315 lines
35 KiB
XML
315 lines
35 KiB
XML
<document id="9C85444BA82E852BC3DB01D34CE9B8F3" ID-DOI="10.11646/zootaxa.4755.1.4" ID-GBIF-Dataset="70aae18d-e6ec-4bb1-abe0-27d1f24fbab8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724261" ID-ZooBank="53E41147-52AA-4A16-BB69-3A9279A303AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968507449" checkinUser="plazi" docAuthor="Barr, Cheryl B. & Shepard, William D." docDate="2020" docId="039CB74BFFCC3A46F9DFFE65FDC791A9" docLanguage="en" docName="zootaxa.4755.1.4.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parygrus lengua Barr and Shepard 2020, new species" docType="treatment" docUuid="8995529C-3873-46EC-B9F1-E030A1A07509" docUuidSource="ZooBank" docVersion="6" lastPageNumber="119" masterDocId="FFA5CF33FFDE3A52F948FFB8FF87956E" masterDocTitle="Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)" masterLastPageNumber="128" masterPageNumber="99" pageNumber="117" updateTime="1698730760387" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="30B5FB667C402BC0AB049F1A4485F0C9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9F4FE3698F6E27B51EEB355189FA0FD4">
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<mods:title id="E8AE53C3D5C30A489077DDB97E2FE7EC">Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B7C5F034693BAEF43FCE44A51AB65726" type="personal">
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<mods:roleTerm id="9E379A296CFA9C799D3997A300467B09">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="91071307307E157A86BB53B582D28586">Barr, Cheryl B.</mods:namePart>
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<mods:name id="23BD7CAC0C0B8DC122ABF9A6BBB6D30B" type="personal">
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<mods:roleTerm id="6EF2EAF40216A316FF4A5D3EC174312A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="703A403F235CFF95994DD6945D9E36AC">Shepard, William D.</mods:namePart>
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<mods:typeOfResource id="151A515C58EC946A2471278105A2FB1D">text</mods:typeOfResource>
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<mods:title id="9D3BD79BA8D1818C38C760174B749F69">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F6DF5B5C3F8FA88A86250269B7FA3382">
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<mods:date id="C79E4392B681B7A6320DBB96AA4B3733">2020</mods:date>
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<mods:number id="79FAA30AE6689F84E66923754E68FB35">2020-03-23</mods:number>
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<mods:number id="9893329FE001656820EF24C39659B91D">4755</mods:number>
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<mods:detail id="3FDAC7EF2AAEAD1E774A0295DDEBE230" type="issue">
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<mods:classification id="BBE8915B0C7AC1A1EA9B73717C806781">journal article</mods:classification>
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<mods:identifier id="11363AFDD98038C396F0D2C008D77E55" type="DOI">10.11646/zootaxa.4755.1.4</mods:identifier>
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<mods:identifier id="185E6BEF45C841E1050126D9479CEBFC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039CB74BFFCC3A46F9DFFE65FDC791A9" ID-DOI="http://doi.org/10.5281/zenodo.3730213" ID-GBIF-Taxon="162696916" ID-Zenodo-Dep="3730213" LSID="urn:lsid:zoobank.org:act:8995529C-3873-46EC-B9F1-E030A1A07509" httpUri="http://treatment.plazi.org/id/039CB74BFFCC3A46F9DFFE65FDC791A9" lastPageId="20" lastPageNumber="119" pageId="18" pageNumber="117">
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<subSubSection id="C32F55D6FFCC3A40F9DFFE65FEC49775" pageId="18" pageNumber="117" type="nomenclature">
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<paragraph id="8B8A065DFFCC3A40F9DFFE65FD6D9498" blockId="18.[151,884,476,611]" box="[151,746,476,503]" pageId="18" pageNumber="117">
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<heading id="D0C2B131FFCC3A40F9DFFE65FD6D9498" bold="true" box="[151,746,476,503]" fontSize="11" level="1" pageId="18" pageNumber="117" reason="1">
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<emphasis id="B941DA4FFFCC3A40F9DFFE65FD6D9498" bold="true" box="[151,746,476,503]" pageId="18" pageNumber="117">
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<taxonomicName id="4C357DDEFFCC3A40F9DFFE65FDC99498" authority="Barr and Shepard" authorityName="Barr and Shepard" authorityYear="2020" box="[151,590,476,503]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="117" phylum="Arthropoda" rank="species" species="lengua" status="sp. nov.">
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<emphasis id="B941DA4FFFCC3A40F9DFFE65FEE79499" bold="true" box="[151,352,476,503]" italics="true" pageId="18" pageNumber="117">Parygrus lengua</emphasis>
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||
Barr and Shepard
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="A2726734FFCC3A40FB12FE65FD6D9498" box="[602,746,476,503]" pageId="18" pageNumber="117" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
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||
</paragraph>
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||
<paragraph id="8B8A065DFFCC3A40F9DFFDB8FEC49775" blockId="18.[151,884,476,611]" box="[151,323,512,539]" pageId="18" pageNumber="117">(Figs 3, 16, 17)</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C32F55D6FFCC3A40F9DFFDF0FCF3970D" box="[151,884,584,611]" pageId="18" pageNumber="117" type="description">
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<paragraph id="8B8A065DFFCC3A40F9DFFDF0FCF3970D" blockId="18.[151,884,476,611]" box="[151,884,584,611]" pageId="18" pageNumber="117">
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http://zoobank.org/
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||
<uuid id="FF933C88FFCC3A40F838FDF0FCF3970D" box="[368,884,584,611]" pageId="18" pageNumber="117">8995529C-3873-46EC-B9F1-E030A1A07509</uuid>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32F55D6FFCC3A40F9DFFD29FA9796A5" pageId="18" pageNumber="117" type="materials_examined">
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<paragraph id="8B8A065DFFCC3A40F9DFFD29FB109631" blockId="18.[151,1437,656,2015]" pageId="18" pageNumber="117">
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||
<emphasis id="B941DA4FFFCC3A40F9DFFD29FEC197C5" bold="true" box="[151,326,656,683]" pageId="18" pageNumber="117">Type material.</emphasis>
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||
<materialsCitation id="3B5D0C00FFCC3A40F806FD29FF7C979D" ID-GBIF-Occurrence="2579576885" collectingDate="2006-12-02" collectionCode="EMEC" country="Paraguay" location="Puerto La Esperanza" pageId="18" pageNumber="117" specimenCount="2" stateProvince="Alto Paraguay" typeStatus="holotype">
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<emphasis id="B941DA4FFFCC3A40F806FD29FD2B97C5" bold="true" box="[334,684,656,683]" pageId="18" pageNumber="117">
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||
<typeStatus id="548EB8FFFFCC3A40F806FD29FE3C97C5" box="[334,443,657,683]" pageId="18" pageNumber="117" type="holotype">Holotype</typeStatus>
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||
male.
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||
<collectingCountry id="F32246CDFFCC3A40FB44FD28FD2097C5" box="[524,679,656,683]" name="Paraguay" pageId="18" pageNumber="117">PARAGUAY</collectingCountry>
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||
.
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||
</emphasis>
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||
“
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<collectingCountry id="F32246CDFFCC3A40FB88FD29FCDE97C5" box="[704,857,657,683]" name="Paraguay" pageId="18" pageNumber="117">PARAGUAY</collectingCountry>
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||
:
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||
<collectingRegion id="49F1C8BFFFCC3A40FA2BFD28FB8297C5" box="[867,1029,656,683]" country="Paraguay" name="Alto Paraguay" pageId="18" pageNumber="117">Alto Paraguay</collectingRegion>
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||
/
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||
<location id="8EEA5086FFCC3A40FD55FD29FA8097C5" LSID="urn:lsid:plazi:treatment:039CB74BFFCC3A46F9DFFE65FDC791A9:8EEA5086FFCC3A40FD55FD29FA8097C5" box="[1053,1287,657,683]" country="Paraguay" name="Puerto La Esperanza" pageId="18" pageNumber="117" stateProvince="Alto Paraguay">Puerto La Esperanza</location>
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||
/
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||
<date id="FF8B209DFFCC3A40FC57FD29FA1B97C4" box="[1311,1436,656,683]" pageId="18" pageNumber="117" value="2006-12-02">
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||
<collectingDate id="EFCFD975FFCC3A40FC57FD29FA1B97C4" box="[1311,1436,656,683]" pageId="18" pageNumber="117" value="2006-12-02">2 XII 2006</collectingDate>
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||
</date>
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||
/ Carlos Aguilar Julio //
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<typeStatus id="548EB8FFFFCC3A40F8EFFD0DFDBC97A1" box="[423,571,693,719]" pageId="18" pageNumber="117" type="holotype">HOLOTYPE</typeStatus>
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||
/
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<taxonomicName id="4C357DDEFFCC3A40FB19FD0DFC8197A0" authorityName="Barr and Shepard" authorityYear="2020" box="[593,774,692,719]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="117" phylum="Arthropoda" rank="species" species="lengua">Parygrus lengua</taxonomicName>
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||
/ Barr & Shepard [red label, handwritten] // EMEC49557” (
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||
<collectionCode id="ED249E98FFCC3A40F9D7FD61FF74979D" box="[159,243,729,755]" country="USA" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="18" pageNumber="117" type="University or college">EMEC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B5D0C00FFCC3A40F84DFD61FEE69630" ID-GBIF-Occurrence="2579576902" collectingDate="1996-01-20" collectingDateMax="2006-12-02" collectingDateMin="1996-01-20" collectionCode="EMEC, NHMUK, USNM" country="Paraguay" location="Carlos Aguilar Julio" municipality="Puerto La Esperanza" pageId="18" pageNumber="117" specimenCount="2" stateProvince="Alto Paraguay" typeStatus="paratype">
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||
<emphasis id="B941DA4FFFCC3A40F84DFD61FD06979D" bold="true" box="[261,641,728,755]" pageId="18" pageNumber="117">
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||
<typeStatus id="548EB8FFFFCC3A40F84DFD61FEFA979D" box="[261,381,729,755]" pageId="18" pageNumber="117" type="paratype">Paratypes</typeStatus>
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||
(10 M).
|
||
<collectingCountry id="F32246CDFFCC3A40F8A9FD60FDFD979D" box="[481,634,728,755]" name="Paraguay" pageId="18" pageNumber="117">PARAGUAY</collectingCountry>
|
||
.
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||
</emphasis>
|
||
<collectingCountry id="F32246CDFFCC3A40FBC0FD61FCA6979D" box="[648,801,729,755]" name="Paraguay" pageId="18" pageNumber="117">PARAGUAY</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F1C8BFFFCC3A40FA61FD60FC4E979D" box="[809,969,728,755]" country="Paraguay" name="Alto Paraguay" pageId="18" pageNumber="117">Alto Paraguay</collectingRegion>
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||
/
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||
<collectingMunicipality id="6BEE9C27FFCC3A40FA97FD61FB41979D" box="[991,1222,729,755]" pageId="18" pageNumber="117">Puerto La Esperanza</collectingMunicipality>
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||
/
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||
<date id="FF8B209DFFCC3A40FD93FD61FAD1979C" box="[1243,1366,728,755]" pageId="18" pageNumber="117" value="2006-12-02">
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||
<collectingDate id="EFCFD975FFCC3A40FD93FD61FAD1979C" box="[1243,1366,728,755]" pageId="18" pageNumber="117" value="2006-12-02">2 XII 2006</collectingDate>
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||
</date>
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||
/
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||
<location id="8EEA5086FFCC3A40FC23FD61FEC89678" LSID="urn:lsid:plazi:treatment:039CB74BFFCC3A46F9DFFE65FDC791A9:8EEA5086FFCC3A40FC23FD61FEC89678" country="Paraguay" municipality="Puerto La Esperanza" name="Carlos Aguilar Julio" pageId="18" pageNumber="117" stateProvince="Alto Paraguay">Carlos Aguilar Julio</location>
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(6 M,
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||
<collectionCode id="ED249E98FFCC3A40F8D3FD45FE6B9679" box="[411,492,765,791]" country="USA" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="18" pageNumber="117" type="University or college">EMEC</collectionCode>
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||
; 1 M, LSAM; 1 M,
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<collectionCode id="ED249E98FFCC3A40FB82FD45FCB09679" box="[714,823,765,791]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="117" type="Museum">NHMUK</collectionCode>
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||
; 1 M,
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||
<collectionCode id="ED249E98FFCC3A40FA36FD45FC539679" box="[894,980,765,791]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="117" type="Museum">USNM</collectionCode>
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||
);
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<collectingCountry id="F32246CDFFCC3A40FAAEFD45FBF89679" box="[998,1151,765,791]" name="Paraguay" pageId="18" pageNumber="117">PARAGUAY</collectingCountry>
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||
: Pdte. [Presidente] Hayes / Cruce los Pioneros /
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<date id="FF8B209DFFCC3A40F8D9FC99FE7A9654" box="[401,509,800,827]" pageId="18" pageNumber="117" value="1996-01-20">
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||
<collectingDate id="EFCFD975FFCC3A40F8D9FC99FE7A9654" box="[401,509,800,827]" pageId="18" pageNumber="117" value="1996-01-20">20 I 1996</collectingDate>
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||
</date>
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||
// Purchased from / Alfredo Ugarte (1 M,
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||
<collectionCode id="ED249E98FFCC3A40FA90FC99FBAC9655" box="[984,1067,801,827]" country="USA" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="18" pageNumber="117" type="University or college">EMEC</collectionCode>
|
||
). All
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||
<typeStatus id="548EB8FFFFCC3A40FD2FFC99FB559655" box="[1127,1234,801,827]" pageId="18" pageNumber="117" type="paratype">paratypes</typeStatus>
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||
also have the following final label
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||
</materialsCitation>
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||
:
|
||
<typeStatus id="548EB8FFFFCC3A40F824FCFDFE709631" box="[364,503,837,863]" pageId="18" pageNumber="117" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C357DDEFFCC3A40FB44FCFDFD409630" authorityName="Barr and Shepard" authorityYear="2020" box="[524,711,836,862]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="117" phylum="Arthropoda" rank="species" species="lengua">
|
||
<emphasis id="B941DA4FFFCC3A40FB44FCFDFD409630" box="[524,711,836,862]" italics="true" pageId="18" pageNumber="117">Parygrus lengua</emphasis>
|
||
</taxonomicName>
|
||
/ Barr & Shepard [yellow label, printed].
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCC3A40F98FFCD0FA9796A5" blockId="18.[151,1437,656,2015]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FFCD0FC8C96ED" bold="true" box="[199,779,872,899]" pageId="18" pageNumber="117">Additional specimens examined (18; 3 M, 15 F).</emphasis>
|
||
<materialsCitation id="3B5D0C00FFCC3A40FA59FCD0FD2E96C9" ID-GBIF-Occurrence="2579576889" collectingDate="2006-12" collectionCode="EMEC" collectorName="Zubaron Gaston" country="Argentina" location="Federacion" pageId="18" pageNumber="117" specimenCount="1" stateProvince="Entre Rios">
|
||
<emphasis id="B941DA4FFFCC3A40FA59FCD0FC4396ED" bold="true" box="[785,964,872,899]" pageId="18" pageNumber="117">
|
||
<collectingCountry id="F32246CDFFCC3A40FA59FCD0FC3996ED" box="[785,958,872,899]" name="Argentina" pageId="18" pageNumber="117">ARGENTINA</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
ARGENTINA:
|
||
<collectingRegion id="49F1C8BFFFCC3A40FD36FCD1FB7196EC" box="[1150,1270,872,899]" country="Argentina" name="Entre Rios" pageId="18" pageNumber="117">Entre Rios</collectingRegion>
|
||
/
|
||
<location id="8EEA5086FFCC3A40FC58FCD0FA0B96EC" LSID="urn:lsid:plazi:treatment:039CB74BFFCC3A46F9DFFE65FDC791A9:8EEA5086FFCC3A40FC58FCD0FA0B96EC" box="[1296,1420,872,898]" country="Argentina" name="Federacion" pageId="18" pageNumber="117" stateProvince="Entre Rios">Federacion</location>
|
||
/
|
||
<date id="FF8B209DFFCC3A40F9DFFC35FF7A96C8" box="[151,253,908,935]" pageId="18" pageNumber="117" value="2006-12">
|
||
<collectingDate id="EFCFD975FFCC3A40F9DFFC35FF7A96C8" box="[151,253,908,935]" pageId="18" pageNumber="117" value="2006-12">XII 2006</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="26C0638BFFCC3A40F85CFC34FE4B96C9" box="[276,460,908,935]" pageId="18" pageNumber="117">Zubaron Gaston</collectorName>
|
||
, leg. (3 M;
|
||
<collectionCode id="ED249E98FFCC3A40FB06FC35FD2696C9" box="[590,673,909,935]" country="USA" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="18" pageNumber="117" type="University or college">EMEC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B5D0C00FFCC3A40FBFCFC34FA8A96A5" ID-GBIF-Occurrence="2579576903" collectingDate="2006-12-02" collectionCode="NHMUK, USNM" country="Paraguay" location="Puerto La Esperanza" pageId="18" pageNumber="117" specimenCount="1" stateProvince="Asuncion">
|
||
<emphasis id="B941DA4FFFCC3A40FBFCFC34FCD396C9" bold="true" box="[692,852,908,935]" pageId="18" pageNumber="117">
|
||
<collectingCountry id="F32246CDFFCC3A40FBFCFC34FCCA96C9" box="[692,845,908,935]" name="Paraguay" pageId="18" pageNumber="117">PARAGUAY</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
<collectingRegion id="49F1C8BFFFCC3A40FA13FC35FC7396C9" box="[859,1012,909,935]" country="Paraguay" name="Asuncion" pageId="18" pageNumber="117">PARAGUAY</collectingRegion>
|
||
:
|
||
<collectingRegion id="49F1C8BFFFCC3A40FAB4FC34FB1A96C9" box="[1020,1181,908,935]" country="Paraguay" name="Alto Paraguay" pageId="18" pageNumber="117">Alto Paraguay</collectingRegion>
|
||
/
|
||
<location id="8EEA5086FFCC3A40FDFCFC35FA1B96C9" LSID="urn:lsid:plazi:treatment:039CB74BFFCC3A46F9DFFE65FDC791A9:8EEA5086FFCC3A40FDFCFC35FA1B96C9" box="[1204,1436,909,935]" country="Paraguay" name="Puerto La Esperanza" pageId="18" pageNumber="117" stateProvince="Asuncion">Puerto La Esperanza</location>
|
||
/
|
||
<date id="FF8B209DFFCC3A40F9EEFC09FEA696A4" box="[166,289,944,971]" pageId="18" pageNumber="117" value="2006-12-02">
|
||
<collectingDate id="EFCFD975FFCC3A40F9EEFC09FEA696A4" box="[166,289,944,971]" pageId="18" pageNumber="117" value="2006-12-02">2 XII 2006</collectingDate>
|
||
</date>
|
||
/ Carlos Aguilar Julio (10 F, EMEC; 2 F, FSCA; 1 F, LSAM; 1 F,
|
||
<collectionCode id="ED249E98FFCC3A40FD4EFC09FBF496A5" box="[1030,1139,945,971]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="117" type="Museum">NHMUK</collectionCode>
|
||
; 1 F,
|
||
<collectionCode id="ED249E98FFCC3A40FDE7FC09FA8296A5" box="[1199,1285,945,971]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="117" type="Museum">USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFCC3A40F98FFC6CFCC49111" pageId="18" pageNumber="117" type="diagnosis">
|
||
