment" docVersion="19" lastPageNumber="183" masterDocId="FF9178687757FFACFF83A36B907DFF9A" masterDocTitle="Sponges of the Guyana Shelf" masterLastPageNumber="225" masterPageNumber="1" pageNumber="182" updateTime="1645784257244" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Sponges of the Guyana Shelf</mods:title>
</mods:titleInfo>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Van, Rob W. M.</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2017</mods:date>
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<mods:number>1</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3510822" ID-GBIF-Taxon="127419808" ID-Zenodo-Dep="3510822" LSID="urn:lsid:plazi:treatment:03A8001077E2FF1AFF14A6749084F8D0" httpUri="http://treatment.plazi.org/id/03A8001077E2FF1AFF14A6749084F8D0" lastPageId="182" lastPageNumber="183" pageId="181" pageNumber="182">
<subSubSection pageId="181" pageNumber="182" type="nomenclature">
<paragraph blockId="181.[151,876,1311,1371]" box="[151,876,1311,1337]" pageId="181" pageNumber="182">
<heading bold="true" box="[151,876,1311,1337]" fontSize="11" level="1" pageId="181" pageNumber="182" reason="1">
<taxonomicName authority="Carvalho, Carraro, Lerner &amp; Hajdu, 2003" authorityName="Carvalho, Carraro, Lerner &amp; Hajdu" authorityYear="2003" box="[151,876,1311,1337]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="181" pageNumber="182" phylum="Porifera" rank="species" species="alba">
<emphasis bold="true" box="[151,876,1311,1337]" pageId="181" pageNumber="182">
<emphasis bold="true" box="[151,338,1311,1337]" italics="true" pageId="181" pageNumber="182">Ciocalypta alba</emphasis>
<bibRefCitation author="Carvalho" box="[346,876,1311,1337]" pageId="181" pageNumber="182" refString="Carvalho, M. S., Carraro, J. L., Lerner, C. &amp; Hajdu, E. (2003) First record of Ciocalypta (Demospongiae: Halichondrida) from Brazil, southwestern Atlantic, with the description of a new species. Zootaxa, 302, 1 - 8." type="journal article" year="2003">Carvalho, Carraro, Lerner &amp; Hajdu, 2003</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="181.[151,876,1311,1371]" box="[151,319,1346,1371]" pageId="181" pageNumber="182">
<figureCitation box="[151,282,1346,1371]" captionStart="FIGURE 114" captionStartId="182.[151,250,1323,1345]" captionTargetBox="[151,1436,193,1301]" captionTargetId="figure@182.[151,1436,193,1301]" captionTargetPageId="182" captionText="FIGURE 114. Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu, 2003, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, d f, SEM images of the spicules, d, large oxea, d 1, details of d, e, middle-sized oxea, e 1, details of e, f, small oxea, f 1, details of f." httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="181" pageNumber="182">Figures 114</figureCitation>
af
</paragraph>
</subSubSection>
<subSubSection box="[151,660,1418,1440]" pageId="181" pageNumber="182" type="reference_group">
<paragraph blockId="181.[151,660,1418,1440]" box="[151,660,1418,1440]" pageId="181" pageNumber="182">
<treatmentCitationGroup box="[151,660,1418,1440]" pageId="181" pageNumber="182">
<taxonomicName authority="Carvalho et al., 2003: 2" authorityName="Carvalho et al." authorityPageNumber="2" authorityYear="2003" box="[151,560,1418,1440]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="181" pageNumber="182" phylum="Porifera" rank="species" species="alba">
<emphasis box="[151,313,1418,1440]" italics="true" pageId="181" pageNumber="182">Ciocalypta alba</emphasis>
<treatmentCitation ID-DOI="http://doi.org/10.5281/zenodo.6276765" ID-Zenodo-Dep="6276765" author="Carvalho" box="[320,560,1418,1440]" httpUri="http://treatment.plazi.org/id/03855567400AFFA0D8277406698E75D8" page="2" pageId="181" pageNumber="182" year="2003">
<bibRefCitation author="Carvalho" box="[320,536,1418,1440]" pageId="181" pageNumber="182" refString="Carvalho, M. S., Carraro, J. L., Lerner, C. &amp; Hajdu, E. (2003) First record of Ciocalypta (Demospongiae: Halichondrida) from Brazil, southwestern Atlantic, with the description of a new species. Zootaxa, 302, 1 - 8." type="journal article" year="2003">
Carvalho
<emphasis box="[419,470,1418,1440]" italics="true" pageId="181" pageNumber="182">et al.</emphasis>
, 2003
</bibRefCitation>
: 2
</treatmentCitation>
</taxonomicName>
, figs 12.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="181" pageNumber="182" type="materials_examined">
<paragraph blockId="181.[151,1437,1484,2013]" pageId="181" pageNumber="182">
<materialsCitation ID-GBIF-Occurrence="1424726542" collectingDate="1970-08-31" collectionCode="RMNH" country="Guyana" elevation="71" latitude="7.5833" location="Material" longLatPrecision="7" longitude="-57.0667" pageId="181" pageNumber="182" specimenCount="1">
<emphasis bold="true" box="[151,392,1484,1509]" pageId="181" pageNumber="182">
<location LSID="urn:lsid:plazi:treatment:03A8001077E2FF1AFF14A6749084F8D0:8EDEE7DD77E2FF19FF14A6A79082FA7F" box="[151,255,1484,1509]" country="Guyana" latitude="7.5833" longLatPrecision="7" longitude="-57.0667" name="Material" pageId="181" pageNumber="182">Material</location>
examined.
</emphasis>
<collectionCode box="[405,489,1485,1508]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="181" pageNumber="182">RMNH</collectionCode>
Por. 9938,
<collectingCountry box="[639,729,1485,1509]" name="Guyana" pageId="181" pageNumber="182">Guyana</collectingCountry>
, Luymes
<collectingCountry box="[871,958,1485,1509]" name="Guyana" pageId="181" pageNumber="182">Guyana</collectingCountry>
Shelf Expedition, station 63,
<geoCoordinate box="[1328,1436,1484,1509]" direction="north" orientation="latitude" pageId="181" pageNumber="182" precision="5" value="7.5833">7.5833°N</geoCoordinate>
<geoCoordinate box="[151,279,1520,1545]" direction="west" orientation="longitude" pageId="181" pageNumber="182" precision="5" value="-57.0667">57.0667°W</geoCoordinate>
, depth
<quantity box="[360,419,1521,1545]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="181" pageNumber="182" unit="m" value="71.0">
<elevation box="[360,419,1521,1545]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="181" pageNumber="182" unit="m" value="71.0">71 m</elevation>
</quantity>
, sandy bottom,
<date box="[597,780,1521,1545]" pageId="181" pageNumber="182" value="1970-08-31">
<collectingDate box="[597,780,1521,1545]" pageId="181" pageNumber="182" value="1970-08-31">31 August 1970</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="181" pageNumber="182" type="description">
<paragraph blockId="181.[151,1437,1484,2013]" pageId="181" pageNumber="182">
<emphasis bold="true" box="[199,345,1556,1581]" pageId="181" pageNumber="182">Description.</emphasis>
(
<figureCitation box="[368,471,1556,1581]" captionStart="FIGURE 114" captionStartId="182.[151,250,1323,1345]" captionTargetBox="[151,1436,193,1301]" captionTargetId="figure@182.[151,1436,193,1301]" captionTargetPageId="182" captionText="FIGURE 114. Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu, 2003, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, d f, SEM images of the spicules, d, large oxea, d 1, details of d, e, middle-sized oxea, e 1, details of e, f, small oxea, f 1, details of f." httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="181" pageNumber="182">Fig. 114</figureCitation>
a) A single fistule of about
<quantity box="[829,914,1556,1581]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="181" pageNumber="182" unit="cm" value="2.5">2.5 cm</quantity>
long,
<quantity box="[1003,1090,1556,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="181" pageNumber="182" unit="mm" value="15.0">15 mm</quantity>
in diameter, encrusting and consolidating several dead mollusk shells. The shape is equidiametrical ending in a short closed pointed apex. The smooth skin is glassy grey, transparent. Inside a darker colored central axis is visible, surrounded by large subdermal spaces. Consistency elastic but fragile.
</paragraph>
<paragraph blockId="181.[151,1437,1484,2013]" pageId="181" pageNumber="182">
<emphasis bold="true" box="[199,309,1700,1725]" pageId="181" pageNumber="182">Skeleton.</emphasis>
The ectosomal skeleton (
<figureCitation box="[607,705,1700,1725]" captionStart="FIGURE 114" captionStartId="182.[151,250,1323,1345]" captionTargetBox="[151,1436,193,1301]" captionTargetId="figure@182.[151,1436,193,1301]" captionTargetPageId="182" captionText="FIGURE 114. Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu, 2003, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, d f, SEM images of the spicules, d, large oxea, d 1, details of d, e, middle-sized oxea, e 1, details of e, f, small oxea, f 1, details of f." httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="181" pageNumber="182">Fig. 114</figureCitation>
bc) is a tangential halichondrioid reticulation of spicule tracts, 27 spicules in cross section, intercrossing and anastomosing. The central axis is a thick aligned mass of oxeas, giving off side branches running to the surface, supporting the ectosomal skeleton.
</paragraph>
<paragraph blockId="181.[151,1437,1484,2013]" box="[199,603,1808,1833]" pageId="181" pageNumber="182">
<emphasis bold="true" box="[199,304,1808,1833]" pageId="181" pageNumber="182">Spicules.</emphasis>
(
<figureCitation box="[319,415,1808,1833]" captionStart="FIGURE 114" captionStartId="182.[151,250,1323,1345]" captionTargetBox="[151,1436,193,1301]" captionTargetId="figure@182.[151,1436,193,1301]" captionTargetPageId="182" captionText="FIGURE 114. Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu, 2003, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, d f, SEM images of the spicules, d, large oxea, d 1, details of d, e, middle-sized oxea, e 1, details of e, f, small oxea, f 1, details of f." httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="181" pageNumber="182">Figs 114</figureCitation>
df) Oxeas only.
</paragraph>
<paragraph blockId="181.[151,1437,1484,2013]" pageId="181" pageNumber="182">
Oxeas, symmetrical, predominantly slightly curved, sharply pointed, in a large size range, divisible in three almost overlapping size categories, (1) large (
<figureCitation box="[663,767,1880,1905]" captionStart="FIGURE 114" captionStartId="182.[151,250,1323,1345]" captionTargetBox="[151,1436,193,1301]" captionTargetId="figure@182.[151,1436,193,1301]" captionTargetPageId="182" captionText="FIGURE 114. Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu, 2003, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, d f, SEM images of the spicules, d, large oxea, d 1, details of d, e, middle-sized oxea, e 1, details of e, f, small oxea, f 1, details of f." httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="181" pageNumber="182">Figs 114</figureCitation>
d,d1), 576816
<date box="[939,1016,1881,1905]" pageId="181" pageNumber="182" value="1921-10-09">x 921</date>
µm, (2) middle-sized (
<figureCitation box="[1276,1380,1880,1905]" captionStart="FIGURE 114" captionStartId="182.[151,250,1323,1345]" captionTargetBox="[151,1436,193,1301]" captionTargetId="figure@182.[151,1436,193,1301]" captionTargetPageId="182" captionText="FIGURE 114. Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu, 2003, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, d f, SEM images of the spicules, d, large oxea, d 1, details of d, e, middle-sized oxea, e 1, details of e, f, small oxea, f 1, details of f." httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="181" pageNumber="182">Figs 114</figureCitation>
e,e1), 316512 x 6.510 µm, and (3) small (
<figureCitation box="[571,673,1916,1941]" captionStart="FIGURE 114" captionStartId="182.[151,250,1323,1345]" captionTargetBox="[151,1436,193,1301]" captionTargetId="figure@182.[151,1436,193,1301]" captionTargetPageId="182" captionText="FIGURE 114. Ciocalypta alba Carvalho, Carraro, Lerner &amp; Hajdu, 2003, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, d f, SEM images of the spicules, d, large oxea, d 1, details of d, e, middle-sized oxea, e 1, details of e, f, small oxea, f 1, details of f." httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="181" pageNumber="182">Figs 114</figureCitation>
f,f1), 182308
<date box="[836,898,1917,1941]" pageId="181" pageNumber="182">x 47</date>
µm; overall oxea size 182
<emphasis box="[1206,1248,1917,1941]" italics="true" pageId="181" pageNumber="182">453</emphasis>
816
<date box="[1310,1344,1917,1941]" pageId="181" pageNumber="182">x 4</date>
<emphasis box="[1359,1394,1916,1941]" italics="true" pageId="181" pageNumber="182">9.6</emphasis>
21 µm.
</paragraph>
</subSubSection>
<subSubSection box="[199,1185,1988,2013]" pageId="181" pageNumber="182" type="distribution">
<paragraph blockId="181.[151,1437,1484,2013]" box="[199,1185,1988,2013]" pageId="181" pageNumber="182">
<emphasis bold="true" box="[199,498,1988,2013]" pageId="181" pageNumber="182">Distribution and ecology.</emphasis>
<collectingCountry box="[505,592,1989,2013]" name="Guyana" pageId="181" pageNumber="182">Guyana</collectingCountry>
Shelf, SE
<collectingCountry box="[712,783,1988,2013]" name="Brazil" pageId="181" pageNumber="182">Brazil</collectingCountry>
, in sandy bottom at
<quantity box="[1009,1108,1989,2013]" metricMagnitude="1" metricUnit="m" metricValue="4.15" metricValueMax="7.1" metricValueMin="1.2" pageId="181" pageNumber="182" unit="m" value="41.5" valueMax="71.0" valueMin="12.0">1271 m</quantity>
depth.
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/273064/files/figure.png" pageId="182" pageNumber="183" targetBox="[151,1436,193,1301]" targetPageId="182">
<paragraph blockId="182.[151,1436,1323,1406]" pageId="182" pageNumber="183">
<emphasis bold="true" box="[151,299,1323,1345]" pageId="182" pageNumber="183">FIGURE 114.</emphasis>
<taxonomicName authority="Carvalho, Carraro, Lerner &amp; Hajdu, 2003" authorityName="Carvalho, Carraro, Lerner &amp; Hajdu" authorityYear="2003" box="[305,895,1323,1345]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="alba">
<taxonomicName authority="Carvalho, Carraro, Lerner &amp; Hajdu" authorityName="Carvalho, Carraro, Lerner &amp; Hajdu" box="[305,833,1323,1345]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="alba">
<taxonomicName authority="Carvalho, Carraro, Lerner" authorityName="Carvalho, Carraro, Lerner" box="[305,735,1323,1345]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="alba">
<taxonomicName authority="Carvalho, Carraro" authorityName="Carvalho, Carraro" box="[305,659,1323,1345]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="alba">
<taxonomicName authority="Carvalho" authorityName="Carvalho" box="[305,568,1323,1345]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="alba">
<taxonomicName box="[305,467,1323,1345]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="alba">
<taxonomicName box="[305,416,1323,1345]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="genus">
<emphasis box="[305,416,1323,1345]" italics="true" pageId="182" pageNumber="183">Ciocalypta</emphasis>
</taxonomicName>
<taxonomicName authority="Carvalho" authorityName="Carvalho" authorityYear="2003" box="[422,467,1323,1345]" pageId="182" pageNumber="183" rank="species" species="alba">
<emphasis box="[422,467,1323,1345]" italics="true" pageId="182" pageNumber="183">alba</emphasis>
</taxonomicName>
</taxonomicName>
Carvalho
</taxonomicName>
, Carraro
</taxonomicName>
, Lerner
</taxonomicName>
&amp; Hajdu
</taxonomicName>
, 2003
</taxonomicName>
, a, habitus of RMNH Por. 9938 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c, detail of b, df, SEM images of the spicules, d, large oxea, d1, details of d, e, middle-sized oxea, e1, details of e, f, small oxea, f1, details of f.
</paragraph>
</caption>
<subSubSection pageId="182" pageNumber="183" type="discussion">
<paragraph blockId="182.[151,1436,1445,1866]" pageId="182" pageNumber="183">
<emphasis bold="true" box="[199,315,1445,1470]" pageId="182" pageNumber="183">Remarks.</emphasis>
The single fistule matches all details described by
<bibRefCitation author="Carvalho" box="[898,1134,1445,1470]" pageId="182" pageNumber="183" refString="Carvalho, M. S., Carraro, J. L., Lerner, C. &amp; Hajdu, E. (2003) First record of Ciocalypta (Demospongiae: Halichondrida) from Brazil, southwestern Atlantic, with the description of a new species. Zootaxa, 302, 1 - 8." type="journal article" year="2003">
Carvalho
<emphasis box="[1008,1066,1445,1470]" italics="true" pageId="182" pageNumber="183">et al.</emphasis>
2003
</bibRefCitation>
: the transparent skin, the central axis of oxeas and the tangential halichondroid skeleton at the surface, and the sizes and categories of the oxeas. The
<typeStatus box="[279,327,1519,1543]" pageId="182" pageNumber="183">type</typeStatus>
material is much bigger, with dozens of fistules, but the present individual fistule is closely similar to one of the fistules of the
<typeStatus box="[458,510,1554,1578]" pageId="182" pageNumber="183">type</typeStatus>
.
</paragraph>
<paragraph blockId="182.[151,1436,1445,1866]" pageId="182" pageNumber="183">
Another
<collectingRegion box="[302,385,1589,1614]" country="Sri Lanka" name="Central" pageId="182" pageNumber="183">Central</collectingRegion>
<collectingRegion box="[396,451,1591,1614]" country="Tanzania" name="Mjini Magharibi" pageId="182" pageNumber="183">West</collectingRegion>
Atlantic
<taxonomicName box="[564,687,1589,1614]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="genus">
<emphasis box="[564,687,1589,1614]" italics="true" pageId="182" pageNumber="183">Ciocalypta</emphasis>
</taxonomicName>
species is
<taxonomicName authority="Wells, Wells &amp; Gray, 1960" authorityName="Wells, Wells &amp; Gray" authorityYear="1960" box="[820,1272,1589,1614]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="gibbsi">
<emphasis box="[820,925,1589,1614]" italics="true" pageId="182" pageNumber="183">C. gibbsi</emphasis>
(
<bibRefCitation author="Wells" box="[944,1264,1589,1614]" pageId="182" pageNumber="183" refString="Wells, H. W., Wells, M. J. &amp; Gray, I. E. (1960) Marine sponges of North Carolina. Journal of the Elisha Mitchell Scientific Society, 76 (2), 200 - 245." type="journal article" year="1960">Wells, Wells &amp; Gray, 1960</bibRefCitation>
)
</taxonomicName>
(originally as
<taxonomicName box="[151,290,1625,1650]" class="Demospongiae" family="Halichondriidae" genus="Ciocalapata" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="genus">
<emphasis box="[151,290,1625,1650]" italics="true" pageId="182" pageNumber="183">Ciocalapata</emphasis>
</taxonomicName>
), but this has distinctly smaller and thinner spicules.
<bibRefCitation author="Carvalho" box="[917,1155,1625,1650]" pageId="182" pageNumber="183" refString="Carvalho, M. S., Carraro, J. L., Lerner, C. &amp; Hajdu, E. (2003) First record of Ciocalypta (Demospongiae: Halichondrida) from Brazil, southwestern Atlantic, with the description of a new species. Zootaxa, 302, 1 - 8." type="journal article" year="2003">
Carvalho
<emphasis box="[1029,1089,1625,1650]" italics="true" pageId="182" pageNumber="183">et al.</emphasis>
2003
</bibRefCitation>
also assigned
<taxonomicName authority="Diaz, Pomponi &amp; Van Soest, 1993" authorityName="Diaz, Pomponi &amp; Van Soest" authorityYear="1993" class="Demospongiae" family="Halichondriidae" genus="Topsentia" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="pseudoporrecta">
<emphasis italics="true" pageId="182" pageNumber="183">Topsentia pseudoporrecta</emphasis>
<bibRefCitation author="Diaz" box="[340,761,1661,1686]" pageId="182" pageNumber="183" refString="Diaz, M. C., Pomponi, S. A. &amp; Van Soest, R. W. M. (1993) A systematic revision of the central West Atlantic Halichondrida (Demospongiae, Porifera). Part III: Description of valid species. In: Uriz, M. - J. &amp; Rutzler, K. (Eds.), Recent Advances in Ecology and Systematics of Sponges. Scientia Marina, 57 (4), 283 - 306." type="journal article" year="1993">Díaz, Pomponi &amp; Van Soest, 1993</bibRefCitation>
</taxonomicName>
to
<taxonomicName box="[811,934,1661,1686]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="genus">
<emphasis box="[811,934,1661,1686]" italics="true" pageId="182" pageNumber="183">Ciocalypta</emphasis>
</taxonomicName>
, but since its fistules are solid and nontransparent, I do not think this is warranted. It differs also clearly from
<taxonomicName box="[955,1038,1697,1722]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="alba">
<emphasis box="[955,1038,1697,1722]" italics="true" pageId="182" pageNumber="183">C. alba</emphasis>
</taxonomicName>
and
<taxonomicName box="[1094,1195,1697,1722]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="gibbsi">
<emphasis box="[1094,1195,1697,1722]" italics="true" pageId="182" pageNumber="183">C. gibbsi</emphasis>
</taxonomicName>
in having very much larger oxeas (up to 1800 µm). The
<typeStatus box="[544,592,1735,1759]" pageId="182" pageNumber="183">type</typeStatus>
species of
<taxonomicName box="[719,842,1733,1758]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="genus">
<emphasis box="[719,842,1733,1758]" italics="true" pageId="182" pageNumber="183">Ciocalypta</emphasis>
</taxonomicName>
, European
<taxonomicName authority="Bowerbank, 1862" authorityName="Bowerbank" authorityYear="1862" box="[971,1320,1733,1758]" class="Demospongiae" family="Halichondriidae" genus="Ciocalypta" kingdom="Animalia" order="Halichondrida" pageId="182" pageNumber="183" phylum="Porifera" rank="species" species="penicillus">
<emphasis box="[971,1113,1733,1758]" italics="true" pageId="182" pageNumber="183">C. penicillus</emphasis>
<bibRefCitation author="Bowerbank" box="[1120,1320,1733,1758]" pageId="182" pageNumber="183" refString="Bowerbank, J. S. (1862) On the anatomy and physiology of the Spongiadae. Part II. Philosophical Transactions of the Royal Society, 152 (2), 747 - 829." type="journal article" year="1862">Bowerbank, 1862</bibRefCitation>
</taxonomicName>
has styles as the dominant spicule
<typeStatus box="[441,492,1770,1794]" pageId="182" pageNumber="183">type</typeStatus>
, but like
<bibRefCitation box="[610,871,1769,1794]" pageId="182" pageNumber="183" refString="Carvalho, M. S., Carraro, J. L., Lerner, C. &amp; Hajdu, E. (2003) First record of Ciocalypta (Demospongiae: Halichondrida) from Brazil, southwestern Atlantic, with the description of a new species. Zootaxa, 302, 1 - 8." type="journal article">
Carvalho
<emphasis box="[724,785,1769,1794]" italics="true" pageId="182" pageNumber="183">et al.</emphasis>
(2003)
</bibRefCitation>
I follow here Erpenbeck &amp; Van Soests (2002) argument that the genus is characterized by the peculiar fistular shape and species may have oxeas, styles or both as spicules.
</paragraph>
</subSubSection>
</treatment>
</document>