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<document id="F98F02866B44BB88F97E3D98B48B6ED7" ID-DOI="10.1515/aemnp-2017-0097" ID-ISSN="0374-1036" ID-Zenodo-Dep="5318416" ID-ZooBank="638421F2-985E-43CC-8438-E871F96B1DB8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630129638815" checkinUser="marcus" docAuthor="Magoga, Giulia, Montagna, Matteo, Marziali, Laura, Rossaro, Bruno &amp; Disaa" docDate="2017" docId="039435190B55C86BFEA8FACBFE8F5B9E" docLanguage="en" docName="ActaEntMusNatPra.2017.57.2.723-749.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57 (2)" docSource="http://dx.doi.org/10.1515/aemnp-2017-0097" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Orthocladius (Orthocladius) rhyacobius Kieffer 1911" docType="treatment" docVersion="4" lastPageNumber="741" masterDocId="FFAD4D610B45C879FFFFFFB9FD53590D" masterDocTitle="Revision of type and non-type material assigned to the genus Orthocladius by Goetghebuer (1940 - 1950), deposited in the Royal Belgian Institute of Natural Sciences (Diptera: Chironomidae)" masterLastPageNumber="749" masterPageNumber="723" pageNumber="739" updateTime="1699162318611" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F3B78AD01FE88438B4EE060131B29EF5">Revision of type and non-type material assigned to the genus Orthocladius by Goetghebuer (1940 - 1950), deposited in the Royal Belgian Institute of Natural Sciences (Diptera: Chironomidae)</mods:title>
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<mods:namePart id="B3E7C6C5EE81733F656A0A9B57A6762A">Magoga, Giulia</mods:namePart>
<mods:affiliation id="869151EE0EDCC730F4F8A557E77FB0BE">) &amp;) &amp;) Università degli Studi di Milano, Dipartimento di Scienze Agrarie e Ambientali (DISAA), via Celoria 2, I 20133, Milano, Italia; e-mails: giulia. magoga @ unimi. it; matteo. montagna @ unimi. it</mods:affiliation>
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<mods:namePart id="173FF93FD4C7ACB3D8714A76100F52B7">Montagna, Matteo</mods:namePart>
<mods:affiliation id="2ACFE1EDB9F9C793788884B554CC9719">) &amp;) &amp;) Università degli Studi di Milano, Dipartimento di Scienze Agrarie e Ambientali (DISAA), via Celoria 2, I 20133, Milano, Italia; e-mails: giulia. magoga @ unimi. it; matteo. montagna @ unimi. it</mods:affiliation>
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<mods:namePart id="60085A6F23B82BF7D2B2F590A1F78A1B">Marziali, Laura</mods:namePart>
<mods:affiliation id="CC05DDF2F2AC4E315988C40D957E2128">) &amp;) Water Research Institute, National Research Council (IRSA-CNR), Via del Mulino 19, I 20861 Brugherio (MB), Italy; e-mail: marziali @ irsa. cnr. it</mods:affiliation>
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<mods:namePart id="AB400857856F3154001C21340BA65D8D">Rossaro, Bruno</mods:namePart>
<mods:affiliation id="DEBE7BB5D539E613EF1E236FBA217287">) &amp;) Università degli Studi di Milano, Dipartimento di Scienze per gli Alimenti, la Nutrizione e l’Ambiente (DeFENS), Via Celoria 2, I 20133 Milan, Italy; e-mail: bruno. rossaro @ unimi. it</mods:affiliation>
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<mods:title id="BF15900CDBC03F8C9A508B1FF83431E0">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="D858BAF3F9F1F0DE521BE59E5B5F9C8E">2017</mods:date>
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<treatment id="039435190B55C86BFEA8FACBFE8F5B9E" ID-DOI="http://doi.org/10.5281/zenodo.5335068" ID-Zenodo-Dep="5335068" LSID="urn:lsid:plazi:treatment:039435190B55C86BFEA8FACBFE8F5B9E" httpUri="http://treatment.plazi.org/id/039435190B55C86BFEA8FACBFE8F5B9E" lastPageId="18" lastPageNumber="741" pageId="16" pageNumber="739">
<subSubSection id="C327D7840B55C869FEA8FACBFEF55C87" box="[343,934,1394,1418]" pageId="16" pageNumber="739" type="nomenclature">
<paragraph id="8B82840F0B55C869FEA8FACBFEF55C87" blockId="16.[343,934,1394,1418]" box="[343,934,1394,1418]" pageId="16" pageNumber="739">
<heading id="D0CA33630B55C869FEA8FACBFEF55C87" bold="true" box="[343,934,1394,1418]" centered="true" fontSize="8" level="2" pageId="16" pageNumber="739" reason="3">
<taxonomicName id="4C3DFF8C0B55C869FEA8FACBFEF55C87" ID-CoL="89MBN" authority="Kieffer, 1911" authorityName="Kieffer" authorityYear="1911" box="[343,934,1394,1418]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="739" phylum="Arthropoda" rank="species" species="rhyacobius" subGenus="Orthocladius">
<emphasis id="B949581D0B55C869FEA8FACBFEF55C87" bold="true" box="[343,934,1394,1418]" pageId="16" pageNumber="739">
<emphasis id="B949581D0B55C869FEA8FACBFCB45C87" bold="true" box="[343,487,1394,1418]" italics="true" pageId="16" pageNumber="739">Orthocladius</emphasis>
(
<emphasis id="B949581D0B55C869FE08FACBFFD45C87" bold="true" box="[503,647,1394,1418]" italics="true" pageId="16" pageNumber="739">Orthocladius</emphasis>
)
<emphasis id="B949581D0B55C869FD69FACBFE5C5C87" bold="true" box="[662,783,1394,1418]" italics="true" pageId="16" pageNumber="739">rhyacobius</emphasis>
<bibRefCitation id="EFACF9FE0B55C869FCE9FACAFEF55C87" author="KIEFFER J. J." box="[790,934,1395,1418]" pageId="16" pageNumber="739" pagination="181 - 187" refId="ref9890" refString="KIEFFER J. J. 1911: Nouveaux Tendipedides du groupe Orthocladius [Dipt.]. (1 re note). Bulletin de la Societe Entomologique de France 1911: 181 - 187." type="journal article" year="1911">Kieffer, 1911</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C327D7840B55C869FDB7FA28FFE65CA9" box="[584,693,1425,1444]" pageId="16" pageNumber="739" type="description">
<paragraph id="8B82840F0B55C869FDB7FA28FFE65CA9" blockId="16.[151,1127,1425,1573]" box="[584,693,1425,1444]" pageId="16" pageNumber="739">
(
<figureCitation id="1306988A0B55C869FDB1FA28FFFC5CA9" box="[590,687,1425,1444]" captionStart="Figs 53‒58" captionStartId="17.[115,152,1280,1299]" captionTargetBox="[113,1088,220,1265]" captionTargetId="figure-115@17.[113,1088,220,1265]" captionTargetPageId="17" captionText="Figs 53‒58. Orthocladius (Orthocladius) rhyacobius Kieffer, 1911. 53 ‒ antenna, 54 ‒ thorax, 55 ‒ palps, 56 ‒ hypopygium, 57 ‒ inferior volsella, 58 ‒ virga and superior volsella." figureDoi="http://doi.org/10.5281/zenodo.5318442" httpUri="https://zenodo.org/record/5318442/files/figure.png" pageId="16" pageNumber="739">Figs 53‒58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C327D7840B55C869FF68FA0AFFCB5CCB" box="[151,664,1459,1478]" pageId="16" pageNumber="739" type="reference_group">
<paragraph id="8B82840F0B55C869FF68FA0AFFCB5CCB" blockId="16.[151,1127,1425,1573]" box="[151,664,1459,1478]" pageId="16" pageNumber="739">
<treatmentCitationGroup id="AB2DA3210B55C869FF68FA0AFFCB5CCB" box="[151,664,1459,1478]" pageId="16" pageNumber="739">
<treatmentCitation id="0A9CA21E0B55C869FF68FA0AFFC05CCB" author="KIEFFER J. J." box="[151,659,1459,1478]" page="181" pageId="16" pageNumber="739" year="1911">
<taxonomicName id="4C3DFF8C0B55C869FF68FA0AFFC05CCB" ID-CoL="89MBN" authority="Kieffer, 1911: 181" authorityName="Kieffer" authorityPageNumber="181" authorityYear="1911" box="[151,659,1459,1478]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="739" phylum="Arthropoda" rank="species" species="rhyacobius" subGenus="Orthocladius">
<emphasis id="B949581D0B55C869FF68FA0AFC5B5CCB" box="[151,264,1459,1478]" italics="true" pageId="16" pageNumber="739">Orthocladius</emphasis>
(
<emphasis id="B949581D0B55C869FEEBFA0AFCD65CCB" box="[276,389,1459,1478]" italics="true" pageId="16" pageNumber="739">Orthocladius</emphasis>
)
<emphasis id="B949581D0B55C869FE6EFA0AFCBC5CCB" box="[401,495,1459,1478]" italics="true" pageId="16" pageNumber="739">rhyacobius</emphasis>
<bibRefCitation id="EFACF9FE0B55C869FE0AFA0AFFC05CCB" author="KIEFFER J. J." box="[501,659,1459,1478]" pageId="16" pageNumber="739" pagination="181 - 187" refId="ref9890" refString="KIEFFER J. J. 1911: Nouveaux Tendipedides du groupe Orthocladius [Dipt.]. (1 re note). Bulletin de la Societe Entomologique de France 1911: 181 - 187." type="journal article" year="1911">Kieffer, 1911: 181</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C327D7840B55C869FF68FA64FFC35F28" pageId="16" pageNumber="739" type="materials_examined">
<paragraph id="8B82840F0B55C869FF68FA64FFC35F28" blockId="16.[151,1127,1425,1573]" pageId="16" pageNumber="739">
<emphasis id="B949581D0B55C869FF68FA64FC165CFD" bold="true" box="[151,325,1501,1520]" pageId="16" pageNumber="739">Material examined</emphasis>
. 1 ♁ pinned labelled: ‘
<taxonomicName id="4C3DFF8C0B55C869FDF0FA64FE015CFD" authority="Edw. Brno 1937" authorityName="Edw. Brno" authorityYear="1937" box="[527,850,1501,1520]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="739" phylum="Arthropoda" rank="species" species="oblidens">
<emphasis id="B949581D0B55C869FDF0FA64FF215CFD" box="[527,626,1501,1520]" italics="true" pageId="16" pageNumber="739">O. oblidens</emphasis>
Edw. Brno 1937 (Zavrel)
</taxonomicName>
//
<collectionCode id="ED2C1CCA0B55C869FC94FA64FE2F5CFD" box="[875,892,1501,1520]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="739">R</collectionCode>
.I.SC.N.B. 18.073 / Coll. et det. M. Goetghebuer’; 1 ♁ in isinglass labelled: ‘
<taxonomicName id="4C3DFF8C0B55C869FDC6FA41FEFA5F06" authority="K. Brno, Zavrel" authorityName="K. Brno, Zavrel" box="[569,937,1528,1547]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="739" phylum="Arthropoda" rank="species" species="rhyacophilus">
<emphasis id="B949581D0B55C869FDC6FA41FE4E5F06" box="[569,797,1528,1547]" italics="true" pageId="16" pageNumber="739">Orthocladius rhyacophilus</emphasis>
K. Brno, Zavrel
</taxonomicName>
//
<collectionCode id="ED2C1CCA0B55C869FC40FA41FE835F06" box="[959,976,1528,1547]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="739">R</collectionCode>
.I.SC.N.B. 18.073 / Coll. et det. M. Goetghebuer’, all now mounted on slides.
</paragraph>
</subSubSection>
<caption id="DF42D4870B54C868FF8CFAB9FFF65C23" ID-DOI="http://doi.org/10.5281/zenodo.5318442" ID-Zenodo-Dep="5318442" httpUri="https://zenodo.org/record/5318442/files/figure.png" pageId="17" pageNumber="740" startId="17.[115,152,1280,1299]" targetBox="[113,1088,220,1265]" targetPageId="17">
<paragraph id="8B82840F0B54C868FF8CFAB9FFF65C23" blockId="17.[115,1088,1280,1326]" pageId="17" pageNumber="740">
Figs 53‒58.
<taxonomicName id="4C3DFF8C0B54C868FF22FAB9FFFE5C1E" authority="Kieffer, 1911" authorityName="Kieffer" authorityYear="1911" box="[221,685,1280,1299]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="740" phylum="Arthropoda" rank="species" species="rhyacobius" subGenus="Orthocladius">
<emphasis id="B949581D0B54C868FF22FAB9FC1D5C1E" box="[221,334,1280,1299]" italics="true" pageId="17" pageNumber="740">Orthocladius</emphasis>
(
<emphasis id="B949581D0B54C868FEA5FAB9FC985C1E" box="[346,459,1280,1299]" italics="true" pageId="17" pageNumber="740">Orthocladius</emphasis>
)
<emphasis id="B949581D0B54C868FE28FAB9FF665C1E" box="[471,565,1280,1299]" italics="true" pageId="17" pageNumber="740">rhyacobius</emphasis>
<bibRefCitation id="EFACF9FE0B54C868FDC5FAB9FFFE5C1E" author="KIEFFER J. J." box="[570,685,1280,1299]" pageId="17" pageNumber="740" pagination="181 - 187" refId="ref9890" refString="KIEFFER J. J. 1911: Nouveaux Tendipedides du groupe Orthocladius [Dipt.]. (1 re note). Bulletin de la Societe Entomologique de France 1911: 181 - 187." type="journal article" year="1911">Kieffer, 1911</bibRefCitation>
</taxonomicName>
. 53 ‒ antenna, 54 ‒ thorax, 55 ‒ palps, 56 ‒ hypopygium, 57 ‒ inferior volsella, 58 ‒ virga and superior volsella.
</paragraph>
</caption>
<subSubSection id="C327D7840B54C86BFF8EFAE0FE8F5B9E" lastPageId="18" lastPageNumber="741" pageId="17" pageNumber="740" type="discussion">
<paragraph id="8B82840F0B54C868FF8EFAE0F9125CBD" blockId="17.[113,1089,1369,1552]" pageId="17" pageNumber="740">
<emphasis id="B949581D0B54C868FF8EFAE0FDA05C7D" bold="true" box="[113,243,1369,1392]" pageId="17" pageNumber="740">Comments.</emphasis>
The
<specimenCount id="9D3B4F860B54C868FED8FAE0FC975C7D" box="[295,452,1369,1392]" pageId="17" pageNumber="740" type="generic">two specimens</specimenCount>
apparently come from the same locality and are conspecific. The dorsal lobes of the inferior volsellae and the absence of virga fit the description of
<taxonomicName id="4C3DFF8C0B54C868FBD8FAC0FC2E5CBD" authority="Kieffer, 1911" authorityName="Kieffer" authorityYear="1911" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="740" phylum="Arthropoda" rank="species" species="rhyacobius">
<emphasis id="B949581D0B54C868FBD8FAC0FDB45CBD" italics="true" pageId="17" pageNumber="740">O. rhyacobius</emphasis>
<bibRefCitation id="EFACF9FE0B54C868FF11FA20FC2E5CBD" author="KIEFFER J. J." box="[238,381,1433,1456]" pageId="17" pageNumber="740" pagination="181 - 187" refId="ref9890" refString="KIEFFER J. J. 1911: Nouveaux Tendipedides du groupe Orthocladius [Dipt.]. (1 re note). Bulletin de la Societe Entomologique de France 1911: 181 - 187." type="journal article" year="1911">Kieffer, 1911</bibRefCitation>
</taxonomicName>
, not with that of
<taxonomicName id="4C3DFF8C0B54C868FDCEFA20F96E5CBD" authority="(Walker, 1856) (ROSSARO et al. 2003)" authorityName="ROSSARO et al." authorityYear="2003" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[561,1085,1433,1456]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="740" phylum="Arthropoda" rank="species" species="oblidens">
<emphasis id="B949581D0B54C868FDCEFA20FFF85CBD" box="[561,683,1433,1456]" italics="true" pageId="17" pageNumber="740">O. oblidens</emphasis>
(Walker, 1856) (
<bibRefCitation id="EFACF9FE0B54C868FCA0FA20F9655CBD" author="ROSSARO B. &amp; LENCIONI V. &amp; CASALEGNO C." box="[863,1078,1433,1456]" pageId="17" pageNumber="740" pagination="213 - 241" refId="ref10485" refString="ROSSARO B., LENCIONI V. &amp; CASALEGNO C. 2003: Revision of West Palaearctic species of Orthocladius s. str. van der Wulp, 1874 (Diptera: Chironomidae: Orthocladiinae), with a new key to species. Studi Trentini di Scienze Naturali, Acta Biologica 79: 213 - 241." type="journal article" year="2003">ROSSARO et al. 2003</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B82840F0B54C86BFF6FFA00FF0158DE" blockId="17.[113,1089,1369,1552]" lastBlockId="18.[151,1127,220,659]" lastPageId="18" lastPageNumber="741" pageId="17" pageNumber="740">
<bibRefCitation id="EFACF9FE0B54C868FF6FFA00FCC55CDD" author="KIEFFER J. J." box="[144,406,1465,1488]" pageId="17" pageNumber="740" pagination="181 - 187" refId="ref9890" refString="KIEFFER J. J. 1911: Nouveaux Tendipedides du groupe Orthocladius [Dipt.]. (1 re note). Bulletin de la Societe Entomologique de France 1911: 181 - 187." type="journal article" year="1911">KIEFFER (1911: 181‒182)</bibRefCitation>
separated
<taxonomicName id="4C3DFF8C0B54C868FDFBFA00FFCA5CDD" authorityName="Kieffer" authorityYear="1911" box="[516,665,1465,1488]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="740" phylum="Arthropoda" rank="species" species="rhyacobius">
<emphasis id="B949581D0B54C868FDFBFA00FFCA5CDD" box="[516,665,1465,1488]" italics="true" pageId="17" pageNumber="740">O. rhyacobius</emphasis>
</taxonomicName>
from
<taxonomicName id="4C3DFF8C0B54C868FD29FA00FE2C5CDD" box="[726,895,1465,1488]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="740" phylum="Arthropoda" rank="species" species="rhyacophilus">
<emphasis id="B949581D0B54C868FD29FA00FE2C5CDD" box="[726,895,1465,1488]" italics="true" pageId="17" pageNumber="740">O. rhyacophilus</emphasis>
</taxonomicName>
by the color of the thoracic vittae (black in
<taxonomicName id="4C3DFF8C0B54C868FE94FA60FF415CFD" box="[363,530,1497,1520]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="740" phylum="Arthropoda" rank="species" species="rhyacophilus">
<emphasis id="B949581D0B54C868FE94FA60FF415CFD" box="[363,530,1497,1520]" italics="true" pageId="17" pageNumber="740">O. rhyacophilus</emphasis>
</taxonomicName>
versus yolk-colored ‘vitellines’ in
<taxonomicName id="4C3DFF8C0B54C868FC86FA60F95E5CFD" authorityName="Kieffer" authorityYear="1911" box="[889,1037,1497,1520]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="740" phylum="Arthropoda" rank="species" species="rhyacobius">
<emphasis id="B949581D0B54C868FC86FA60F95E5CFD" box="[889,1037,1497,1520]" italics="true" pageId="17" pageNumber="740">O. rhyacobius</emphasis>
</taxonomicName>
) and by the relative lengths of palpomeres (Pm) 3 and 4, with the Pm 3 being longer than Pm
<quantity id="4CC529EA0B54C868FBE7FA40F9125F1D" box="[1048,1089,1529,1552]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="17" pageNumber="740" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C3DFF8C0B57C86BFF68FF65FC7F59FE" authorityName="Kieffer" authorityYear="1911" box="[151,300,220,243]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="species" species="rhyacobius">
<emphasis id="B949581D0B57C86BFF68FF65FC7F59FE" box="[151,300,220,243]" italics="true" pageId="18" pageNumber="741">O. rhyacobius</emphasis>
</taxonomicName>
(‘articles 2 des palpes plus long che le 3’) versus the reverse in
<taxonomicName id="4C3DFF8C0B57C86BFC42FF65F93559FE" box="[957,1126,220,243]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="species" species="rhyacophilus">
<emphasis id="B949581D0B57C86BFC42FF65F93559FE" box="[957,1126,220,243]" italics="true" pageId="18" pageNumber="741">O. rhyacophilus</emphasis>
</taxonomicName>
(‘articles 2 des palpes plus court che le 3’). It is emphasized (
<bibRefCitation id="EFACF9FE0B57C86BFCDCFF45F909581E" author="SPIES M. &amp; SAETHER O. A." box="[803,1114,252,275]" pageId="18" pageNumber="741" pagination="1 - 90" refId="ref10840" refString="SPIES M. &amp; SAETHER O. A. 2004: Notes and recommendations on taxonomy and nomenclature of Chironomidae (Diptera). Zootaxa 752: 1 - 90." type="journal article" year="2004">SPIES &amp; SAETHER 2004: 25‒26</bibRefCitation>
), that Pm 3 and Pm 4 correspond to articles 2 and 3 of
<bibRefCitation id="EFACF9FE0B57C86BFD35FEA5FE38583E" author="KIEFFER J. J." box="[714,875,284,307]" pageId="18" pageNumber="741" pagination="181 - 187" refId="ref9890" refString="KIEFFER J. J. 1911: Nouveaux Tendipedides du groupe Orthocladius [Dipt.]. (1 re note). Bulletin de la Societe Entomologique de France 1911: 181 - 187." type="journal article" year="1911">KIEFFER (1911)</bibRefCitation>
. All the examined adult specimens of various species in
<taxonomicName id="4C3DFF8C0B57C86BFE06FE85FFD5585E" authorityName="sensu Goetghebuer" authorityYear="1940" box="[505,646,316,339]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B949581D0B57C86BFE06FE85FFD5585E" box="[505,646,316,339]" italics="true" pageId="18" pageNumber="741">Orthocladius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27AE5660B57C86BFD70FE85FF99585E" box="[655,714,316,339]" pageId="18" pageNumber="741" sensu="stricto">s. str.</taxonomicNameLabel>
that are present in the Department of Food, Environmental and Nutritional Science at the University of Milano (DeFENS) have Pm 3 longer than or subequal to Pm 4, never shorter, with the sole possible exception of
<taxonomicName id="4C3DFF8C0B57C86BFBB3FEC5FCB358BE" authority="Brundin, 1956" authorityName="Brundin" authorityYear="1956" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="species" species="wetterensis" subGenus="Orthocladius">
<emphasis id="B949581D0B57C86BFBB3FEC5F934589E" box="[1100,1127,380,403]" italics="true" pageId="18" pageNumber="741">O.</emphasis>
(
<emphasis id="B949581D0B57C86BFF5FFE25FDE958BE" box="[160,186,412,435]" italics="true" pageId="18" pageNumber="741">O.</emphasis>
)
<emphasis id="B949581D0B57C86BFF37FE25FC6C58BE" box="[200,319,412,435]" italics="true" pageId="18" pageNumber="741">wetterensis</emphasis>
Brundin, 1956
</taxonomicName>
. The color of the vittae is variable, winter specimens are often black, while spring specimens are lighter.
</paragraph>
<paragraph id="8B82840F0B57C86BFF4AFE65FE8F5B9E" blockId="18.[151,1127,220,659]" pageId="18" pageNumber="741">
The RBINS males examined are in poor condition, the color of the vittae was not registered in the pinned specimens and cannot be evaluated in the mounted slide. Consequently, in light of the problems discussed in detail by
<bibRefCitation id="EFACF9FE0B57C86BFDA0FDA5FEF15B3E" author="SPIES M. &amp; SAETHER O. A." box="[607,930,540,563]" pageId="18" pageNumber="741" pagination="1 - 90" refId="ref10840" refString="SPIES M. &amp; SAETHER O. A. 2004: Notes and recommendations on taxonomy and nomenclature of Chironomidae (Diptera). Zootaxa 752: 1 - 90." type="journal article" year="2004">SPIES &amp; SAETHER (2004: 24‒26)</bibRefCitation>
, the status of these two species (
<taxonomicName id="4C3DFF8C0B57C86BFED8FD85FCE15B5E" box="[295,434,572,595]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="species" species="rhycobius">
<emphasis id="B949581D0B57C86BFED8FD85FCE15B5E" box="[295,434,572,595]" italics="true" pageId="18" pageNumber="741">O. rhycobius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3DFF8C0B57C86BFE15FD85FFCB5B5E" box="[490,664,572,595]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="species" species="rhyacophilus">
<emphasis id="B949581D0B57C86BFE15FD85FFCB5B5E" box="[490,664,572,595]" italics="true" pageId="18" pageNumber="741">O. rhyacophilus</emphasis>
</taxonomicName>
) remains unresolved. In the present work, the solution proposed by
<bibRefCitation id="EFACF9FE0B57C86BFE55FDE5FF965B7E" author="ASHE P. &amp; O'CONNOR J. P." box="[426,709,604,627]" pageId="18" pageNumber="741" refId="ref9623" refString="ASHE P. &amp; O'CONNOR J. P. 2012: A World Catalogue of Chironomidae (Diptera) Part 2. Orthocladiinae. The Irish Biographical Society and National Museum of Ireland, Dublin, 968 pp." type="book" year="2012">ASHE &amp; O’CONNOR (2012)</bibRefCitation>
is followed, considering
<taxonomicName id="4C3DFF8C0B57C86BFC25FDE5FF385B9E" authority="Kieffer, 1911" authorityName="Kieffer" authorityYear="1911" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="species" species="rhyacophilous" subGenus="Orthocladius">
<emphasis id="B949581D0B57C86BFC25FDE5F9345B7E" box="[986,1127,604,627]" italics="true" pageId="18" pageNumber="741">Orthocladius</emphasis>
(
<emphasis id="B949581D0B57C86BFF5FFDC5FC7E5B9E" box="[160,301,636,659]" italics="true" pageId="18" pageNumber="741">Orthocladius</emphasis>
)
<emphasis id="B949581D0B57C86BFEC3FDC5FC875B9E" box="[316,468,636,659]" italics="true" pageId="18" pageNumber="741">rhyacophilous</emphasis>
<bibRefCitation id="EFACF9FE0B57C86BFE24FDC5FF385B9E" author="KIEFFER J. J." box="[475,619,636,659]" pageId="18" pageNumber="741" pagination="181 - 187" refId="ref9890" refString="KIEFFER J. J. 1911: Nouveaux Tendipedides du groupe Orthocladius [Dipt.]. (1 re note). Bulletin de la Societe Entomologique de France 1911: 181 - 187." type="journal article" year="1911">Kieffer, 1911</bibRefCitation>
</taxonomicName>
, a
<emphasis id="B949581D0B57C86BFD77FDC5FE755B9E" box="[648,806,636,659]" italics="true" pageId="18" pageNumber="741">nomen dubium</emphasis>
in
<emphasis id="B949581D0B57C86BFCB7FDC5FE8F5B9E" box="[840,988,636,659]" italics="true" pageId="18" pageNumber="741">
<taxonomicName id="4C3DFF8C0B57C86BFCB7FDC5FE8B5B9E" authorityName="sensu Goetghebuer" authorityYear="1940" box="[840,984,636,659]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="741" phylum="Arthropoda" rank="genus">Orthocladius</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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