<paragraph id="8B8A065DFFCC3A40F98FFC6CFCC49111" blockId="18.[151,1437,656,2015]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FFC6CFEC59681" bold="true" box="[199,322,980,1007]" pageId="18" pageNumber="117">Diagnosis.</emphasis>
|
||
The male genitalia of
|
||
<taxonomicName id="4C357DDEFFCC3A40FB74FC6DFD709680" authority="Barr & Shepard, 2020" authorityName="Barr & Shepard" authorityYear="2020" box="[572,759,980,1006]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="117" phylum="Arthropoda" rank="species" species="lengua" status="sp. nov.">
|
||
<emphasis id="B941DA4FFFCC3A40FB74FC6DFD709680" box="[572,759,980,1006]" italics="true" pageId="18" pageNumber="117">Parygrus lengua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B941DA4FFFCC3A40FBB6FC6DFCBB9681" bold="true" box="[766,828,981,1007]" pageId="18" pageNumber="117">
|
||
<taxonomicNameLabel id="A2726734FFCC3A40FBB6FC6DFCBB9681" box="[766,828,981,1007]" pageId="18" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are distinctive (Fig. 17); none of the other species (for which the genitalia are known) have parameres both with inner ridges and without teeth. An external character not seen in other species is that the
|
||
<taxonomicName id="4C357DDEFFCC3A40FB40FBA4FDF29158" box="[520,629,1052,1078]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="18" pageNumber="123" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
|
||
is weakly depressed at the anterior half and convex near the posterior angle. The females were assigned to this species due to their association with the males, their similar scutellar morphology, and body size; they are not designated as
|
||
<typeStatus id="548EB8FFFFCC3A40FB98FBDDFCB89111" box="[720,831,1125,1151]" pageId="18" pageNumber="117" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFCC3A46F98FFB30FF659135" lastPageId="20" lastPageNumber="119" pageId="18" pageNumber="117" type="description">
|
||
<paragraph id="8B8A065DFFCC3A40F98FFB30FD0B9185" blockId="18.[151,1437,656,2015]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FFB30FD9091CD" bold="true" box="[199,535,1160,1187]" pageId="18" pageNumber="117">
|
||
Description.
|
||
<typeStatus id="548EB8FFFFCC3A40F829FB31FE4991CD" box="[353,462,1161,1187]" pageId="18" pageNumber="117" type="holotype">Holotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
Cuticle brown, legs, antennae and mouthparts lighter; elongate, subcylindrical; length
|
||
<quantity id="4CCDABB8FFCC3A40F9AEFB14FECC91A9" box="[230,331,1196,1223]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" pageId="18" pageNumber="117" unit="mm" value="4.35">4.35 mm</quantity>
|
||
(pronotum + elytra), width
|
||
<quantity id="4CCDABB8FFCC3A40FBC2FB14FD7491A9" box="[650,755,1196,1223]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="117" unit="mm" value="1.6">1.60 mm</quantity>
|
||
; body covered with erect and semi-erect, long, golden setae and much shorter, recumbent setae (
|
||
<figureCitation id="130E1AD8FFCC3A40FB63FB68FDF99185" box="[555,638,1232,1259]" captionStart="FIGURE 16" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[261,1325,181,1920]" captionTargetId="figure@19.[261,1327,181,1920]" captionTargetPageId="19" captionText="FIGURE 16. Parygrus lengua n. sp., male holotype; 4.35 mm long, 1.65 mm wide: a) dorsal habitus; b) lateral habitus. FIGURE 17. Parygrus lengua n. sp., male genitalia: a) dorsal view; b) lateral view; c) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3724273" httpUri="https://zenodo.org/record/3724273/files/figure.png" pageId="18" pageNumber="117">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCC3A40F98FFB4CFC439364" blockId="18.[151,1437,656,2015]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FFB4CFE839060" box="[199,260,1268,1294]" italics="true" pageId="18" pageNumber="117">Head</emphasis>
|
||
dark brown; punctuation fine, dense, punctures separated by at least 1x puncture diameter; setae moderately long; vertex with moderately deep median depression; frons raised and protruding slightly forward between antennal bases, anterior margin arcuate, distance between antennal bases less than length of antennomere 1. Antenna with antennomeres 1 and 2 light brown, setose, antennomeres 3–11 yellow, densely setose; antennomere 1 nearly as long as antennomere 2; antennomere 2 covering antennomeres 3–5. Clypeus densely setose, coarsely punctate, barely emarginate. Maxillary palpus with terminal palpomere elongate, subcylindrical, weakly curved. Labrum emarginate, densely setose, border fringed with long setae. Labial palpus with terminal palpomere subrectangular, slightly flattened; length about 2/3 that of terminal maxillary palpomere.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCC3A40F98FF9ADFB51938C" blockId="18.[151,1437,656,2015]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FF9ADFEB19340" box="[199,310,1557,1582]" italics="true" pageId="18" pageNumber="117">Pronotum</emphasis>
|
||
dark brown; weakly convex, disc weakly flattened at center; dark brown; length
|
||
<quantity id="4CCDABB8FFCC3A40FDFEF9ADFA9C9341" box="[1206,1307,1557,1583]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="117" unit="mm" value="0.9">0.90 mm</quantity>
|
||
, width
|
||
<quantity id="4CCDABB8FFCC3A40FC23F9ACFF42933D" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="18" pageNumber="117" unit="mm" value="1.25">1.25 mm</quantity>
|
||
, widest at basal 1/3. Anterior margin straight except at strongly acute, depressed, anterolateral angles; lateral margins weakly arcuate, barely sinuate at lateral angles, narrowly margined, fringed with erect setae; posterolateral angles acute, explanate; posterior border trisinuate. Disc punctuation slightly coarser than on head; setae moderately long.
|
||
<taxonomicName id="4C357DDEFFCC3A40F99EF91CFEC393D0" box="[214,324,1700,1726]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="18" pageNumber="117" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFCC3A40F99EF91CFEC393D0" box="[214,324,1700,1726]" italics="true" pageId="18" pageNumber="117">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
dark brown; subcordate, wider than long; anterior margin weakly arcuate between two anterolateral notches; disc weakly depressed at anterior 1/2, convex near posterior angle; punctation very fine.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCC3A40F98FF954FEF992F9" blockId="18.[151,1437,656,2015]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FF954FE9D9268" box="[199,282,1772,1798]" italics="true" pageId="18" pageNumber="117">Elytron</emphasis>
|
||
red-brown; length
|
||
<quantity id="4CCDABB8FFCC3A40F8BAF954FDDF9269" box="[498,600,1772,1799]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" pageId="18" pageNumber="117" unit="mm" value="3.45">3.45 mm</quantity>
|
||
, width
|
||
<quantity id="4CCDABB8FFCC3A40FBE2F955FC969269" box="[682,785,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="117" unit="mm" value="0.8">0.80 mm</quantity>
|
||
; convex; narrowest at basal 1/3, widest at apical 1/3. Base depressed near margin; humeral angle rounded; lateral margin widened at basal 1/3, narrowly margined; apex acute. Disc strongly punctate and striate with deep, closely spaced punctures distinct from near base to apex, intervals weakly convex; setae slightly shorter than on pronotum, semi-erect and recumbent, uniformly distributed.
|
||
<emphasis id="B941DA4FFFCC3A40FC2CF8E0FF5092F8" italics="true" pageId="18" pageNumber="117">Hind wing:</emphasis>
|
||
macropterous.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCC3A46F98FF819FEAF9409" blockId="18.[151,1437,656,2015]" lastBlockId="20.[151,1437,153,1223]" lastPageId="20" lastPageNumber="119" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FF819FE8392D4" box="[199,260,1953,1978]" italics="true" pageId="18" pageNumber="117">Legs.</emphasis>
|
||
Profemur red-brown, covered with fine, evenly spaced punctures and long, recumbent golden setae; protibia yellow-brown, mostly bare and shiny, dorsal surface with sparse, very long setae, ventral surface with dense row of short, stiff setae, weakly arcuate with narrow spines at ventral apex; protarsus yellow-brown, shiny, dorsal surface with sparse, very long setae, ventral surface with row of shorter, semi-erect setae, protarsomeres 4 and 5 each with a few much-longer dorsal setae. Mesofemur similar to profemur, mesotibia yellow-brown, ventral setae much longer than those of protibia, weakly arcuate with spinose ventral apex; mesotarsus similar to protarsus except ventral setae much longer. Metafemur and metatarsus similar to those of mesoleg, metatibia straight with spines at ventral apex.
|
||
</paragraph>
|
||
<caption id="DF4A56D5FFCD3A41F9DFF821FB1592B8" ID-DOI="http://doi.org/10.5281/zenodo.3724273" ID-Zenodo-Dep="3724273" httpUri="https://zenodo.org/record/3724273/files/figure.png" pageId="19" pageNumber="118" startId="19.[151,250,1945,1970]" targetBox="[261,1325,181,1920]" targetPageId="19">
|
||
<paragraph id="8B8A065DFFCD3A41F9DFF821FB1592B8" blockId="19.[151,1369,1945,2006]" pageId="19" pageNumber="118">
|
||
<emphasis id="B941DA4FFFCD3A41F9DFF821FEA792DC" bold="true" box="[151,288,1945,1970]" pageId="19" pageNumber="118">FIGURE 16.</emphasis>
|
||
<taxonomicName id="4C357DDEFFCD3A41F86EF822FE5792DC" authority="Barr & Shepard, 2020" authorityName="Barr & Shepard" authorityYear="2020" box="[294,464,1946,1970]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="118" phylum="Arthropoda" rank="species" species="lengua" status="sp. nov.">
|
||
<emphasis id="B941DA4FFFCD3A41F86EF822FE5792DC" box="[294,464,1946,1970]" italics="true" pageId="19" pageNumber="118">Parygrus lengua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B941DA4FFFCD3A41F89EF822FD8892DC" bold="true" box="[470,527,1946,1970]" pageId="19" pageNumber="118">
|
||
<taxonomicNameLabel id="A2726734FFCD3A41F89EF822FD8892DC" box="[470,527,1946,1970]" pageId="19" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype; 4.35 mm long, 1.65 mm wide: a) dorsal habitus; b) lateral habitus.
|
||
<emphasis id="B941DA4FFFCD3A41F9DFF805FEA792B8" bold="true" box="[151,288,1981,2006]" pageId="19" pageNumber="118">FIGURE 17.</emphasis>
|
||
<taxonomicName id="4C357DDEFFCD3A41F86EF806FE5792B8" authority="Barr & Shepard, 2020" authorityName="Barr & Shepard" authorityYear="2020" box="[294,464,1982,2006]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="118" phylum="Arthropoda" rank="species" species="lengua" status="sp. nov.">
|
||
<emphasis id="B941DA4FFFCD3A41F86EF806FE5792B8" box="[294,464,1982,2006]" italics="true" pageId="19" pageNumber="118">Parygrus lengua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B941DA4FFFCD3A41F89EF806FD8892BB" bold="true" box="[470,527,1982,2006]" pageId="19" pageNumber="118">
|
||
<taxonomicNameLabel id="A2726734FFCD3A41F89EF806FD8892BB" box="[470,527,1982,2006]" pageId="19" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male genitalia: a) dorsal view; b) lateral view; c) ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8A065DFFCA3A46F98FFECAFCE897E9" blockId="20.[151,1437,153,1223]" pageId="20" pageNumber="119">
|
||
<emphasis id="B941DA4FFFCA3A46F98FFECAFE8894E5" box="[199,271,370,395]" italics="true" pageId="20" pageNumber="119">Venter</emphasis>
|
||
red-brown; heavily setose; finely to coarsely punctate. Prosternum with anterior border narrowly margined; prosternal process wide between procoxae, swollen over coxal bases, lateral margins weakly arcuate; process with median longitudinal carina terminating in a low, elongate protuberance near apex. Metaventrite with anterior process margined and depressed between mesocoxae; posterior disc depressed at junction of sulcate metakatepisternal suture and metathoracic discrimen. Abdomen with some setae longer than on rest of body except legs; ventrite 1 with triangular intercoxal process weakly depressed, margined laterally; ventrites 2–4 of equal length, finely and evenly punctate at center of disc, coarser laterally, aligned with faint, transverse strigae; ventrite 5 longest, disc coarsely punctate, covered with long setae prominent at margins.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCA3A46F98FFD29FBEB9631" blockId="20.[151,1437,153,1223]" pageId="20" pageNumber="119">
|
||
<emphasis id="B941DA4FFFCA3A46F98FFD29FEBE97C5" box="[199,313,657,683]" italics="true" pageId="20" pageNumber="119">Genitalia.</emphasis>
|
||
Aedeagus moderately elongate (Fig. 17). Phallobase same length as parameres and same width as paramere bases together. Parameres each blade-like, inner surface concave; in dorsal view narrow, lateral margins nearly straight with a slight inward curve at midlength; in dorsolateral view sclerotized inner dorsal margin with short parallel ridges nearly perpendicular to margin; in lateral view each paramere wide, dorsal and ventral surfaces broadly arcuate, paramere tip acuminate and angled ventrally. Penis nearly as long as parameres, bottle-shaped, base constricted, basal 1/3–1/2 bulbous, apical 1/2 evenly tapered to narrowly rounded apex.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCA3A46F98FFCD0FF659135" blockId="20.[151,1437,153,1223]" pageId="20" pageNumber="119">
|
||
<emphasis id="B941DA4FFFCA3A46F98FFCD0FE5C96ED" bold="true" box="[199,475,872,899]" pageId="20" pageNumber="119">Intraspecific variation.</emphasis>
|
||
The males from
|
||
<collectingCountry id="F32246CDFFCA3A46FBE8FCD1FC8F96ED" box="[672,776,873,899]" name="Paraguay" pageId="20" pageNumber="119">Paraguay</collectingCountry>
|
||
vary from
|
||
<quantity id="4CCDABB8FFCA3A46FAC1FCD0FBA996ED" box="[905,1070,872,899]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.45" metricValueMin="3.85" pageId="20" pageNumber="119" unit="mm" value="4.15" valueMax="4.45" valueMin="3.85">3.85–4.45 mm</quantity>
|
||
long (n=5); females from
|
||
<quantity id="4CCDABB8FFCA3A46FC15FCD0FF7B96C9" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="20" pageNumber="119" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.50– 4.70 mm</quantity>
|
||
long (n=4). The three male specimens from
|
||
<collectingCountry id="F32246CDFFCA3A46FBBBFC35FCE596C9" box="[755,866,909,935]" name="Argentina" pageId="20" pageNumber="119">Argentina</collectingCountry>
|
||
are much larger (
|
||
<quantity id="4CCDABB8FFCA3A46FD62FC34FB4896C9" box="[1066,1231,908,935]" metricMagnitude="-3" metricUnit="m" metricValue="5.075" metricValueMax="5.2" metricValueMin="4.95" pageId="20" pageNumber="119" unit="mm" value="5.075" valueMax="5.2" valueMin="4.95">4.95–5.20 mm</quantity>
|
||
long), as are their genitalia, but otherwise differences between the groups are minor. Nevertheless, they have not been designated as
|
||
<typeStatus id="548EB8FFFFCA3A46F9DFFC6DFE819681" box="[151,262,981,1007]" pageId="20" pageNumber="119" type="paratype">paratypes</typeStatus>
|
||
. The morphology of the prosternal process is somewhat variable among all of the males examined. Some have the process swollen over coxal bases and not margined; others are not swollen, but margined and have the longitudinal carina bordered laterally by shallow sulci. Among males, the degree to which the protibiae are curved varies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFCA3A46F98FFBDCFBB39111" box="[199,1076,1124,1151]" pageId="20" pageNumber="119" type="distribution">
|
||
<paragraph id="8B8A065DFFCA3A46F98FFBDCFBB39111" blockId="20.[151,1437,153,1223]" box="[199,1076,1124,1151]" pageId="20" pageNumber="119">
|
||
<emphasis id="B941DA4FFFCA3A46F98FFBDCFEE69111" bold="true" box="[199,353,1124,1151]" pageId="20" pageNumber="119">Distribution.</emphasis>
|
||
Known from three localities in
|
||
<collectingCountry id="F32246CDFFCA3A46FB8DFBDDFCB39111" box="[709,820,1125,1151]" name="Argentina" pageId="20" pageNumber="119">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32246CDFFCA3A46FA22FBDDFC559111" box="[874,978,1125,1151]" name="Paraguay" pageId="20" pageNumber="119">Paraguay</collectingCountry>
|
||
(Fig. 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFCA3A46F98FFB31FDC791A9" pageId="20" pageNumber="119" type="etymology">
|
||
<paragraph id="8B8A065DFFCA3A46F98FFB31FDC791A9" blockId="20.[151,1437,153,1223]" pageId="20" pageNumber="119">
|
||
<emphasis id="B941DA4FFFCA3A46F98FFB31FEC991CD" bold="true" box="[199,334,1161,1187]" pageId="20" pageNumber="119">Etymology.</emphasis>
|
||
This new species is named in honor of the
|
||
<emphasis id="B941DA4FFFCA3A46FA76FB32FC1591CD" box="[830,914,1162,1187]" italics="true" pageId="20" pageNumber="119">Lengua</emphasis>
|
||
, one of the indigenous peoples who inhabited the Gran Chaco of northern
|
||
<collectingCountry id="F32246CDFFCA3A46F89AFB15FDBB91A9" box="[466,572,1197,1223]" name="Paraguay" pageId="20" pageNumber="119">Paraguay</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |