154 lines
19 KiB
XML
154 lines
19 KiB
XML
<document id="691D0464A6656CD91A1604EC4D6335D4" ID-DOI="10.11646/zootaxa.4006.3.1" ID-GBIF-Dataset="9b8d3194-5435-46fb-92ee-301fd23b0782" ID-ISSN="1175-5326" ID-Zenodo-Dep="240836" ID-ZooBank="136F727A-1628-449F-B762-254371D27360" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461211198470" checkinUser="plazi" docAuthor="Köhler, Jörn, Glaw, Frank, Pabijan, Maciej & Vences, Miguel" docDate="2015" docId="039D87F4FFB4C80E08C1FDF9FA76E417" docLanguage="en" docName="zt04006p438.pdf" docOrigin="Zootaxa 4006 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aglyptodactylus laticeps Glaw, Vences & Bohme 1998" docType="treatment" docVersion="9" lastPageNumber="417" masterDocId="FFA4FF8CFFBBC81E0856FFDBFFFCE677" masterDocTitle="Integrative taxonomic revision of mantellid frogs of the genus Aglyptodactylus (Anura: Mantellidae)" masterLastPageNumber="438" masterPageNumber="401" pageNumber="416" updateTime="1698602080951" updateUser="plazi">
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<mods:title id="75811E47EDC61600DD0AA5404AD39C6B">Integrative taxonomic revision of mantellid frogs of the genus Aglyptodactylus (Anura: Mantellidae)</mods:title>
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<mods:namePart id="96527875EB31AEDA8C404D398091A526">Köhler, Jörn</mods:namePart>
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<mods:namePart id="F82121209D69B8F6CA53272FE88949E3">Glaw, Frank</mods:namePart>
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<mods:namePart id="07B4AD2BFC8D95BA464ACFF82CFDEA0E">Pabijan, Maciej</mods:namePart>
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<mods:namePart id="06229675EBB30A985721F7B7E7FD55B8">Vences, Miguel</mods:namePart>
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<mods:title id="FF87FE4208A8CF427310A85F74BDCCEB">Zootaxa</mods:title>
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<mods:date id="036DDF7B2045A9107110966E94DF9B13">2015</mods:date>
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<treatment id="039D87F4FFB4C80E08C1FDF9FA76E417" ID-DOI="http://doi.org/10.5281/zenodo.5684804" ID-GBIF-Taxon="119630505" ID-Zenodo-Dep="5684804" LSID="urn:lsid:plazi:treatment:039D87F4FFB4C80E08C1FDF9FA76E417" httpUri="http://treatment.plazi.org/id/039D87F4FFB4C80E08C1FDF9FA76E417" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
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<subSubSection id="C32E6569FFB4C80E08C1FDF9FE7AE782" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416" type="nomenclature">
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<paragraph id="8B8B36E2FFB4C81108C1FDF9FCD2E44B" blockId="15.[151,814,546,572]" box="[151,814,546,572]" pageId="15" pageNumber="416">
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<heading id="D0C3818EFFB4C81108C1FDF9FCD2E44B" bold="true" box="[151,814,546,572]" fontSize="11" level="1" pageId="15" pageNumber="416" reason="1">
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<taxonomicName id="4C344D61FFB4C81108C1FDF9FCD2E44B" ID-CoL="5TRB8" authority="Glaw, Vences & Bohme, 1998" authorityName="Glaw, Vences & Bohme" authorityYear="1998" box="[151,814,546,572]" class="Amphibia" family="Mantellidae" genus="Aglyptodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="416" phylum="Chordata" rank="species" species="laticeps">
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<emphasis id="B940EAF0FFB4C81108C1FDF9FCD2E44B" bold="true" box="[151,814,546,572]" pageId="15" pageNumber="416">
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<emphasis id="B940EAF0FFB4C81108C1FDF9FE45E44B" bold="true" box="[151,441,546,572]" italics="true" pageId="15" pageNumber="416">Aglyptodactylus laticeps</emphasis>
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<bibRefCitation id="EFA54B13FFB4C8110997FDF9FCD2E44B" author="Glaw" box="[449,814,546,572]" pageId="15" pageNumber="416" refString="Glaw, F., Vences, M. & Bohme, W. (1998) Systematic revision of the genus Aglyptodactylus Boulenger, 1919 (Amphibia: Ranidae), and analysis of its phylogenetic relationships to other Madagascan ranid genera (Tomopterna, Boophis, Mantidactylus, and Mantella). Journal of Zoological Systematics and Evolutionary Research, 36 (1), 17 - 37. http: // dx. doi. org / 10.1111 / j. 1439 - 0469.1998. tb 00775. x" type="journal article" year="1998">Glaw, Vences & Böhme, 1998</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B8B36E2FFB4C81108C1FDB2FF0EE565" blockId="15.[151,1437,617,786]" pageId="15" pageNumber="416">
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<emphasis id="B940EAF0FFB4C81108C1FDB2FF07E4F5" bold="true" box="[151,251,617,642]" pageId="15" pageNumber="416">Identity.</emphasis>
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This species was described and diagnosed from Kirindy Forest, western
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<collectingCountry id="F3237672FFB4C8110CFFFDB2FACFE4F5" box="[1193,1331,617,642]" name="Madagascar" pageId="15" pageNumber="416">Madagascar</collectingCountry>
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. Among
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<taxonomicName id="4C344D61FFB4C81108C1FD56FEB7E4D2" box="[151,331,653,677]" class="Amphibia" family="Mantellidae" genus="Aglyptodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="416" phylum="Chordata" rank="genus">
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<emphasis id="B940EAF0FFB4C81108C1FD56FEB7E4D2" box="[151,331,653,677]" italics="true" pageId="15" pageNumber="416">Aglyptodactylus</emphasis>
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</taxonomicName>
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, this species can be unequivocally identified by its different external morphology alone (
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<figureCitation id="130F2A67FFB4C8110D39FD56FF59E4BD" captionStart="FIGURE 6" captionStartId="10.[151,250,1734,1756]" captionTargetBox="[151,1436,525,1712]" captionTargetId="figure@10.[151,1436,517,1713]" captionTargetPageId="10" captionText="FIGURE 6. Photographs in life of Aglyptodactylus laticeps and A. securifer: A. laticeps from Kirindy (a – c), with (b) being a female, and (c) probably being a male. Frontodorsal and ventral views of male holotype of A. securifer (ZFMK 59975) from Kirindy (d, g); dorsal and ventral views of a male of A. securifer from Ankarana (e, h); dorsal and ventral views of a female of A. securifer from Ankarana (ZSM 762 / 2003) (f, i); A. securifer from Berara (j); A. securifer from Tsingy de Bemaraha (k – l), with (k) corresponding to ZSM 14 / 2006. Photographed individuals not unequivocally assignable to preserved voucher specimens unless otherwise mentioned." httpUri="https://zenodo.org/record/240842/files/figure.png" pageId="15" pageNumber="416">Fig. 6</figureCitation>
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a‒c). It is morphologically distinguished from all other known species in the genus by its broad head, large inner metatarsal tubercle, comparatively poorly developed webbing on feet, and largely connected lateral metatarsalia (
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<figureCitation id="130F2A67FFB4C81108C9FD22FF19E565" box="[159,229,761,786]" captionStart="FIGURE 4" captionStartId="8.[151,250,1780,1802]" captionTargetBox="[151,1436,1129,1758]" captionTargetId="figure@8.[151,1436,1129,1758]" captionTargetPageId="8" captionText="FIGURE 4. Ventral view of feet of representative specimens of Aglyptodactylus (not to scale; see Table 3 for measurements). Arrows point at the large inner metatarsal tubercles of A. laticeps and A. securifer, and at the lateral metatarsalia of A. laticeps connected by tissue rather than separated by webbing. Asterisks mark mirrored images." httpUri="https://zenodo.org/record/240840/files/figure.png" pageId="15" pageNumber="416">Fig. 4</figureCitation>
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).
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</paragraph>
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<caption id="DF4B666AFFB4C81108C1F932FEF8E169" httpUri="https://zenodo.org/record/240843/files/figure.png" pageId="15" pageNumber="416" targetBox="[164,1401,838,1744]" targetPageId="15">
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<paragraph id="8B8B36E2FFB4C81108C1F932FEF8E169" blockId="15.[151,1436,1769,1822]" pageId="15" pageNumber="416">
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<emphasis id="B940EAF0FFB4C81108C1F932FEF0E088" bold="true" box="[151,268,1769,1791]" pageId="15" pageNumber="416">FIGURE 7</emphasis>
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. Audiospectrogram (above) and oscillogram (below) of an advertisement call of
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<taxonomicName id="4C344D61FFB4C8110C69F932FACBE088" box="[1087,1335,1769,1791]" class="Amphibia" family="Mantellidae" genus="Aglyptodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="416" phylum="Chordata" rank="species" species="laticeps">
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<emphasis id="B940EAF0FFB4C8110C69F932FACBE088" box="[1087,1335,1769,1791]" italics="true" pageId="15" pageNumber="416">Aglyptodactylus laticeps</emphasis>
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</taxonomicName>
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, recorded at Kirindy.
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</paragraph>
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</caption>
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<paragraph id="8B8B36E2FFB4C80E0891F89EFA6AE7FF" blockId="15.[151,1436,1861,2030]" lastBlockId="16.[151,1436,151,608]" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
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<emphasis id="B940EAF0FFB4C8110891F89EFEFCE129" bold="true" box="[199,256,1861,1886]" pageId="15" pageNumber="416">Call.</emphasis>
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The advertisement call of the species has been described by Glaw
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<emphasis id="B940EAF0FFB4C8110BB9F89DFBD4E12A" box="[1007,1064,1861,1885]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
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(1998) from recordings obtained in 1995 from Kirindy. Our analysis of calls recorded by J. Glos at the same locality in 2001 (Vences
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<emphasis id="B940EAF0FFB4C8110D4CF8B1FAA8E1F6" box="[1306,1364,1897,1921]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
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2006: CD1/Track1) generally confirmed the call characters previously reported. The call consists of a series of pulsed notes repeated at somewhat irregular intervals (
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<figureCitation id="130F2A67FFB4C8110AFAF86AFD0DE1BD" box="[684,753,1969,1994]" captionStart="FIGURE 7" captionStartId="15.[151,250,1769,1791]" captionTargetBox="[164,1401,838,1744]" captionTargetId="figure@15.[164,1402,838,1745]" captionTargetPageId="15" captionText="FIGURE 7. Audiospectrogram (above) and oscillogram (below) of an advertisement call of Aglyptodactylus laticeps, recorded at Kirindy." httpUri="https://zenodo.org/record/240843/files/figure.png" pageId="15" pageNumber="416">Fig. 7</figureCitation>
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). Notes are composed of 7‒12 well-separated pulses emitted with increasing repetition rate from beginning towards the end of the note. Slight differences among recordings are apparent in distinctly longer inter-note intervals (170‒319 vs. 31‒133 ms) in the more recent recordings, and might be the result of differences in calling motivation and/or temperature (temperature not available for 2001 recordings). Furthermore, our analysis revealed an amplitude modulation within notes (
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<figureCitation id="130F2A67FFABC80E0CD6FF04FB2EE68F" box="[1152,1234,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1772,1794]" captionTargetBox="[151,1436,193,1750]" captionTargetId="figure@6.[151,1436,193,1750]" captionTargetPageId="6" captionText="FIGURE 3. Comparative oscillograms of the calls of known species of Aglyptodactylus at the same time scale (2000 milliseconds section each)." httpUri="https://zenodo.org/record/240839/files/figure.png" pageId="16" pageNumber="417">Figs. 3</figureCitation>
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,
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<figureCitation id="130F2A67FFABC80E0CB6FF3BFB13E68F" box="[1248,1263,224,248]" captionStart="FIGURE 7" captionStartId="15.[151,250,1769,1791]" captionTargetBox="[164,1401,838,1744]" captionTargetId="figure@15.[164,1402,838,1745]" captionTargetPageId="15" captionText="FIGURE 7. Audiospectrogram (above) and oscillogram (below) of an advertisement call of Aglyptodactylus laticeps, recorded at Kirindy." httpUri="https://zenodo.org/record/240843/files/figure.png" pageId="16" pageNumber="417">7</figureCitation>
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) that is unique among known calls in
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<taxonomicName id="4C344D61FFABC80E09CDFEDFFDB3E76B" box="[411,591,260,284]" class="Amphibia" family="Mantellidae" genus="Aglyptodactylus" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="genus">
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<emphasis id="B940EAF0FFABC80E09CDFEDFFDB3E76B" box="[411,591,260,284]" italics="true" pageId="16" pageNumber="417">Aglyptodactylus</emphasis>
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</taxonomicName>
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, with amplitude increasing towards the middle of the note and decreasing again towards its end. This amplitude modulation is not evident from the former call description (Glaw
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<emphasis id="B940EAF0FFABC80E0D40FEF2FAB1E737" box="[1302,1357,296,320]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
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1998), likely due to the processing/filtering of the poor-quality recording, as in the oscillogram figured in Glaw
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<emphasis id="B940EAF0FFABC80E0D37FE96FA60E713" box="[1377,1436,332,356]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
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(1998) the amplitude appears to be cut off at a certain level. For call parameters of the 2001 recording see Table 2.
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</paragraph>
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<paragraph id="8B8B36E2FFABC80E0891FE4FFE7AE782" blockId="16.[151,1436,151,608]" pageId="16" pageNumber="417">
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<emphasis id="B940EAF0FFABC80E0891FE4FFE7AE7DA" bold="true" box="[199,390,404,429]" pageId="16" pageNumber="417">Natural history.</emphasis>
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Glaw
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<emphasis id="B940EAF0FFABC80E0982FE4EFDF2E7DB" box="[468,526,404,428]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
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(1998),
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<bibRefCitation id="EFA54B13FFABC80E0A26FE4FFD05E7DB" author="Glos" box="[624,761,404,429]" pageId="16" pageNumber="417" refString="Glos, J. (2003) The amphibian fauna of the Kirindy dry forest in western Madagascar. Salamandra, 39, 75 - 90." type="journal article" year="2003">Glos (2003)</bibRefCitation>
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and
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<bibRefCitation id="EFA54B13FFABC80E0B64FE4FFB9DE7DB" author="Glos" box="[818,1121,404,429]" pageId="16" pageNumber="417" refString="Glos, J. & Linsenmair, K. E. (2004) Descriptions of the tadpoles of Aglyptodactylus laticeps and Aglyptodactylus securifer from western Madagascar, with notes on life history and ecology. Journal of Herpetology, 38, 131 - 136." type="journal article" year="2004">Glos & Linsenmair (2004)</bibRefCitation>
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provided life history notes, and the latter authors also described the tadpole.
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<bibRefCitation id="EFA54B13FFABC80E0AE3FE6CFC1CE7A7" box="[693,992,439,464]" pageId="16" pageNumber="417" refString="Randrianiaina, R. - D., Kohler, J., Glos, J., Vences, M. & Glaw, F. (2011) Where to grow in the Tsingy? Limestone rock pools as breeding habitats of the relict frog Tsingymantis antitra from Madagascar and description of its tadpole. Salamandra, 47, 77 - 89." type="journal article">
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Randrianiaina
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<emphasis id="B940EAF0FFABC80E0B0FFE62FC6CE7A7" box="[857,912,440,464]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
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(2011)
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</bibRefCitation>
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provided figures of
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<taxonomicName id="4C344D61FFABC80E0C95FE63FAC4E7A7" box="[1219,1336,440,464]" class="Amphibia" family="Mantellidae" genus="Aglyptodactylus" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="laticeps">
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<emphasis id="B940EAF0FFABC80E0C95FE63FAC4E7A7" box="[1219,1336,440,464]" italics="true" pageId="16" pageNumber="417">A. laticeps</emphasis>
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</taxonomicName>
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tadpoles and their mouthparts.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6569FFABC80E0891FE24FA76E417" pageId="16" pageNumber="417" type="distribution">
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<paragraph id="8B8B36E2FFABC80E0891FE24FA76E417" blockId="16.[151,1436,151,608]" pageId="16" pageNumber="417">
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<emphasis id="B940EAF0FFABC80E0891FE24FE9CE46F" bold="true" box="[199,352,511,536]" pageId="16" pageNumber="417">Distribution.</emphasis>
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The species likely occurs at various localities of dry western
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<collectingCountry id="F3237672FFABC80E0C7BFE24FB4AE46F" box="[1069,1206,511,536]" name="Madagascar" pageId="16" pageNumber="417">Madagascar</collectingCountry>
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, but so far has only been reliably recorded from Kirindy and Tsingy de Bemaraha (Glaw
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<emphasis id="B940EAF0FFABC80E0BF5FDFEFC20E44B" box="[931,988,548,572]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
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1998;
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<bibRefCitation id="EFA54B13FFABC80E0C7DFDFFFB5FE44B" author="Glos" box="[1067,1187,548,573]" pageId="16" pageNumber="417" refString="Glos, J. (2003) The amphibian fauna of the Kirindy dry forest in western Madagascar. Salamandra, 39, 75 - 90." type="journal article" year="2003">Glos 2003</bibRefCitation>
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;
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<bibRefCitation id="EFA54B13FFABC80E0CF9FDFFFA6DE44B" author="Raselimanana" box="[1199,1425,548,573]" pageId="16" pageNumber="417" refString="Raselimanana, A. P. (2008) Herpetofaune des forets seches malgaches. Malagasy Nature, 1, 46 - 75." type="journal article" year="2008">Raselimanana 2008</bibRefCitation>
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).
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<bibRefCitation id="EFA54B13FFABC80E08C1FD9CFE4CE417" author="Rakotondravony" box="[151,432,583,608]" pageId="16" pageNumber="417" refString="Rakotondravony, H. A. (2006 a) Patterns de la diversite des reptiles et amphibiens de la region de Loky-Manambato. In: Goodman, S. M. & Wilme, L. (Eds.), Inventaires de la faune et de la flore du nord de Madagascar dans la region Loky- Manambato, Analamerana et Andavakoera. Recherche pour le Developpement, Serie Sciences Biologiques, 23, 101 - 148." type="journal article" year="2006" yearSuffix="a">Rakotondravony (2006a)</bibRefCitation>
|
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listed the species from Antsahabe in the Loky-Manombato region in north
|
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<collectingCountry id="F3237672FFABC80E0CAAFD9CFA7AE417" box="[1276,1414,583,608]" name="Madagascar" pageId="16" pageNumber="417">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |