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<document id="A03DFE6CB1EB522DB2447FDA9B854126" ID="10.11646/zootaxa.4169.2.1" ID-DOI="10.11646/zootaxa.4169.2.1" ID-GBIF-Dataset="d1fbe4b3-de10-4d72-a117-aa6bfedf210d" ID-ISSN="1175-5326" ID-Zenodo-Dep="262585" ID-ZooBank="B8B067BF-F6E6-4122-B884-AA385FF04421" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474351831659" checkinUser="plazi" docAuthor="Andrew Hamilton, K. G." docDate="2016" docId="03A5742DFFDAAE05FF76AB38CEFFFAC4" docLanguage="en" docName="zootaxa.4169.2.1.pdf" docOrigin="Zootaxa 4169 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microsargane (Microtholia) habrotes Hamilton, 2016, sp.nov." docType="treatment" docVersion="7" lastPageNumber="216" masterDocId="FF9C0C55FFD5AE0AFFE1A955CD70FFD1" masterDocTitle="Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae" masterLastPageNumber="250" masterPageNumber="201" pageNumber="216" updateTime="1698386375821" updateUser="plazi">
<mods:mods id="BED1C76F8C71ABEE161727E6C3A50816" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="09016E6EFB4B0621FF89CCC584D0B755">Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae</mods:title>
</mods:titleInfo>
<mods:name id="305BC5176B9F2DEBC44FD503D52931B9" type="personal">
<mods:role id="01E94CDE22571378E0929EAC11FF7ECD">
<mods:roleTerm id="D772B1FE0B7220F2EEBE3920ECB4B8B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9476F9585315C01837EB4BCFFAB56DA">Andrew Hamilton, K. G.</mods:namePart>
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<mods:typeOfResource id="2A628B527C0CE76C3939981DC6E149F7">text</mods:typeOfResource>
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<mods:title id="893FED6EB2D20F1E39A5FB49C97CCF78">Zootaxa</mods:title>
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<mods:part id="2894CF25DBAD0609CDF860EC311AB775">
<mods:date id="BE381D29015C468FE7310C5DA15056D1">2016</mods:date>
<mods:detail id="859DF13A1BE76001791FDB41E650E767" type="volume">
<mods:number id="7B0F852D9DE242C75706F3AF1D4D5D8B">4169</mods:number>
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<mods:start id="DAB6DD70B1E9C1FA6D53FD3A6C4FE80F">201</mods:start>
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<mods:classification id="65988082FE834A03191FA3590AA92953">journal article</mods:classification>
<mods:identifier id="1767E2E103C154E45EDA3E706B1F0E76" type="DOI">10.11646/zootaxa.4169.2.1</mods:identifier>
<mods:identifier id="FDC76477938FF5E52C44B28A2FC59D42" type="GBIF-Dataset">d1fbe4b3-de10-4d72-a117-aa6bfedf210d</mods:identifier>
<mods:identifier id="41AF629F14471E24210B67FEB13838EB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="269B7C44268246C8CDF38A95BCA160F5" type="Zenodo-Dep">262585</mods:identifier>
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<treatment id="03A5742DFFDAAE05FF76AB38CEFFFAC4" ID-DOI="http://doi.org/10.5281/zenodo.5613341" ID-GBIF-Taxon="124552086" ID-Zenodo-Dep="5613341" LSID="urn:lsid:plazi:treatment:03A5742DFFDAAE05FF76AB38CEFFFAC4" httpUri="http://treatment.plazi.org/id/03A5742DFFDAAE05FF76AB38CEFFFAC4" lastPageNumber="216" pageId="15" pageNumber="216">
<subSubSection id="C31696B0FFDAAE05FF76AB38CFC7FD56" box="[151,695,621,647]" pageId="15" pageNumber="216" type="nomenclature">
<paragraph id="8BB3C53BFFDAAE05FF76AB38CFC7FD56" blockId="15.[151,695,621,647]" box="[151,695,621,647]" pageId="15" pageNumber="216">
<heading id="D0FB7257FFDAAE05FF76AB38CFC7FD56" bold="true" box="[151,695,621,647]" fontSize="11" level="1" pageId="15" pageNumber="216" reason="1">
<emphasis id="B9781929FFDAAE05FF76AB38CFC7FD56" bold="true" box="[151,695,621,647]" pageId="15" pageNumber="216">
<taxonomicName id="4C0CBEB8FFDAAE05FF76AB38CF2AFD56" box="[151,602,621,647]" class="Insecta" family="Aphrophoridae" genus="Microsargane" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="habrotes" status="sp. nov." subGenus="Microtholia">
<emphasis id="B9781929FFDAAE05FF76AB38CC30FD56" bold="true" box="[151,320,621,647]" italics="true" pageId="15" pageNumber="216">Microsargane</emphasis>
(
<emphasis id="B9781929FFDAAE05FEB0AB38CC90FD56" bold="true" box="[337,480,621,647]" italics="true" pageId="15" pageNumber="216">Microtholia</emphasis>
)
<emphasis id="B9781929FFDAAE05FE13AB38CF2AFD56" bold="true" box="[498,602,621,647]" italics="true" pageId="15" pageNumber="216">habrotes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFDAAE05FD80AB38CFC7FD56" box="[609,695,621,647]" pageId="15" pageNumber="216" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFDAAE05FF76ABE1C94FFD1C" box="[151,1087,692,717]" pageId="15" pageNumber="216" type="etymology">
<paragraph id="8BB3C53BFFDAAE05FF76ABE1C94FFD1C" blockId="15.[151,1437,692,1301]" box="[151,1087,692,717]" pageId="15" pageNumber="216">
<emphasis id="B9781929FFDAAE05FF76ABE1CC68FD1C" bold="true" box="[151,280,692,717]" pageId="15" pageNumber="216">Etymology</emphasis>
<emphasis id="B9781929FFDAAE05FEFEABE1CCC0FD1D" box="[287,432,692,717]" italics="true" pageId="15" pageNumber="216">habros, -otes</emphasis>
(adjective) pretty; the most colorful of the known species.
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFDAAE05FF26AB8DCD9FFC8E" pageId="15" pageNumber="216" type="diagnosis">
<paragraph id="8BB3C53BFFDAAE05FF26AB8DCD9FFC8E" blockId="15.[151,1437,692,1301]" pageId="15" pageNumber="216">
<emphasis id="B9781929FFDAAE05FF26AB8DCC31FD20" bold="true" box="[199,321,728,753]" pageId="15" pageNumber="216">Diagnosis.</emphasis>
Redbrown, paler on venter, tegminal tips suffused with red in male, marked with yellow midline of dorsum and two tegminal stripes, one extending from sides of pronotum along claval suture, and another extending from base of costa to midlength of central anteapical cell; 2 contrasting black spots on 1st and 2nd apical cells (
<figureCitation id="1337D9BEFFDAAE05FA8BAA76CDD5FC8E" captionStart="FIGURES 9" captionStartId="12.[151,264,1652,1674]" captionTargetBox="[169,1435,208,1618]" captionTargetId="figure@12.[167,1452,195,1619]" captionTargetPageId="12" captionText="FIGURES 9. New species of Microsargane subgenera Microtholia subgen. nov. (A – G) and Microrhaphe subgen. nov. (H – K): A, M. ios sp. nov.; B, M. karykroua sp. nov., male; C, same, female; D, M. bipunctura sp. nov.; E, M. sobria sp. nov.; F, M. habrote sp. nov. s, male; G, same, female; H – K, M. martialis sp. nov., color varieties." httpUri="https://zenodo.org/record/262587/files/figure.png" pageId="15" pageNumber="216">Figs 9</figureCitation>
F–G).
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFDAAE05FF26AA3ECEFFFAC4" pageId="15" pageNumber="216" type="description">
<paragraph id="8BB3C53BFFDAAE05FF26AA3ECCE4FBAE" blockId="15.[151,1437,692,1301]" pageId="15" pageNumber="216">
<emphasis id="B9781929FFDAAE05FF26AA3ECC29FC55" bold="true" box="[199,345,875,900]" pageId="15" pageNumber="216">Description.</emphasis>
Venter marked only with narrow black lines across frontal arcs and dark tarsi; legs and genital capsule golden yellow. Coronal margin carinate at antennal ledge, becoming only angular on tylus (first muscle arc of face visible from dorsal aspect). Male paler than female, with red on tegmina confined to veins outlining apical cells. Male subgenital plates as wide at tip as at base, mesally concave and laterally notched just beyond short, thumblike lateral processes; style stout, tapered to bluntly pointed tip bearing sharp preapical spine and short setae basad of this on dorsal edge (
<figureCitation id="1337D9BEFFDAAE05FE3EAD4BCF42FBE6" box="[479,562,1054,1079]" captionStart="FIGURES 55 – 60" captionStartId="41.[151,264,1837,1859]" captionTargetBox="[199,1382,198,1775]" captionTargetId="figure@41.[187,1392,193,1791]" captionTargetPageId="41" captionText="FIGURES 55 – 60. Male genitalia of Microsargane new subgenera Microrhaphe (55) and Microtholia (56 – 58) compared to Microclimax gen. nov. (59): A, genital capsule, lateral aspect; B, same, theca and phallobase; C, theca, caudal aspect; D, fused subgenital plates; E, pygofer process, dorsal aspect; F, style tip, dorsal aspect). 55, M. martialis; 56, M. sobria sp. nov.; 57, M. karykroua sp. nov.; 58, M. habrotes sp. nov. (genital capsule as in M. karykroua); 59, M. ios (genital capsule, style and theca as in M. habrotes); 60, Microclimax luteosignata (Valdés Ragués) showing entire genital capsule (D) in ventral aspect at smaller scale." httpUri="https://zenodo.org/record/262598/files/figure.png" pageId="15" pageNumber="216">Fig. 58</figureCitation>
E); thecal shaft broad, 4 × as long as wide in lateral aspect (
<figureCitation id="1337D9BEFFDAAE05FB31AD4BC853FBE6" box="[1232,1315,1054,1079]" captionStart="FIGURES 55 – 60" captionStartId="41.[151,264,1837,1859]" captionTargetBox="[199,1382,198,1775]" captionTargetId="figure@41.[187,1392,193,1791]" captionTargetPageId="41" captionText="FIGURES 55 – 60. Male genitalia of Microsargane new subgenera Microrhaphe (55) and Microtholia (56 – 58) compared to Microclimax gen. nov. (59): A, genital capsule, lateral aspect; B, same, theca and phallobase; C, theca, caudal aspect; D, fused subgenital plates; E, pygofer process, dorsal aspect; F, style tip, dorsal aspect). 55, M. martialis; 56, M. sobria sp. nov.; 57, M. karykroua sp. nov.; 58, M. habrotes sp. nov. (genital capsule as in M. karykroua); 59, M. ios (genital capsule, style and theca as in M. habrotes); 60, Microclimax luteosignata (Valdés Ragués) showing entire genital capsule (D) in ventral aspect at smaller scale." httpUri="https://zenodo.org/record/262598/files/figure.png" pageId="15" pageNumber="216">Fig. 58</figureCitation>
B), 12 × as long as wide in caudal aspect, bearing 5–6 pairs of filiform appendages (
<figureCitation id="1337D9BEFFDAAE05FC3FAD16C944FB8D" box="[990,1076,1091,1116]" captionStart="FIGURES 55 – 60" captionStartId="41.[151,264,1837,1859]" captionTargetBox="[199,1382,198,1775]" captionTargetId="figure@41.[187,1392,193,1791]" captionTargetPageId="41" captionText="FIGURES 55 – 60. Male genitalia of Microsargane new subgenera Microrhaphe (55) and Microtholia (56 – 58) compared to Microclimax gen. nov. (59): A, genital capsule, lateral aspect; B, same, theca and phallobase; C, theca, caudal aspect; D, fused subgenital plates; E, pygofer process, dorsal aspect; F, style tip, dorsal aspect). 55, M. martialis; 56, M. sobria sp. nov.; 57, M. karykroua sp. nov.; 58, M. habrotes sp. nov. (genital capsule as in M. karykroua); 59, M. ios (genital capsule, style and theca as in M. habrotes); 60, Microclimax luteosignata (Valdés Ragués) showing entire genital capsule (D) in ventral aspect at smaller scale." httpUri="https://zenodo.org/record/262598/files/figure.png" pageId="15" pageNumber="216">Fig. 58</figureCitation>
C). Length: male 9.0–
<quantity id="4CF468DEFFDAAE05FAD1AD16C8E8FB8A" box="[1328,1432,1091,1115]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="15" pageNumber="216" unit="mm" value="10.4">10.4 mm</quantity>
; female 10.0–
<quantity id="4CF468DEFFDAAE05FECBAD32CCE0FBAE" box="[298,400,1127,1151]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="15" pageNumber="216" unit="mm" value="10.7">10.7 mm</quantity>
.
</paragraph>
<paragraph id="8BB3C53BFFDAAE05FF26ADD8CEFFFAC4" blockId="15.[151,1437,692,1301]" pageId="15" pageNumber="216">
<materialsCitation id="3B64CF66FFDAAE05FF26ADD8CF18FB77" ID-GBIF-Occurrence="1313797436" box="[199,616,1165,1190]" country="Panama" location="PANAMA" pageId="15" pageNumber="216" specimenCount="2" typeStatus="holotype">
<emphasis id="B9781929FFDAAE05FF26ADD8CC63FB77" bold="true" box="[199,275,1165,1190]" pageId="15" pageNumber="216">
<typeStatus id="54B77B99FFDAAE05FF26ADD8CC7FFB77" box="[199,271,1165,1190]" pageId="15" pageNumber="216">Types</typeStatus>
.
</emphasis>
<typeStatus id="54B77B99FFDAAE05FEC5ADD8CCFCFB77" box="[292,396,1165,1190]" pageId="15" pageNumber="216" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F31B85ABFFDAAE05FE0DADDBCF18FB77" box="[492,616,1166,1190]" name="Panama" pageId="15" pageNumber="216">PANAMA</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B64CF66FFDAAE05FD9AADD8CC77FB1B" ID-GBIF-Occurrence="1313797430" collectingDate="1978-05-17" collectorName="O'Brien &amp; Marshall" country="Panama" latitude="8.733334" location="Fortuna" longLatPrecision="1300" longitude="-82.25" municipality="La" pageId="15" pageNumber="216" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<emphasis id="B9781929FFDAAE05FD9AADD8CF9EFB77" box="[635,750,1165,1190]" italics="true" pageId="15" pageNumber="216">
<collectingRegion id="49C80BD9FFDAAE05FD9AADD8CFA9FB77" box="[635,729,1165,1190]" country="Panama" name="Chiriqui" pageId="15" pageNumber="216">Chiriqui</collectingRegion>
─
</emphasis>
[
<collectingMunicipality id="6BD75F41FFDAAE05FD15ADDBCE63FB74" box="[756,787,1166,1189]" pageId="15" pageNumber="216">La</collectingMunicipality>
]
<location id="8ED393E0FFDAAE05FCF8ADDBCE05FB74" LSID="urn:lsid:plazi:treatment:03A5742DFFDAAE05FF76AB38CEFFFAC4:8ED393E0FFDAAE05FCF8ADDBCE05FB74" box="[793,885,1166,1189]" country="Panama" latitude="8.733334" longLatPrecision="1300" longitude="-82.25" municipality="La" name="Fortuna" pageId="15" pageNumber="216" stateProvince="Chiriqui">Fortuna</location>
<geoCoordinate id="EE38A3FCFFDAAE05FC67ADDBCEA1FB77" box="[902,977,1162,1190]" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="925" value="8.733334">8o44'N</geoCoordinate>
<geoCoordinate id="EE38A3FCFFDAAE05FC03ADDBC931FB77" box="[994,1089,1162,1190]" direction="west" orientation="longitude" pageId="15" pageNumber="216" precision="925" value="-82.25">82o15'W</geoCoordinate>
,
<collectingDate id="EFF61A13FFDAAE05FBB7ADDBC98FFB77" box="[1110,1279,1166,1190]" pageId="15" pageNumber="216" value="1978-05-17">
<date id="FFB2E3FBFFDAAE05FBB7ADDBC98FFB77" box="[1110,1279,1166,1190]" pageId="15" pageNumber="216" value="1978-05-17">17 May 1978</date>
</collectingDate>
(
<collectorName id="26F9A0EDFFDAAE05FAF9ADD8C805FB77" box="[1304,1397,1165,1190]" pageId="15" pageNumber="216">O’Brien</collectorName>
&amp;
<collectorName id="26F9A0EDFFDAAE05FF76ADE4CC70FB1B" box="[151,256,1201,1226]" pageId="15" pageNumber="216">Marshall</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B64CF66FFDAAE05FEF5ADE7CC92FB20" ID-GBIF-Occurrence="1313797448" collectingDate="1992-06-22" collectionCode="AHCNC" collectorName="Jean &amp; Keve Ribardo" country="Panama" location="Fortuna watershed" pageId="15" pageNumber="216" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui" typeStatus="paratype">
<typeStatus id="54B77B99FFDAAE05FEF5ADE7CCF5FB1B" box="[276,389,1202,1226]" pageId="15" pageNumber="216" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0A0EB2FFDAAE05FE73ADE7CC88FB1B" box="[402,504,1201,1226]" pageId="15" pageNumber="216" type="female">1 female</specimenCount>
,
<quantity id="4CF468DEFFDAAE05FDE4ADE4CF3EFB1B" box="[517,590,1201,1226]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="216" unit="km" value="45.0">45 km</quantity>
N
<collectingRegion id="49C80BD9FFDAAE05FD94ADE4CFA2FB1B" box="[629,722,1201,1226]" country="Panama" name="Chiriqui" pageId="15" pageNumber="216">Chiriqui</collectingRegion>
[in]
<location id="8ED393E0FFDAAE05FCECADE7CE91FB1B" LSID="urn:lsid:plazi:treatment:03A5742DFFDAAE05FF76AB38CEFFFAC4:8ED393E0FFDAAE05FCECADE7CE91FB1B" box="[781,993,1201,1226]" country="Panama" name="Fortuna watershed" pageId="15" pageNumber="216" stateProvince="Chiriqui">Fortuna watershed</location>
,
<collectingDate id="EFF61A13FFDAAE05FC0EADE7C9F8FB1B" box="[1007,1160,1202,1226]" pageId="15" pageNumber="216" value="1992-06-22">
<date id="FFB2E3FBFFDAAE05FC0EADE7C9F8FB1B" box="[1007,1160,1202,1226]" pageId="15" pageNumber="216" value="1992-06-22">22 June 1992</date>
</collectingDate>
(
<collectorName id="26F9A0EDFFDAAE05FB78ADE7C9BDFB1B" box="[1177,1229,1202,1226]" pageId="15" pageNumber="216">Jean</collectorName>
&amp;
<collectorName id="26F9A0EDFFDAAE05FB15ADE7C8E5FB1B" box="[1268,1429,1201,1226]" pageId="15" pageNumber="216">Keve Ribardo</collectorName>
) Barcode of Life
<collectionCode id="ED1D5DFEFFDAAE05FEAEAD8DCCC8FB20" box="[335,440,1240,1265]" pageId="15" pageNumber="216">AHCNC</collectionCode>
162
</materialsCitation>
-
<materialsCitation id="3B64CF66FFDAAE05FE08AD8DCC74FAC4" ID-GBIF-Occurrence="1313797447" collectingDate="2000-05-30" collectionCode="AHCNC" collectorName="B. &amp; J. Gill" country="Panama" latitude="8.722333" location="Fortuna watershed" longLatPrecision="13" longitude="-82.2375" pageId="15" pageNumber="216" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
12;
<specimenCount id="9D0A0EB2FFDAAE05FDEEAD8CCF2DFB20" box="[527,605,1240,1265]" pageId="15" pageNumber="216" type="male">1 male</specimenCount>
,
<specimenCount id="9D0A0EB2FFDAAE05FD89AD8CCFBCFB20" box="[616,716,1240,1265]" pageId="15" pageNumber="216" type="female">1 female</specimenCount>
, same data as holotype,
<geoCoordinate id="EE38A3FCFFDAAE05FC3AAD8CC939FB20" box="[987,1097,1236,1265]" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="9" value="8.722333">8o43.34'N</geoCoordinate>
<geoCoordinate id="EE38A3FCFFDAAE05FBAEAD8CC9A1FB20" box="[1103,1233,1236,1265]" direction="west" orientation="longitude" pageId="15" pageNumber="216" precision="9" value="-82.2375">82o14.25'W</geoCoordinate>
,
<collectingDate id="EFF61A13FFDAAE05FB3CAD8CC802FB20" box="[1245,1394,1241,1265]" pageId="15" pageNumber="216" value="2000-05-30">
<date id="FFB2E3FBFFDAAE05FB3CAD8CC802FB20" box="[1245,1394,1241,1265]" pageId="15" pageNumber="216" value="2000-05-30">30 May 2000</date>
</collectingDate>
(
<collectorName id="26F9A0EDFFDAAE05FA62AD8CCD8CFAC4" pageId="15" pageNumber="216">B. &amp; J. Gill</collectorName>
)
</materialsCitation>
. Holotype and 1 paratype in CAAS; 2 paratypes in CNCI.
</paragraph>
</subSubSection>
</treatment>
</document>
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