159 lines
19 KiB
XML
159 lines
19 KiB
XML
<document id="DBA137D938E681DC8212E8962440B5C0" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří & Bordoni, Arnaldo" docDate="2015" docId="039D87F5556EFF94FF4AFEEAFC37BA2A" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thyreocephalus camerunensis Janák & Bordoni, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="86" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="86" updateTime="1698618126966" updateUser="plazi">
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<mods:mods id="D9E342CF2CF8B9598B14E63F76F442E5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B679DF2E04B2BD9F28BCF948FC84CE85">
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<mods:title id="C63E1379D500A0E65507B690176F7ED1">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:name id="805C9C3FCF60A30C16F3EC6E654238E0" type="personal">
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<mods:roleTerm id="46C5C1612886C2A43FADD3DF1DF59A69">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D66C46949882BF1E2C7405B7905638C3">Janák, Jiří</mods:namePart>
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</mods:name>
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<mods:name id="5A7FC706751645A4B7245282E0DEF726" type="personal">
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<mods:role id="FD59B2C84276BEE555045A7821B3A154">
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<mods:roleTerm id="7609D14CE2856124D2A60FDD9295CD19">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DC5067549F4D6480F564A043A866457B">Bordoni, Arnaldo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="957DFBF6AA81103A1D197449B76E6309">text</mods:typeOfResource>
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<mods:titleInfo id="C4DFCDC2F43BDE23B5311ED24BB4A396">
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<mods:title id="7C9C2C06352069E519985BCC1C607918">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="69E57426E04AC96EEA60C5B25BB21D88">
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<mods:date id="0FD723DDA3F2ADF6AF241C4004027D65">2015</mods:date>
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<mods:detail id="CE33A8748C1F892B2D74726FF895026D" type="volume">
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<mods:number id="14A205F23B7CCD03966E27AEB6B06A63">4038</mods:number>
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</mods:detail>
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<mods:detail id="EF0B5F5B1A07BD1EA9EC4AAA67233C74" type="issue">
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<mods:number id="5DC1EE7C253C0BE882853821BDBF489E">1</mods:number>
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<mods:end id="09B03E6A75F1A297BF5EF59EA34F21C9">94</mods:end>
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<mods:classification id="763DE0C2ECEC59FFA478473CC98F3A8A">journal article</mods:classification>
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<mods:identifier id="BD9209F3ED501F375699F3F1CE60FB0F" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
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<mods:identifier id="3DDC6F67808D58208EB0894E21FE49D8" type="GBIF-Dataset">066e84f1-206b-40cb-a615-2126696b813d</mods:identifier>
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<mods:identifier id="F7B1169C199D83C8C3506EFCC83563FC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7A942CBB9DB4F2BFE7F63E296D7231B8" type="Zenodo-Dep">289876</mods:identifier>
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</mods:mods>
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<treatment id="039D87F5556EFF94FF4AFEEAFC37BA2A" ID-DOI="http://doi.org/10.5281/zenodo.6101713" ID-GBIF-Taxon="127693934" ID-Zenodo-Dep="6101713" LSID="urn:lsid:plazi:treatment:039D87F5556EFF94FF4AFEEAFC37BA2A" httpUri="http://treatment.plazi.org/id/039D87F5556EFF94FF4AFEEAFC37BA2A" lastPageNumber="86" pageId="85" pageNumber="86">
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<subSubSection id="C32E6568556EFF94FF4AFEEAFE18BFA2" pageId="85" pageNumber="86" type="nomenclature">
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<paragraph id="8B8B36E3556EFF94FF4AFEEAFD93BF43" blockId="85.[151,620,332,391]" box="[151,620,332,358]" pageId="85" pageNumber="86">
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<heading id="D0C3818F556EFF94FF4AFEEAFD93BF43" bold="true" box="[151,620,332,358]" fontSize="11" level="1" pageId="85" pageNumber="86" reason="1">
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<emphasis id="B940EAF1556EFF94FF4AFEEAFD93BF43" bold="true" box="[151,620,332,358]" pageId="85" pageNumber="86">
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<taxonomicName id="4C344D60556EFF94FF4AFEEAFDF7BF43" ID-CoL="7C9FS" box="[151,520,332,358]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="camerunensis" status="sp. nov.">
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<emphasis id="B940EAF1556EFF94FF4AFEEAFDF7BF43" bold="true" box="[151,520,332,358]" italics="true" pageId="85" pageNumber="86">Thyreocephalus camerunensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273578A556EFF94FDD2FEEAFD93BF43" box="[527,620,332,358]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8B36E3556EFF94FF4AFEC8FE18BFA2" blockId="85.[151,620,332,391]" box="[151,487,366,391]" pageId="85" pageNumber="86">
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(
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<figureCitation id="130F2A66556EFF94FF42FEC8FEC7BFA2" box="[159,312,366,391]" captionStart="FIGURES 183 – 188. 183, 184" captionStartId="71.[151,264,1662,1684]" captionTargetBox="[174,1420,209,1626]" captionTargetId="figure@71.[151,1436,193,1640]" captionTargetPageId="71" captionText="FIGURES 183 – 188. 183, 184, Thyreocephalus camerunensis sp. nov.; 185, 186, T. marginipennis sp. nov.; 187, 188, T. semipiceus Bernhauer; 183, 185, 187, habitus; 184, 186, 188, head. Scale 1 mm." httpUri="https://zenodo.org/record/289918/files/figure.png" pageId="85" pageNumber="86">Fig. 183, 184</figureCitation>
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,
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<figureCitation id="130F2A66556EFF94FE99FEC8FE8EBFA2" box="[324,369,366,391]" captionStart="FIGURE 206" captionStartId="79.[151,250,1606,1628]" captionTargetBox="[158,1431,199,1578]" captionTargetId="figure@79.[151,1436,193,1584]" captionTargetPageId="79" captionText="FIGURE 206. Distribution of species in Thyreocephalus mocquerysi group (★ = T. atlanticus (Bernhauer), ▲ = T. burgeoni (Bernhauer), ■ = T. camerunensis sp. nov., Δ = T. marginipennis sp. nov., ● = T. mokaensis (Bernhauer), ○ = T. strinatii (Bernhauer)." httpUri="https://zenodo.org/record/289924/files/figure.png" pageId="85" pageNumber="86">206</figureCitation>
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,
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<figureCitation id="130F2A66556EFF94FEA1FEC9FE1FBFA2" box="[380,480,367,391]" captionStart="FIGURES 223 – 227" captionStartId="86.[151,264,1970,1992]" captionTargetBox="[213,1377,207,1932]" captionTargetId="figure@86.[195,1391,193,1949]" captionTargetPageId="86" captionText="FIGURES 223 – 227. Thyreocephalus camerunensis sp. nov.; 223, head and pronotum; 224, labrum; 225, male tergite 10; 226, male sternite 9; 227, aedeagus." httpUri="https://zenodo.org/record/289928/files/figure.png" pageId="85" pageNumber="86">223–227</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6568556EFF94FF4AFE10FC70BC1E" pageId="85" pageNumber="86" type="materials_examined">
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<paragraph id="8B8B36E3556EFF94FF4AFE10FDD5BFEA" blockId="85.[151,554,438,463]" box="[151,554,438,463]" pageId="85" pageNumber="86">
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<emphasis id="B940EAF1556EFF94FF4AFE10FECABFEA" bold="true" box="[151,309,438,463]" pageId="85" pageNumber="86">
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<typeStatus id="548F8841556EFF94FF4AFE10FF2DBFEA" box="[151,210,438,463]" pageId="85" pageNumber="86">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3237673556EFF94FEE1FE11FE4BBFEA" box="[316,436,439,463]" name="Cameroon" pageId="85" pageNumber="86">Cameroon</collectingCountry>
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, Lolodorf.
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</paragraph>
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<paragraph id="8B8B36E3556EFF94FF4AFE58FC70BC1E" blockId="85.[151,1437,510,1039]" pageId="85" pageNumber="86">
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<emphasis id="B940EAF1556EFF94FF4AFE58FE3BBC32" bold="true" box="[151,452,510,535]" pageId="85" pageNumber="86">
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<typeStatus id="548F8841556EFF94FF4AFE58FF2DBC32" box="[151,210,510,535]" pageId="85" pageNumber="86">Type</typeStatus>
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material.
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<typeStatus id="548F8841556EFF94FE8AFE58FE3BBC32" box="[343,452,510,535]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
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</emphasis>
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♂:
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<collectingCountry id="F3237673556EFF94FE26FE58FD5BBC32" box="[507,676,510,535]" name="Cameroon" pageId="85" pageNumber="86">
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<emphasis id="B940EAF1556EFF94FE26FE58FD5BBC32" bold="true" box="[507,676,510,535]" pageId="85" pageNumber="86">CAMEROON</emphasis>
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</collectingCountry>
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: “SO
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<collectingCountry id="F3237673556EFF94FD2BFE59FC9FBC32" box="[758,864,511,535]" name="Cameroon" pageId="85" pageNumber="86">Camerun</collectingCountry>
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, Lolodorf, L. Conradt 1895” (
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<collectionCode id="ED25AE26556EFF94FB13FE59FAFDBC32" LSID="urn:lsid:biocol.org:col:33211" box="[1230,1282,511,535]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="85" pageNumber="86">DEI</collectionCode>
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), “
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<typeStatus id="548F8841556EFF94FAF8FE58FA63BC32" box="[1317,1436,510,535]" pageId="85" pageNumber="86" type="holotype">Holotypus</typeStatus>
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<taxonomicName id="4C344D60556EFF94FF4AFD84FE15BC1E" box="[151,490,546,571]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="camerunensis" status="sp. nov.">
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<emphasis id="B940EAF1556EFF94FF4AFD84FE15BC1E" box="[151,490,546,571]" italics="true" pageId="85" pageNumber="86">Thyreocephalus camerunensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273578A556EFF94FE2CFD85FDB6BC1F" box="[497,585,547,570]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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, Janák & Bordoni det. 2015”
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6568556EFF94FF1AFDE0FAE8BD5A" pageId="85" pageNumber="86" type="description">
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<paragraph id="8B8B36E3556EFF94FF1AFDE0FCF3BD12" blockId="85.[151,1437,510,1039]" pageId="85" pageNumber="86">
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<emphasis id="B940EAF1556EFF94FF1AFDE0FEA6BC7A" bold="true" box="[199,345,582,607]" pageId="85" pageNumber="86">Description.</emphasis>
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Body length
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<quantity id="4CCC9B06556EFF94FE2FFDE0FDA5BC7B" box="[498,602,582,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="85" pageNumber="86" unit="mm" value="12.5">12.5 mm</quantity>
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; length from anterior margin of head to posterior margin of elytra:
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<quantity id="4CCC9B06556EFF94FA8EFDE0FA67BC7B" box="[1363,1432,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="85" pageNumber="86" unit="mm" value="6.0">6 mm</quantity>
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. Head and pronotum shiny, with micro-punctation. Head black; pronotum, elytra and abdomen reddish brown; antennae brown, legs reddish brown (
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<figureCitation id="130F2A66556EFF94FD9BFD28FD55BC82" box="[582,682,654,679]" captionStart="FIGURES 183 – 188. 183, 184" captionStartId="71.[151,264,1662,1684]" captionTargetBox="[174,1420,209,1626]" captionTargetId="figure@71.[151,1436,193,1640]" captionTargetPageId="71" captionText="FIGURES 183 – 188. 183, 184, Thyreocephalus camerunensis sp. nov.; 185, 186, T. marginipennis sp. nov.; 187, 188, T. semipiceus Bernhauer; 183, 185, 187, habitus; 184, 186, 188, head. Scale 1 mm." httpUri="https://zenodo.org/record/289918/files/figure.png" pageId="85" pageNumber="86">Fig. 183</figureCitation>
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). Head and pronotum and related punctation as in
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<figureCitation id="130F2A66556EFF94FB2DFD28FAA3BC82" box="[1264,1372,654,679]" captionStart="FIGURES 183 – 188. 183, 184" captionStartId="71.[151,264,1662,1684]" captionTargetBox="[174,1420,209,1626]" captionTargetId="figure@71.[151,1436,193,1640]" captionTargetPageId="71" captionText="FIGURES 183 – 188. 183, 184, Thyreocephalus camerunensis sp. nov.; 185, 186, T. marginipennis sp. nov.; 187, 188, T. semipiceus Bernhauer; 183, 185, 187, habitus; 184, 186, 188, head. Scale 1 mm." httpUri="https://zenodo.org/record/289918/files/figure.png" pageId="85" pageNumber="86">Figs. 184</figureCitation>
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,
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<figureCitation id="130F2A66556EFF94FAB6FD29FA67BC82" box="[1387,1432,655,679]" captionStart="FIGURES 223 – 227" captionStartId="86.[151,264,1970,1992]" captionTargetBox="[213,1377,207,1932]" captionTargetId="figure@86.[195,1391,193,1949]" captionTargetPageId="86" captionText="FIGURES 223 – 227. Thyreocephalus camerunensis sp. nov.; 223, head and pronotum; 224, labrum; 225, male tergite 10; 226, male sternite 9; 227, aedeagus." httpUri="https://zenodo.org/record/289928/files/figure.png" pageId="85" pageNumber="86">223</figureCitation>
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; labrum as in
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<figureCitation id="130F2A66556EFF94FEF5FD14FE76BCEE" box="[296,393,690,715]" captionStart="FIGURES 223 – 227" captionStartId="86.[151,264,1970,1992]" captionTargetBox="[213,1377,207,1932]" captionTargetId="figure@86.[195,1391,193,1949]" captionTargetPageId="86" captionText="FIGURES 223 – 227. Thyreocephalus camerunensis sp. nov.; 223, head and pronotum; 224, labrum; 225, male tergite 10; 226, male sternite 9; 227, aedeagus." httpUri="https://zenodo.org/record/289928/files/figure.png" pageId="85" pageNumber="86">Fig. 224</figureCitation>
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. Elytra sub-rectangular, with sub-rectilinear and sub-parallel sides, shorter and slightly wider than pronotum, with rounded humeral angles. Surface with distinct, fine, spaced punctation, arranged in three series, one near suture, one median and one lateral. Abdomen with fine and very dense, transverse micro-striation and fine and deep punctation, arranged in several series.
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</paragraph>
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<paragraph id="8B8B36E3556EFF94FF1AFCE4FAE8BD5A" blockId="85.[151,1437,510,1039]" pageId="85" pageNumber="86">
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<collectingCountry id="F3237673556EFF94FF1AFCE4FEFEBD7E" box="[199,257,834,859]" name="Malta" pageId="85" pageNumber="86">Male</collectingCountry>
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.Tergite 10 and sternite 9 of male genital segment as in
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<figureCitation id="130F2A66556EFF94FCB1FCE4FBF0BD7E" box="[876,1039,834,859]" captionStart="FIGURES 223 – 227" captionStartId="86.[151,264,1970,1992]" captionTargetBox="[213,1377,207,1932]" captionTargetId="figure@86.[195,1391,193,1949]" captionTargetPageId="86" captionText="FIGURES 223 – 227. Thyreocephalus camerunensis sp. nov.; 223, head and pronotum; 224, labrum; 225, male tergite 10; 226, male sternite 9; 227, aedeagus." httpUri="https://zenodo.org/record/289928/files/figure.png" pageId="85" pageNumber="86">Figs. 225, 226</figureCitation>
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. Aedeagus (
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<figureCitation id="130F2A66556EFF94FB45FCE4FB07BD7E" box="[1176,1272,834,859]" captionStart="FIGURES 223 – 227" captionStartId="86.[151,264,1970,1992]" captionTargetBox="[213,1377,207,1932]" captionTargetId="figure@86.[195,1391,193,1949]" captionTargetPageId="86" captionText="FIGURES 223 – 227. Thyreocephalus camerunensis sp. nov.; 223, head and pronotum; 224, labrum; 225, male tergite 10; 226, male sternite 9; 227, aedeagus." httpUri="https://zenodo.org/record/289928/files/figure.png" pageId="85" pageNumber="86">Fig. 227</figureCitation>
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)
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||
<quantity id="4CCC9B06556EFF94FADAFCE5FAA2BD7F" box="[1287,1373,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="85" pageNumber="86" unit="mm" value="1.7">1.7 mm</quantity>
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long, ovoid, with narrow, sub-triangular median lobe; parameres long; inner sac very narrow, sub-rectilinear.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6568556EFF94FF1AFC2CFAF3BDCF" pageId="85" pageNumber="86" type="etymology">
|
||
<paragraph id="8B8B36E3556EFF94FF1AFC2CFCE7BD86" blockId="85.[151,1437,510,1039]" box="[199,792,906,931]" pageId="85" pageNumber="86">
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<emphasis id="B940EAF1556EFF94FF1AFC2CFEB1BD86" bold="true" box="[199,334,906,931]" pageId="85" pageNumber="86">Etymology.</emphasis>
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The specific epithet refers to
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<collectingCountry id="F3237673556EFF94FD41FC2DFCEABD86" box="[668,789,907,931]" name="Cameroon" pageId="85" pageNumber="86">Cameroon</collectingCountry>
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.
|
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</paragraph>
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<paragraph id="8B8B36E3556EFF94FF1AFC08FAF3BDCF" blockId="85.[151,1437,510,1039]" pageId="85" pageNumber="86">
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<emphasis id="B940EAF1556EFF94FF1AFC08FE2ABDE2" bold="true" box="[199,469,942,967]" pageId="85" pageNumber="86">Differential diagnosis.</emphasis>
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<taxonomicName id="4C344D60556EFF94FE39FC08FCC1BDE3" box="[484,830,942,967]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="camerunensis" status="sp. nov.">
|
||
<emphasis id="B940EAF1556EFF94FE39FC08FCC1BDE3" box="[484,830,942,967]" italics="true" pageId="85" pageNumber="86">Thyreocephalus camerunensis</emphasis>
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</taxonomicName>
|
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<taxonomicNameLabel id="A273578A556EFF94FC91FC08FC51BDE2" box="[844,942,942,967]" pageId="85" pageNumber="86" rank="species">
|
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<emphasis id="B940EAF1556EFF94FC91FC08FC51BDE2" bold="true" box="[844,942,942,967]" pageId="85" pageNumber="86">sp. nov.</emphasis>
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</taxonomicNameLabel>
|
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differs from the similar
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<taxonomicName id="4C344D60556EFF94FB3BFC09FA88BDE2" box="[1254,1399,942,967]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="atlanticus">
|
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<emphasis id="B940EAF1556EFF94FB3BFC09FA88BDE2" box="[1254,1399,942,967]" italics="true" pageId="85" pageNumber="86">T. atlanticus</emphasis>
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</taxonomicName>
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in distinctly punctate disc of head and in the aedeagus with narrower median lobe and longer parameres.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6568556EFF94FF1AFC50FC37BA2A" box="[199,968,1014,1039]" pageId="85" pageNumber="86" type="distribution">
|
||
<paragraph id="8B8B36E3556EFF94FF1AFC50FC37BA2A" blockId="85.[151,1437,510,1039]" box="[199,968,1014,1039]" pageId="85" pageNumber="86">
|
||
<emphasis id="B940EAF1556EFF94FF1AFC50FE9FBA2A" bold="true" box="[199,352,1014,1039]" pageId="85" pageNumber="86">Distribution.</emphasis>
|
||
The species is known from
|
||
<collectingCountry id="F3237673556EFF94FD41FC51FCEEBA2A" box="[668,785,1015,1039]" name="Cameroon" pageId="85" pageNumber="86">Cameroon</collectingCountry>
|
||
only (
|
||
<figureCitation id="130F2A66556EFF94FC85FC50FC44BA2A" box="[856,955,1014,1039]" captionStart="FIGURE 206" captionStartId="79.[151,250,1606,1628]" captionTargetBox="[158,1431,199,1578]" captionTargetId="figure@79.[151,1436,193,1584]" captionTargetPageId="79" captionText="FIGURE 206. Distribution of species in Thyreocephalus mocquerysi group (★ = T. atlanticus (Bernhauer), ▲ = T. burgeoni (Bernhauer), ■ = T. camerunensis sp. nov., Δ = T. marginipennis sp. nov., ● = T. mokaensis (Bernhauer), ○ = T. strinatii (Bernhauer)." httpUri="https://zenodo.org/record/289924/files/figure.png" pageId="85" pageNumber="86">Fig. 206</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |