435 lines
46 KiB
XML
435 lines
46 KiB
XML
<document id="E4F8074D9A5705464AE565499F32F83D" ID-DOI="10.11646/zootaxa.4755.1.4" ID-GBIF-Dataset="70aae18d-e6ec-4bb1-abe0-27d1f24fbab8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724261" ID-ZooBank="53E41147-52AA-4A16-BB69-3A9279A303AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968507449" checkinUser="plazi" docAuthor="Barr, Cheryl B. & Shepard, William D." docDate="2020" docId="039CB74BFFDC3A57F9DFFA11FA1A93B6" docLanguage="en" docName="zootaxa.4755.1.4.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parygrus , Erichson 1847" docType="treatment" docVersion="5" lastPageNumber="104" masterDocId="FFA5CF33FFDE3A52F948FFB8FF87956E" masterDocTitle="Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)" masterLastPageNumber="128" masterPageNumber="99" pageNumber="101" updateTime="1698730760387" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="8B382D02FEC8F6FE7E0B44D608864A66">
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<mods:title id="2055E24966F18A49E725F98734DF1304">Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)</mods:title>
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<mods:namePart id="0D4AAC648C05A9C5DE27B021FD0A9716">Barr, Cheryl B.</mods:namePart>
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<mods:namePart id="BFE32C1776B007DFD7E41BA50089B4FE">Shepard, William D.</mods:namePart>
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<mods:title id="6EA4E6689D6E3C9C4484C0209D101110">Zootaxa</mods:title>
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<mods:part id="86DDF12C92F3EC70219642775B25FE8F">
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<mods:date id="225962FF35B97FB804DACC81B575CDDE">2020</mods:date>
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<mods:number id="87AE5499DFD2D1412CD61CDA012D0DAA">2020-03-23</mods:number>
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<mods:number id="0379617B3B8F128BC49A04E7C39854EC">4755</mods:number>
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<mods:identifier id="C2CFF59D1B912CD1DD0AB6A3122EBAF3" type="DOI">10.11646/zootaxa.4755.1.4</mods:identifier>
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<treatment id="039CB74BFFDC3A57F9DFFA11FA1A93B6" ID-DOI="http://doi.org/10.5281/zenodo.3728587" ID-GBIF-Taxon="162696908" ID-Zenodo-Dep="3728587" LSID="urn:lsid:plazi:treatment:039CB74BFFDC3A57F9DFFA11FA1A93B6" httpUri="http://treatment.plazi.org/id/039CB74BFFDC3A57F9DFFA11FA1A93B6" lastPageId="5" lastPageNumber="104" pageId="2" pageNumber="101">
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<subSubSection id="C32F55D6FFDC3A50F9DFFA11FD9890AD" box="[151,543,1449,1475]" pageId="2" pageNumber="101" type="nomenclature">
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<paragraph id="8B8A065DFFDC3A50F9DFFA11FD9890AD" blockId="2.[151,543,1377,1475]" box="[151,543,1449,1475]" pageId="2" pageNumber="101">
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<heading id="D0C2B131FFDC3A50F9DFFA11FD9890AD" bold="true" box="[151,543,1449,1475]" fontSize="11" level="1" pageId="2" pageNumber="101" reason="1">
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<emphasis id="B941DA4FFFDC3A50F9DFFA11FD9890AD" bold="true" box="[151,543,1449,1475]" pageId="2" pageNumber="101">
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Genus
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<taxonomicName id="4C357DDEFFDC3A50F9A6FA11FD9890AD" authority="Erichson, 1847" authorityName=", Erichson" authorityYear="1847" box="[238,543,1449,1475]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50F9A6FA11FEDA90AD" bold="true" box="[238,349,1449,1475]" italics="true" pageId="2" pageNumber="101">Parygrus</emphasis>
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<bibRefCitation id="EFA47BACFFDC3A50F82DFA11FD9890AD" author="Erichson, W. F." box="[357,543,1449,1475]" pageId="2" pageNumber="101" refId="ref21285" refString="Erichson, W. F. (1847) Naturgeschichte der Insecten Deutschlands, Erste Abtheilung, Coleoptera. Vol. 3. Verlag der Nicolaischen Buchhandlung, Berlin, 968 pp. https: // doi. org / 10.5962 / bhl. title. 8270" type="book" year="1847">Erichson, 1847</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F55D6FFDC3A50F9DFFA49FB639365" box="[151,1252,1521,1547]" pageId="2" pageNumber="101" type="type_taxon">
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<paragraph id="8B8A065DFFDC3A50F9DFFA49FB639365" blockId="2.[151,1252,1521,1547]" box="[151,1252,1521,1547]" pageId="2" pageNumber="101">
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<emphasis id="B941DA4FFFDC3A50F9DFFA49FEBE9365" bold="true" box="[151,313,1521,1547]" pageId="2" pageNumber="101">
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<typeStatus id="548EB8FFFFDC3A50F9DFFA49FF539365" box="[151,212,1521,1547]" pageId="2" pageNumber="101">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C357DDEFFDC3A50F809FA4AFC8B9365" authority="Waterhouse, 1876" authorityName="Waterhouse" authorityYear="1876" box="[321,780,1521,1547]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="erichsoni">
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<emphasis id="B941DA4FFFDC3A50F809FA4AFDA59365" box="[321,546,1521,1547]" italics="true" pageId="2" pageNumber="101">Parygrus erichsoni</emphasis>
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<bibRefCitation id="EFA47BACFFDC3A50FB61FA49FC8B9365" author="Waterhouse, C. O." box="[553,780,1521,1547]" pageId="2" pageNumber="101" pagination="11 - 25" refId="ref22177" refString="Waterhouse, C. O. (1876) III. On various new genera and species of Coleoptera. Transactions of the Entomological Society of London, 1876 (Part I), 11 - 25. https: // doi. org / 10.1111 / j. 1365 - 2311.1876. tb 01122. x" type="journal article" year="1876">
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<emphasis id="B941DA4FFFDC3A50FB61FA49FC8B9365" bold="true" box="[553,780,1521,1547]" pageId="2" pageNumber="101">Waterhouse, 1876</emphasis>
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</bibRefCitation>
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</taxonomicName>
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<emphasis id="B941DA4FFFDC3A50FA44FA49FB639365" bold="true" box="[780,1252,1521,1547]" pageId="2" pageNumber="101">; designated by subsequent monotypy.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F55D6FFDC3A51F9DFF980FF329245" lastPageId="3" lastPageNumber="102" pageId="2" pageNumber="101" type="diagnosis">
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<paragraph id="8B8A065DFFDC3A50F9DFF980FD579221" blockId="2.[151,1437,1592,1979]" pageId="2" pageNumber="101">
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<emphasis id="B941DA4FFFDC3A50F9DFF980FE95933D" bold="true" box="[151,274,1592,1619]" pageId="2" pageNumber="101">Diagnosis.</emphasis>
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<taxonomicName id="4C357DDEFFDC3A50F852F982FE05933D" authorityName=", Erichson" authorityYear="1847" box="[282,386,1594,1619]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50F852F982FE05933D" box="[282,386,1594,1619]" italics="true" pageId="2" pageNumber="101">Parygrus</emphasis>
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</taxonomicName>
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shares antennal morphology with
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<taxonomicName id="4C357DDEFFDC3A50FA46F981FCF5933D" authorityName="Erichson" authorityYear="1847" box="[782,882,1593,1619]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FA46F981FCF5933D" box="[782,882,1593,1619]" italics="true" pageId="2" pageNumber="101">Helichus</emphasis>
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</taxonomicName>
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, but the two genera can be separated by the following characters:
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<taxonomicName id="4C357DDEFFDC3A50F823F9E6FE549319" authorityName=", Erichson" authorityYear="1847" box="[363,467,1630,1655]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50F823F9E6FE549319" box="[363,467,1630,1655]" italics="true" pageId="2" pageNumber="101">Parygrus</emphasis>
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</taxonomicName>
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is very pubescent, covered with erect and semi-erect setae, including the eyes; plastron setae are not present; the enlarged 2nd antennomere covers antennomeres 3–4 or 3–5.
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<taxonomicName id="4C357DDEFFDC3A50FD37F939FB6493F5" authorityName="Erichson" authorityYear="1847" box="[1151,1251,1665,1691]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FD37F939FB6493F5" box="[1151,1251,1665,1691]" italics="true" pageId="2" pageNumber="101">Helichus</emphasis>
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</taxonomicName>
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is covered with recumbent plastron setae and lacks erect setae, the eyes are bare; the enlarged 2nd antennomere covers all of the distal antennomeres. Because of its pubescence,
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<taxonomicName id="4C357DDEFFDC3A50FBF3F972FCA4938D" authorityName=", Erichson" authorityYear="1847" box="[699,803,1738,1763]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FBF3F972FCA4938D" box="[699,803,1738,1763]" italics="true" pageId="2" pageNumber="101">Parygrus</emphasis>
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</taxonomicName>
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appears most similar to
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<taxonomicName id="4C357DDEFFDC3A50FD70F971FB33938D" authorityName="Erichson" authorityYear="1847" box="[1080,1204,1737,1763]" class="Insecta" family="Curculionidae" genus="Pelonomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FD70F971FB33938D" box="[1080,1204,1737,1763]" italics="true" pageId="2" pageNumber="101">Pelonomus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C357DDEFFDC3A50FD8BF972FAD3938D" authorityName="Shepard" authorityYear="2019" box="[1219,1364,1738,1763]" class="Hexapoda" family="Dryopidae" genus="Microparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FD8BF972FAD3938D" box="[1219,1364,1738,1763]" italics="true" pageId="2" pageNumber="101">Microparnus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C357DDEFFDC3A50FC2BF971FED59269" authority="Spangler" authorityName="Spangler" authorityYear="1980" class="Insecta" family="Dryopidae" genus="Onopelmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FC2BF971FF619269" italics="true" pageId="2" pageNumber="101">Onopelmus</emphasis>
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Spangler
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</taxonomicName>
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,
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<taxonomicName id="4C357DDEFFDC3A50F815F956FD829269" authority="Olivier" authorityName="Olivier" authorityYear="1791" box="[349,517,1773,1799]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50F815F956FE289269" box="[349,431,1774,1799]" italics="true" pageId="2" pageNumber="101">Dryops</emphasis>
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Olivier
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</taxonomicName>
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and
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<taxonomicName id="4C357DDEFFDC3A50FB72F955FD449269" authorityName="Shepard and Barr" authorityYear="2018" box="[570,707,1773,1799]" class="Insecta" family="Dryopidae" genus="Platyparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FB72F955FD449269" box="[570,707,1773,1799]" italics="true" pageId="2" pageNumber="101">Platyparnus</emphasis>
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</taxonomicName>
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. These genera differ from
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<taxonomicName id="4C357DDEFFDC3A50FAA3F956FBD49269" authorityName=", Erichson" authorityYear="1847" box="[1003,1107,1774,1799]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FAA3F956FBD49269" box="[1003,1107,1774,1799]" italics="true" pageId="2" pageNumber="101">Parygrus</emphasis>
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</taxonomicName>
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by the following:
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<taxonomicName id="4C357DDEFFDC3A50FC68F955FA1B9269" authorityName="Erichson" authorityYear="1847" box="[1312,1436,1773,1799]" class="Insecta" family="Curculionidae" genus="Pelonomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FC68F955FA1B9269" box="[1312,1436,1773,1799]" italics="true" pageId="2" pageNumber="101">Pelonomus</emphasis>
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</taxonomicName>
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does not have an enlarged 2nd antennomere;
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<taxonomicName id="4C357DDEFFDC3A50FBC3F8AAFC9B9245" authorityName="Shepard" authorityYear="2019" box="[651,796,1810,1835]" class="Hexapoda" family="Dryopidae" genus="Microparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FBC3F8AAFC9B9245" box="[651,796,1810,1835]" italics="true" pageId="2" pageNumber="101">Microparnus</emphasis>
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</taxonomicName>
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has 10 antennomeres and
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<taxonomicName id="4C357DDEFFDC3A50FD08F8A9FB479245" authorityName="Spangler" authorityYear="1980" box="[1088,1216,1809,1835]" class="Insecta" family="Dryopidae" genus="Onopelmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FD08F8A9FB479245" box="[1088,1216,1809,1835]" italics="true" pageId="2" pageNumber="101">Onopelmus</emphasis>
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</taxonomicName>
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has 13;
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<taxonomicName id="4C357DDEFFDC3A50FC53F8AAFAEA9245" authorityName="Olivier" authorityYear="1791" box="[1307,1389,1810,1835]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50FC53F8AAFAEA9245" box="[1307,1389,1810,1835]" italics="true" pageId="2" pageNumber="101">Dryops</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C357DDEFFDC3A50F9DFF88DFEA79221" authorityName="Shepard and Barr" authorityYear="2018" box="[151,288,1845,1871]" class="Insecta" family="Dryopidae" genus="Platyparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
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<emphasis id="B941DA4FFFDC3A50F9DFF88DFEA79221" box="[151,288,1845,1871]" italics="true" pageId="2" pageNumber="101">Platyparnus</emphasis>
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</taxonomicName>
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have pronotal sublateral carinae/sulci.
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</paragraph>
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<paragraph id="8B8A065DFFDC3A50F98FF8E0FD1B92D5" blockId="2.[151,1437,1592,1979]" pageId="2" pageNumber="101">
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<emphasis id="B941DA4FFFDC3A50F98FF8E0FE54921D" bold="true" box="[199,467,1880,1907]" pageId="2" pageNumber="101">Generic redescription.</emphasis>
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Size from
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<quantity id="4CCDABB8FFDC3A50FB1AF8E1FD36921D" box="[594,689,1881,1907]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="2" pageNumber="101" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
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long; elongate, parallel-sided or somewhat wider at posterior 1/3, moderately convex in lateral view; color uniformly brown; very setose, covered with long, stout, erect and/or semierect setae and shorter, finer, recumbent setae.
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</paragraph>
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<paragraph id="8B8A065DFFDD3A51F98FFF21FB4794E5" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
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<emphasis id="B941DA4FFFDD3A51F98FFF21FE8395DD" box="[199,260,153,179]" italics="true" pageId="3" pageNumber="102">Head</emphasis>
|
||
covered with erect setae; frons projecting slightly or prominently forward between antennal bases; vertex flattened or shallowly depressed between eyes. Eyes hemispherical, protuberant, covered with erect setae, finely faceted. Antennae densely setose, pectinate, with 11 antennomeres; antennomere 2 enlarged, trapezoid, covering antennomeres 3–5 or 3–6; antennomere 3 shortest; antennomeres 4–11 forming a loose club; subantennal grooves deep below eyes; insertions approximate. Clypeus sub-triangular, wider than long. Labrum transverse, shorter than clypeus. Mandible with four teeth; ventral two teeth larger, acutely pointed; dorsal two teeth smaller, blunt, closely appressed. Maxillary palpi each with four palpomeres; labial palpi each with three palpomeres.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FFE2EFD569775" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FFE2EFEB194C1" box="[199,310,406,431]" italics="true" pageId="3" pageNumber="102">Pronotum</emphasis>
|
||
wider (~1.25x) than long; narrowly marginate and sometimes explanate laterally; anterior angles very acute; posterior angles barely acute, depressed; carinae absent, disc slightly flattened; punctate with large, distinct punctures interspersed with very small, fine punctures. Hypomeron flat, wide, narrower near anterior margin.
|
||
<emphasis id="B941DA4FFFDD3A51F9DFFDB9FE889775" box="[151,271,513,539]" italics="true" pageId="3" pageNumber="102">
|
||
<taxonomicName id="4C357DDEFFDD3A51F9DFFDB9FE8F9775" box="[151,264,513,539]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
shape variable, anterior margin arcuate.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FFD9DFD5397E6" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FFD9DFE9D9751" box="[199,282,549,575]" italics="true" pageId="3" pageNumber="102">Elytron</emphasis>
|
||
without carinae; finely or coarsely punctate-striate, with nine rows of punctures; narrowly margined. Epipleuron sinuate, broad in basal 1/3, narrow in apical 2/3 with groove to receive lateral edge of abdominal ventrite 5.
|
||
<emphasis id="B941DA4FFFDD3A51F9FBFDD5FEB597E9" box="[179,306,621,647]" italics="true" pageId="3" pageNumber="102">Hind wing:</emphasis>
|
||
macropterous (in species examined).
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FFD2AFD069631" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FFD2AFF7B97C5" box="[199,252,658,683]" italics="true" pageId="3" pageNumber="102">Legs</emphasis>
|
||
with major structures similarly proportioned; proleg shortest, metaleg longest. Procoxae oblong and flat, more widely separated than meso- and metacoxae; mesocoxae globular, separated by about one coxal width; metacoxae transverse, separated by about same distance as mesocoxae, excavate to receive femora; metatrochanters swollen; protibia weakly to strongly arcuate, sometimes deflexed (males); mesotibia weakly to moderately arcuate; metatibia straight; tarsomeres increasingly longer from base to apex, tarsomeres 1 and 2 very short, tarsomere 5 as long as previous 4 combined; claws simple.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FFCD2FCE991CD" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FFCD2FE8896ED" box="[199,271,874,899]" italics="true" pageId="3" pageNumber="102">Venter</emphasis>
|
||
with short, fine, recumbent setae present on lateral surfaces including hypomera, pro-, meso- and metepisterna, mesepimera, epipleura; long, semi-recumbent setae present medially. Prosternum long; lateral surface angled at about 45° to central disc; prosternal process broad between procoxae, laterally margined, with a median, longitudinal carina terminating in a low protuberance. Mesoventrite very short, deeply excavate to receive prosternal spine. Metaventrite long, projecting anteriorly between mesocoxae, margined; lateral surface angled at about 45° to posterior disc; metathoracic discrimen distinct; posterior border bidentate between metacoxae; metanepisternum wider anteriorly than posteriorly. Abdomen convex, with 5 ventrites; punctures aligned with faint, transverse strigae; ventrite 1 with margined, triangular intercoxal process, posterior border straight; ventrites 2–4 transverse, subequal in length; ventrite 5 longer, with broadly rounded apex.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FFB15FB6591A6" blockId="3.[151,1437,153,1835]" box="[199,1250,1197,1224]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FFB15FEBE91A9" box="[199,313,1197,1223]" italics="true" pageId="3" pageNumber="102">Genitalia.</emphasis>
|
||
Male genitalia, trilobate
|
||
<typeStatus id="548EB8FFFFDD3A51FB1AFB15FD0191A9" box="[594,646,1197,1223]" pageId="3" pageNumber="102">type</typeStatus>
|
||
; female genitalia, common
|
||
<typeStatus id="548EB8FFFFDD3A51FAF4FB15FC6B91A9" box="[956,1004,1197,1223]" pageId="3" pageNumber="102">type</typeStatus>
|
||
(
|
||
<bibRefCitation id="EFA47BACFFDD3A51FAB3FB15FB5191A6" author="Kodada, J. & Jach, M. A. & Ciampor, F." box="[1019,1238,1197,1224]" pageId="3" pageNumber="102" pagination="590 - 602" refId="ref21609" refString="Kodada, J., Jach, M. A. & Ciampor, F. (2016) 19.3 Dryopidae Bilberg, 1820 (1817). In: Beutel, R. G. & Leschen, R. A. B. (Eds.), Coleoptera, Beetles. Vol. 1. Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Beutel, R. G. & Kristensen, N. P. (Eds.), Handbook of Zoology, Arthropoda: Insecta, Second Edition. Walter de Gruyter, Berlin, pp. 590 - 602. https: // doi. org / 10.1515 / 9783110373929" type="book chapter" year="2016">
|
||
Kodada
|
||
<emphasis id="B941DA4FFFDD3A51FD12FB16FB1491A9" box="[1114,1171,1197,1223]" italics="true" pageId="3" pageNumber="102">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FFB68FC5D90F1" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FFB68FE469185" bold="true" box="[199,449,1232,1259]" pageId="3" pageNumber="102">Original description.</emphasis>
|
||
<bibRefCitation id="EFA47BACFFDD3A51F880FB69FDF99185" author="Erichson, W. F." box="[456,638,1233,1259]" pageId="3" pageNumber="102" refId="ref21285" refString="Erichson, W. F. (1847) Naturgeschichte der Insecten Deutschlands, Erste Abtheilung, Coleoptera. Vol. 3. Verlag der Nicolaischen Buchhandlung, Berlin, 968 pp. https: // doi. org / 10.5962 / bhl. title. 8270" type="book" year="1847">Erichson’s 1847</bibRefCitation>
|
||
descriptions of
|
||
<taxonomicName id="4C357DDEFFDD3A51FA7CFB6AFC1B9185" authorityName=", Erichson" authorityYear="1847" box="[820,924,1234,1259]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51FA7CFB6AFC1B9185" box="[820,924,1234,1259]" italics="true" pageId="3" pageNumber="102">Parygrus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C357DDEFFDD3A51FA9CFB69FBBF9185" authorityName="Erichson" authorityYear="1847" box="[980,1080,1233,1259]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51FA9CFB69FBBF9185" box="[980,1080,1233,1259]" italics="true" pageId="3" pageNumber="102">Helichus</emphasis>
|
||
</taxonomicName>
|
||
were presented in an annotated key. He cited the following characters for separating genera with “auriculate” antennae: Middle legs close to each other (
|
||
<taxonomicName id="4C357DDEFFDD3A51F9ABFAA2FECC905D" authorityName=", Erichson" authorityYear="1847" box="[227,331,1306,1331]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51F9ABFAA2FECC905D" box="[227,331,1306,1331]" italics="true" pageId="3" pageNumber="102">Parygrus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C357DDEFFDD3A51F8CEFAA2FE50905D" authorityName="Fabricius" authorityYear="1792" box="[390,471,1306,1331]" class="Insecta" family="Dryopidae" genus="Parnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51F8CEFAA2FE50905D" box="[390,471,1306,1331]" italics="true" pageId="3" pageNumber="102">Parnus</emphasis>
|
||
</taxonomicName>
|
||
); middle legs further apart (
|
||
<taxonomicName id="4C357DDEFFDD3A51FA56FAA1FC05905D" authorityName="Erichson" authorityYear="1847" box="[798,898,1305,1331]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51FA56FAA1FC05905D" box="[798,898,1305,1331]" italics="true" pageId="3" pageNumber="102">Helichus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C357DDEFFDD3A51FAF4FAA2FB89905D" authorityName="Olivier" authorityYear="1791" box="[956,1038,1306,1331]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51FAF4FAA2FB89905D" box="[956,1038,1306,1331]" italics="true" pageId="3" pageNumber="102">Dryops</emphasis>
|
||
</taxonomicName>
|
||
). In addition, he stated that in
|
||
<taxonomicName id="4C357DDEFFDD3A51FC3CFAA2FF679039" authorityName=", Erichson" authorityYear="1847" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51FC3CFAA2FF679039" italics="true" pageId="3" pageNumber="102">Parygrus</emphasis>
|
||
</taxonomicName>
|
||
the elytra are clearly punctate-striate, and the antenna clearly “11-membered.”
|
||
<bibRefCitation id="EFA47BACFFDD3A51FD3CFA85FAC99036" author="Waterhouse, C. O." box="[1140,1358,1341,1368]" pageId="3" pageNumber="102" pagination="11 - 25" refId="ref22177" refString="Waterhouse, C. O. (1876) III. On various new genera and species of Coleoptera. Transactions of the Entomological Society of London, 1876 (Part I), 11 - 25. https: // doi. org / 10.1111 / j. 1365 - 2311.1876. tb 01122. x" type="journal article" year="1876">Waterhouse (1876)</bibRefCitation>
|
||
, in his paper describing the
|
||
<typeStatus id="548EB8FFFFDD3A51F837FAD9FE289015" box="[383,431,1377,1403]" pageId="3" pageNumber="102">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C357DDEFFDD3A51FB45FADAFD089015" authorityName="Waterhouse" authorityYear="1876" box="[525,655,1377,1403]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="erichsoni">
|
||
<emphasis id="B941DA4FFFDD3A51FB45FADAFD089015" box="[525,655,1377,1403]" italics="true" pageId="3" pageNumber="102">P. erichsoni</emphasis>
|
||
</taxonomicName>
|
||
, stated that Erichson had established the genus to include undescribed species from South America with narrow form and strongly striated elytra.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FFA10FF67933D" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FFA10FDE390AD" bold="true" box="[199,612,1448,1475]" pageId="3" pageNumber="102">Sexual variation and dimorphism.</emphasis>
|
||
Females are generally larger than males, and have a small, angular protuberance at the middle apex of abdominal ventrite 5 which is formed by a ventrally-directed fold. This feature is absent in males and therefore very useful in externally determining the sex of individuals. In the specimens we have examined, the protibiae of females were not deflexed and were usually less arcuate than those of conspecific males.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FF9E4FAF493AE" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FF9E4FE4F9319" bold="true" box="[199,456,1628,1655]" pageId="3" pageNumber="102">Reproductive period.</emphasis>
|
||
Fifteen females were collected along with
|
||
<specimenCount id="9D33CDD4FFDD3A51FA8FF9E5FBA99319" box="[967,1070,1629,1655]" pageId="3" pageNumber="102" type="male">11 males</specimenCount>
|
||
from
|
||
<collectingCountry id="F32246CDFFDD3A51FD33F9E5FB639319" box="[1147,1252,1629,1655]" name="Paraguay" pageId="3" pageNumber="102">Paraguay</collectingCountry>
|
||
,
|
||
<collectingRegion id="49F1C8BFFFDD3A51FDBAF9E5FA1F9319" box="[1266,1432,1629,1655]" country="Paraguay" name="Alto Paraguay" pageId="3" pageNumber="102">Alto Paraguay</collectingRegion>
|
||
, Puerto Esperanza,
|
||
<date id="FF8B209DFFDD3A51F822F939FDCF93F5" box="[362,584,1664,1691]" pageId="3" pageNumber="102" value="2006-12-02">
|
||
2
|
||
<superScript id="7C40AB15FFDD3A51F830F938FE0F93E0" attach="left" box="[376,392,1664,1678]" fontSize="6" pageId="3" pageNumber="102">nd</superScript>
|
||
December 2006
|
||
</date>
|
||
. Dissection found eggs in 13 of the
|
||
<specimenCount id="9D33CDD4FFDD3A51FAA9F938FBE793F5" box="[993,1120,1664,1691]" pageId="3" pageNumber="102" type="female">15 females</specimenCount>
|
||
. Thus the austral summer is the reproductive period. This is also true for
|
||
<taxonomicName id="4C357DDEFFDD3A51FBC3F91DFC8093D1" authorityName="Erichson" authorityYear="1847" box="[651,775,1701,1727]" class="Insecta" family="Curculionidae" genus="Pelonomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51FBC3F91DFC8093D1" box="[651,775,1701,1727]" italics="true" pageId="3" pageNumber="102">Pelonomus</emphasis>
|
||
</taxonomicName>
|
||
spp. from
|
||
<collectingCountry id="F32246CDFFDD3A51FAC8F91DFC6F93D1" box="[896,1000,1701,1727]" name="Paraguay" pageId="3" pageNumber="102">Paraguay</collectingCountry>
|
||
(W.D. Shepard, unpublished data).
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDD3A51F98FF970FF329245" blockId="3.[151,1437,153,1835]" pageId="3" pageNumber="102">
|
||
<emphasis id="B941DA4FFFDD3A51F98FF970FE76938D" bold="true" box="[199,497,1736,1763]" pageId="3" pageNumber="102">Geographic distribution.</emphasis>
|
||
<taxonomicName id="4C357DDEFFDD3A51F8B2F972FDE5938D" authorityName=", Erichson" authorityYear="1847" box="[506,610,1738,1763]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDD3A51F8B2F972FDE5938D" box="[506,610,1738,1763]" italics="true" pageId="3" pageNumber="102">Parygrus</emphasis>
|
||
</taxonomicName>
|
||
is recorded from
|
||
<collectingCountry id="F32246CDFFDD3A51FA78F971FC0F938D" box="[816,904,1737,1763]" name="Mexico" pageId="3" pageNumber="102">Mexico</collectingCountry>
|
||
, south through Central America and the South American countries of
|
||
<collectingCountry id="F32246CDFFDD3A51F8D6F955FD939269" box="[414,532,1773,1799]" name="Venezuela" pageId="3" pageNumber="102">Venezuela</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32246CDFFDD3A51FB6AF955FD149269" box="[546,659,1773,1799]" name="Colombia" pageId="3" pageNumber="102">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32246CDFFDD3A51FBE8F955FD789269" box="[672,767,1773,1799]" name="Ecuador" pageId="3" pageNumber="102">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32246CDFFDD3A51FA45F955FCC49269" box="[781,835,1773,1799]" name="Peru" pageId="3" pageNumber="102">Peru</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32246CDFFDD3A51FA18F955FC229269" box="[848,933,1773,1799]" name="Bolivia" pageId="3" pageNumber="102">Bolivia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32246CDFFDD3A51FAFBF955FB9B9269" box="[947,1052,1773,1799]" name="Paraguay" pageId="3" pageNumber="102">Paraguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32246CDFFDD3A51FD61F955FBF79269" box="[1065,1136,1773,1799]" name="Brazil" pageId="3" pageNumber="102">Brazil</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F32246CDFFDD3A51FDE6F955FA9A9269" box="[1198,1309,1773,1799]" name="Argentina" pageId="3" pageNumber="102">Argentina</collectingCountry>
|
||
(Figs 2, 3, 4
|
||
<emphasis id="B941DA4FFFDD3A51F9EDF8A8FF329245" bold="true" box="[165,181,1808,1835]" pageId="3" pageNumber="102">).</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4A56D5FFDB3A57F9DFFB9DFB1E9150" ID-DOI="http://doi.org/10.5281/zenodo.3724265" ID-Zenodo-Dep="3724265" box="[151,1177,1061,1086]" httpUri="https://zenodo.org/record/3724265/files/figure.png" pageId="5" pageNumber="104" startId="5.[151,250,1061,1086]" targetBox="[231,1355,181,1035]" targetPageId="5">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFB9DFB1E9150" blockId="5.[151,1177,1061,1189]" box="[151,1177,1061,1086]" pageId="5" pageNumber="104">
|
||
<emphasis id="B941DA4FFFDB3A57F9DFFB9DFE949150" bold="true" box="[151,275,1061,1086]" pageId="5" pageNumber="104">FIGURE 4.</emphasis>
|
||
Geographical distribution map of
|
||
<taxonomicName id="4C357DDEFFDB3A57FB39FB9FFC8B9150" authority="Barr & Shepard, 2020" authorityName="Barr & Shepard" authorityYear="2020" box="[625,780,1063,1086]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="maya" status="sp. nov.">
|
||
<emphasis id="B941DA4FFFDB3A57FB39FB9FFC8B9150" box="[625,780,1063,1086]" italics="true" pageId="5" pageNumber="104">Parygrus maya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B941DA4FFFDB3A57FA5BFB9EFCCB9150" bold="true" box="[787,844,1062,1086]" pageId="5" pageNumber="104">
|
||
<taxonomicNameLabel id="A2726734FFDB3A57FA5BFB9EFCCB9150" box="[787,844,1062,1086]" pageId="5" pageNumber="104" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in Mexico and Central America.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32F55D6FFDB3A57F9DFFB33FA1A93B6" pageId="5" pageNumber="104" type="key">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFB33FC7691CB" blockId="5.[151,1177,1061,1189]" box="[151,1009,1163,1189]" pageId="5" pageNumber="104">
|
||
<emphasis id="B941DA4FFFDB3A57F9DFFB33FC7691CB" bold="true" box="[151,1009,1163,1189]" pageId="5" pageNumber="104">
|
||
Key to the genera of Neotropical aquatic and semiaquatic
|
||
<taxonomicName id="4C357DDEFFDB3A57FA25FB33FC7691CB" ID-CoL="9H4" box="[877,1009,1163,1189]" class="Insecta" family="Dryopidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="family">Dryopidae</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<key id="FFA4516CFFDB3A57F9DFFB68FA1A93B6" pageId="5" pageNumber="104">
|
||
<keyStep id="30C11EF8FFDB3A57F9DFFB68FA1B904E" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFB68FA1B906D" blockId="5.[151,1437,1232,1752]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFB68FA1B906D" pageId="5" pageNumber="104">
|
||
1 Body dorsally glabrous; lateral margin of each elytron interrupted near apex by a densely pubescent depression................................................................................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FDA1FB55FA1B906D" authority="Sharp" authorityName="Sharp" authorityYear="1882" box="[1257,1436,1261,1283]" class="Insecta" family="Dryopidae" genus="Elmoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FDA1FB55FAD9906C" box="[1257,1374,1261,1282]" italics="true" pageId="5" pageNumber="104">Elmoparnus</emphasis>
|
||
Sharp
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFAB3FA1B904E" blockId="5.[151,1437,1232,1752]" box="[151,1436,1290,1312]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFAB3FA1B904E" box="[151,1436,1290,1312]" pageId="5" pageNumber="104">– Body dorsally pubescent or with plastron; lateral margin of each elytron without densely pubescent depression.......... 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C11EF8FFDB3A57F9DFFA90FA1B9034" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFA90FA1B9050" blockId="5.[151,1437,1232,1752]" box="[151,1436,1320,1342]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFA90FA1B9050" box="[151,1436,1320,1342]" pageId="5" pageNumber="104">
|
||
2 Pronotum with complete sublateral carina/sulcus on each side.......................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FC44FA90FA1B9050" ID-CoL="48JK" authority="Olivier" authorityName="Olivier" authorityYear="1791" box="[1292,1436,1320,1342]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FC44FA90FAD49053" box="[1292,1363,1320,1341]" italics="true" pageId="5" pageNumber="104">Dryops</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFAFEFA1B9034" blockId="5.[151,1437,1232,1752]" box="[151,1436,1349,1371]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFAFEFA1B9034" box="[151,1436,1349,1371]" pageId="5" pageNumber="104">– Pronotum without sublateral carinae/sulci or with only short carinae/sulci........................................ 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C11EF8FFDB3A57F9DFFADBFA1B90FB" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFADBFA1B9016" blockId="5.[151,1437,1232,1752]" box="[151,1436,1378,1400]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFADBFA1B9016" box="[151,1436,1378,1400]" pageId="5" pageNumber="104">3 Pronotum with sublateral carinae/sulci only in basal 1/4 to 1/3 .................................................. 8</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFA38FA1B90FB" blockId="5.[151,1437,1232,1752]" box="[151,1436,1408,1430]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFA38FA1B90FB" box="[151,1436,1408,1430]" pageId="5" pageNumber="104">– Pronotum without sublateral carinae/sulci.................................................................. 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C11EF8FFDB3A57F9DFFA25FA1B90BE" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFA25FA1B90DD" blockId="5.[151,1437,1232,1752]" box="[151,1436,1437,1459]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFA25FA1B90DD" box="[151,1436,1437,1459]" pageId="5" pageNumber="104">
|
||
4 Each antenna with 13 antennomeres......................................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FD9CFA25FA1B90DD" authority="Spangler" authorityName="Spangler" authorityYear="1980" box="[1236,1436,1437,1459]" class="Insecta" family="Dryopidae" genus="Onopelmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FD9CFA25FAC690DC" box="[1236,1345,1437,1458]" italics="true" pageId="5" pageNumber="104">Onopelmus</emphasis>
|
||
Spangler
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFA03FA1B90BE" blockId="5.[151,1437,1232,1752]" box="[151,1436,1466,1488]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFA03FA1B90BE" box="[151,1436,1466,1488]" pageId="5" pageNumber="104">– Each antenna with 9 or 11 antennomeres.................................................................. 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C11EF8FFDB3A57F9DFFA60FA1B9365" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFA60FA1B9080" blockId="5.[151,1437,1232,1752]" box="[151,1436,1496,1518]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFA60FA1B9080" box="[151,1436,1496,1518]" pageId="5" pageNumber="104">
|
||
5 Each antenna with 9 antennomeres........................................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FD92FA60FA1B9080" authority="Spangler" authorityName="Spangler" authorityYear="1991" box="[1242,1436,1496,1518]" class="Insecta" family="Dryopidae" genus="Guaranius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FD92FA60FAC69083" box="[1242,1345,1496,1517]" italics="true" pageId="5" pageNumber="104">Guaranius</emphasis>
|
||
Spangler
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFFA4EFA1B9365" blockId="5.[151,1437,1232,1752]" box="[151,1436,1525,1547]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFFA4EFA1B9365" box="[151,1436,1525,1547]" pageId="5" pageNumber="104">– Each antenna with 11 antennomeres.................................................................... ... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C11EF8FFDB3A57F9DFF9AAFA1B932B" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFF9AAFA1B9346" blockId="5.[151,1437,1232,1752]" box="[151,1436,1554,1576]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFF9AAFA1B9346" box="[151,1436,1554,1576]" pageId="5" pageNumber="104">
|
||
6 Antennomere 2 enlarged and covering rest of antennomeres.....................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FDA4F9AAFA1B9346" ID-CoL="635L2" authority="Erichson" authorityName="Erichson" authorityYear="1847" box="[1260,1436,1554,1576]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FDA4F9AAFAC69349" box="[1260,1345,1554,1575]" italics="true" pageId="5" pageNumber="104">Helichus</emphasis>
|
||
Erichson
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFF988FA1B932B" blockId="5.[151,1437,1232,1752]" box="[151,1436,1584,1606]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFF988FA1B932B" box="[151,1436,1584,1606]" pageId="5" pageNumber="104">– Antennomere 2 may be enlarged but not covering rest of antennomeres.......................................... 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C11EF8FFDB3A57F9DFF9F6FA1B93EE" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFF9F6FA1B930D" blockId="5.[151,1437,1232,1752]" box="[151,1436,1613,1635]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFF9F6FA1B930D" box="[151,1436,1613,1635]" pageId="5" pageNumber="104">
|
||
7 Antennomere 2 enlarged and at least covering antennomeres 3 and 4................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FDA0F9F6FA1B930D" authority="Erichson" authorityName=", Erichson" authorityYear="1847" box="[1256,1436,1613,1635]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FDA0F9F6FAC6930D" box="[1256,1345,1614,1635]" italics="true" pageId="5" pageNumber="104">Parygrus</emphasis>
|
||
Erichson
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFF9D3FA1B93EE" blockId="5.[151,1437,1232,1752]" box="[151,1436,1642,1664]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFF9D3FA1B93EE" box="[151,1436,1642,1664]" pageId="5" pageNumber="104">
|
||
– Antennomere 2 not enlarged.............................................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FD9EF9D2FA1B93EE" authority="Erichson" authorityName="Erichson" authorityYear="1847" box="[1238,1436,1642,1664]" class="Insecta" family="Curculionidae" genus="Pelonomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FD9EF9D2FAC69311" box="[1238,1345,1642,1663]" italics="true" pageId="5" pageNumber="104">Pelonomus</emphasis>
|
||
Erichson
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C11EF8FFDB3A57F9DFF930FA1A93B6" pageId="5" pageNumber="104">
|
||
<paragraph id="8B8A065DFFDB3A57F9DFF930FA1A93D5" blockId="5.[151,1437,1232,1752]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFF930FA1A93D5" pageId="5" pageNumber="104">
|
||
8 Length
|
||
<quantity id="4CCDABB8FFDB3A57F864F930FE0D93F3" box="[300,394,1672,1694]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="5" pageNumber="104" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 mm</quantity>
|
||
; each antenna with 11 antennomeres; pro- and mesotarsus with tarsomere 2 wide and flattened in males...............................................................................
|
||
<taxonomicName id="4C357DDEFFDB3A57FD31F91DFA1A93D5" authority="Shepard and Barr" authorityName="Shepard and Barr" authorityYear="2018" box="[1145,1437,1701,1723]" class="Insecta" family="Dryopidae" genus="Platyparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FD31F91DFB6993D4" box="[1145,1262,1701,1722]" italics="true" pageId="5" pageNumber="104">Platyparnus</emphasis>
|
||
Shepard and Barr
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFDB3A57F9DFF97BFA1A93B6" blockId="5.[151,1437,1232,1752]" box="[151,1437,1730,1752]" pageId="5" pageNumber="104">
|
||
<keyLead id="30C4A568FFDB3A57F9DFF97BFA1A93B6" box="[151,1437,1730,1752]" pageId="5" pageNumber="104">
|
||
– Length
|
||
<quantity id="4CCDABB8FFDB3A57F864F97BFEF993B6" box="[300,382,1731,1752]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="104" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
; each antenna with 10 antennomeres; no tarsomeres wide and flattened........... ..
|
||
<taxonomicName id="4C357DDEFFDB3A57FD83F97BFA1A93B6" authority="Shepard" authorityName="Shepard" authorityYear="2019" box="[1227,1437,1730,1752]" class="Hexapoda" family="Dryopidae" genus="Microparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFDB3A57FD83F97BFAC093B6" box="[1227,1351,1731,1752]" italics="true" pageId="5" pageNumber="104">Microparnus</emphasis>
|
||
Shepard
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |