302 lines
39 KiB
XML
302 lines
39 KiB
XML
<document id="C366C743CA42844E2AE71424076B6A5A" ID-DOI="10.11646/zootaxa.5109.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6336940" ID-ZooBank="7B6C0651-0935-4C77-B157-3ABF53E3AD81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646727632931" checkinUser="plazi" docAuthor="Jałoszyński, Paweł & Ślipiński, Adam" docDate="2022" docId="038B11419C6D700ED4D8FB77EA05FBA3" docLanguage="en" docName="zootaxa.5109.1.1.pdf.imf" docOrigin="Zootaxa 5109 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5109.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Murmidius campbelli Jałoszyński & Ślipiński 2022, sp. n." docType="treatment" docVersion="5" lastPageNumber="35" masterDocId="FFB269399C4D702CD44FFFD9E928FFB2" masterDocTitle="Revision of the family Murmidiidae (Coleoptera: Coccinelloidea)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="33" updateTime="1699303175474" updateUser="plazi">
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<mods:titleInfo id="4CA514820763238F7500ACDC22AD2BD9">
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<mods:title id="CF2052F375500EFBCAD20B8755A5BB94">Revision of the family Murmidiidae (Coleoptera: Coccinelloidea)</mods:title>
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<mods:name id="EB818FD67E2F1CAD1EAAA0C4B6327BA8" type="personal">
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<mods:roleTerm id="259BAA705ECD26100ED375096152322A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FD127E31563536054F39B01DD480E09E">Jałoszyński, Paweł</mods:namePart>
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<mods:nameIdentifier id="F1ED32FC1A839CBB29F553AB0A1EE074" type="ORCID">0000-0003-2973-1803</mods:nameIdentifier>
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<mods:nameIdentifier id="AAEC4570676AC3401C62FC933A667992" type="email">scydmaenus@yahoo.com</mods:nameIdentifier>
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<mods:name id="72ECBD5F251D5C0CD06A020F00DDE2AB" type="personal">
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<mods:roleTerm id="C1D0AEC0262243F1DC27ED334B951424">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FCD5CBD7528EF10DF818287A1415E1E5">Ślipiński, Adam</mods:namePart>
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<mods:nameIdentifier id="5C02AAF312C2F28CBED5D8642299FA26" type="ORCID">0000-0003-2973-1803</mods:nameIdentifier>
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<mods:nameIdentifier id="0F941C9CDDE566576D5611EDF403E825" type="email">scydmaenus@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="03AB6026E98B7652A155FB9CE7177A75">text</mods:typeOfResource>
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<mods:title id="2BF0FEC04406802D2A1CB897E40BB925">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0D2064C45965C894AC2E7C472EF2C446">
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<mods:date id="AC18AE1DA2B73871AD54B883BCEAE57F">2022</mods:date>
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<mods:number id="984B52D60DC43DF77A02CC4508181861">2022-03-08</mods:number>
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<mods:number id="192367D8C293D2F46BED4492110E7265">5109</mods:number>
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<mods:detail id="9061EEC582C161572AC8F77F68A20FA7" type="issue">
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<mods:number id="6666A6694493C991AC671C7C20EC7382">1</mods:number>
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<mods:url id="FB0323FAF2A1538EF7EC5A26BE7094F7">http://dx.doi.org/10.11646/zootaxa.5109.1.1</mods:url>
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</mods:location>
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<mods:classification id="AFD9A18378885BFEEFDEDE16E573AC2B">journal article</mods:classification>
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<mods:identifier id="083055DCE8EC5D812291907882F26180" type="DOI">10.11646/zootaxa.5109.1.1</mods:identifier>
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<mods:identifier id="13AFF3D023F75FF628417FF07675FA4B" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="038B11419C6D700ED4D8FB77EA05FBA3" LSID="urn:lsid:plazi:treatment:038B11419C6D700ED4D8FB77EA05FBA3" httpUri="http://treatment.plazi.org/id/038B11419C6D700ED4D8FB77EA05FBA3" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
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<subSubSection id="C338F3DC9C6D700CD4D8FB77E8CDFB7B" box="[151,485,1198,1225]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8B9DA0579C6D700CD4D8FB77E8CDFB7B" blockId="32.[151,485,1198,1261]" box="[151,485,1198,1225]" pageId="32" pageNumber="33">
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<heading id="D0D5173B9C6D700CD4D8FB77E8CDFB7B" bold="true" box="[151,485,1198,1225]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
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<emphasis id="B9567C459C6D700CD4D8FB77E8CDFB7B" bold="true" box="[151,485,1198,1225]" pageId="32" pageNumber="33">
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<taxonomicName id="4C22DBD49C6D700CD4D8FB77E8B3FB7B" authority="Jałoszyński & Ślipiński, 2022" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[151,411,1198,1225]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="campbelli" status="sp. nov.">
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<emphasis id="B9567C459C6D700CD4D8FB77E8B3FB7B" bold="true" box="[151,411,1198,1225]" italics="true" pageId="32" pageNumber="33">Murmidius campbelli</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A265C13E9C6D700CD5EDFB76E8CDFB7B" box="[418,485,1199,1225]" pageId="32" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C338F3DC9C6D700CD4D8FB0AE8BFFB5F" box="[151,407,1235,1261]" pageId="32" pageNumber="33" type="description">
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<paragraph id="8B9DA0579C6D700CD4D8FB0AE8BFFB5F" blockId="32.[151,485,1198,1261]" box="[151,407,1235,1261]" pageId="32" pageNumber="33">
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(
|
||
<figureCitation id="1319BCD29C6D700CD4D0FB0AE808FB5F" box="[159,288,1235,1261]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="32" pageNumber="33">Figs 81–87</figureCitation>
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,
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<figureCitation id="1319BCD29C6D700CD563FB0AE870FB5F" box="[300,344,1235,1261]" captionStart="FIGURES 123–128" captionStartId="44.[151,264,1783,1807]" captionTargetBox="[151,1436,181,1735]" captionTargetId="figure-19@44.[151,1436,181,1759]" captionTargetPageId="44" captionText="FIGURES 123–128. Murmidius campbelli group, comparison of microsculpture on heads of M. campbelli sp. n. (123), M. panamanus sp. n. (124), M. araguanus sp. n. (125), M. meridensis sp. n. (126), M. tachiranus sp. n. (127), and M. trujilloensis sp. n. (128)." figureDoi="http://doi.org/10.5281/zenodo.6336983" httpUri="https://zenodo.org/record/6336983/files/figure.png" pageId="32" pageNumber="33">123</figureCitation>
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,
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<figureCitation id="1319BCD29C6D700CD52BFB0AE8A7FB5F" box="[356,399,1235,1261]" captionStart="FIGURES 129–134" captionStartId="45.[151,264,1173,1197]" captionTargetBox="[151,1429,181,1137]" captionTargetId="figure-17@45.[151,1436,181,1149]" captionTargetPageId="45" captionText="FIGURES 129–134. Murmidius campbelli group, comparison of microsculpture on pronota of M. campbelli sp. n. (129), M. panamanus sp. n. (130), M. araguanus sp. n. (131), M. meridensis sp. n. (132), M. tachiranus sp. n. (133), and M. trujilloensis sp. n. (134)." figureDoi="http://doi.org/10.5281/zenodo.6336985" httpUri="https://zenodo.org/record/6336985/files/figure.png" pageId="32" pageNumber="33">129</figureCitation>
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||
)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C338F3DC9C6D700CD4D8FAC2EB50FA77" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8B9DA0579C6D700CD4D8FAC2EB50FA77" blockId="32.[151,1437,1306,2017]" pageId="32" pageNumber="33">
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<emphasis id="B9567C459C6D700CD4D8FAC2E8E9FA86" bold="true" box="[151,449,1306,1333]" pageId="32" pageNumber="33">Type material examined.</emphasis>
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||
<materialsCitation id="3B4AAA0A9C6D700CD586FAC2E89FFAEB" collectionCode="COLOM, CNC" collectorName="J. M. Campbell" country="Colombia" location="Colombia" pageId="32" pageNumber="33" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B9567C459C6D700CD586FAC2EBEDFA86" bold="true" box="[457,709,1306,1333]" pageId="32" pageNumber="33">
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||
<typeStatus id="54991EF59C6D700CD586FAC2EB1EFA87" box="[457,566,1307,1333]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
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||
(
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||
<collectingCountry id="F335E0C79C6D700CD60AFAC3EB95FA86" box="[581,701,1306,1332]" name="Colombia" pageId="32" pageNumber="33">Colombia</collectingCountry>
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)
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||
</emphasis>
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: sex unknown, “
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||
<collectionCode id="ED3338929C6D700CD7CCFAC2EAD9FA87" box="[899,1009,1307,1333]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12877" lsid="urn:lsid:biocol.org:col:12877" name="Colorado State Museum" pageId="32" pageNumber="33" type="Herbarium">COLOM</collectionCode>
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., Valle / Soladito, VII. / 20.1970,6700 /
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||
<collectorName id="26D7C5819C6D700CD4E6FAE6E87CFAEB" box="[169,340,1343,1369]" pageId="32" pageNumber="33">J.M. Campbell</collectorName>
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||
” (
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||
<collectionCode id="ED3338929C6D700CD53FFAE6E887FAEB" box="[368,431,1343,1369]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="32" pageNumber="33">CNC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B4AAA0A9C6D700CD58BFAE6E828FACF" collectionCode="CNC, MNHW" collectorName="J. M. Campbell" country="Colombia" location="Colombia" pageId="32" pageNumber="33" specimenCount="2" specimenCount-male="1" typeStatus="paratype">
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||
<emphasis id="B9567C459C6D700CD58BFAE6EB14FAEB" bold="true" box="[452,572,1343,1369]" pageId="32" pageNumber="33">
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||
<typeStatus id="54991EF59C6D700CD58BFAE6EB14FAEB" box="[452,572,1343,1369]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
(4 exx):
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||
<specimenCount id="9D246BDE9C6D700CD6E5FAE6EBFFFAEB" box="[682,727,1343,1369]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
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||
and
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||
<specimenCount id="9D246BDE9C6D700CD75CFAE6EA65FAEB" box="[787,845,1343,1369]" pageId="32" pageNumber="33" type="generic">1 ex.</specimenCount>
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||
of unknown sex, same data as for holotype (
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||
<collectionCode id="ED3338929C6D700CD115FAE6ECB0FAEB" box="[1370,1432,1343,1369]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="32" pageNumber="33">CNC</collectionCode>
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||
,
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<collectionCode id="ED3338929C6D700CD4D8FABAE9D0FACF" box="[151,248,1379,1405]" pageId="32" pageNumber="33">MNHW</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B4AAA0A9C6D700CD542FABAEB5DFA77" collectingDate="1981-01-21" collectionCode="PAN, FMNH" collectorName="W. Suter" country="Panama" county="Cerro" elevation="1290" location="Gato" municipality="Qda." pageId="32" pageNumber="33" specimenCode="FMHD #81-237, FMHD #81-72" specimenCount="2" stateProvince="Chiriqui" typeStatus="paratype">
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||
<emphasis id="B9567C459C6D700CD542FABAE850FACF" bold="true" box="[269,376,1379,1405]" pageId="32" pageNumber="33">
|
||
<collectingCountry id="F335E0C79C6D700CD542FABAE85BFACF" box="[269,371,1379,1405]" name="Panama" pageId="32" pageNumber="33">Panama</collectingCountry>
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||
:
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||
</emphasis>
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||
<specimenCount id="9D246BDE9C6D700CD5CEFABAE893FACF" box="[385,443,1379,1405]" pageId="32" pageNumber="33" type="generic">1 ex.</specimenCount>
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||
, “
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||
<collectionCode id="ED3338929C6D700CD59DFABAEB39FACF" box="[466,529,1379,1405]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13841" lsid="urn:lsid:biocol.org:col:13841" name="Panjab University" pageId="32" pageNumber="33" type="Herbarium">PAN</collectionCode>
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||
.:
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||
<collectingRegion id="49E66EB59C6D700CD66EFABAEB56FACF" box="[545,638,1379,1405]" country="Panama" name="Chiriqui" pageId="32" pageNumber="33">Chiriqui</collectingRegion>
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||
/ Prov.,
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||
<collectingCounty id="62FCD8DB9C6D700CD6AEFABAEA09FACF" box="[737,801,1379,1405]" pageId="32" pageNumber="33">Cerro</collectingCounty>
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||
Colora- / do (
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||
<quantity id="4CDA0DB29C6D700CD78AFABAED30FACF" box="[965,1048,1379,1405]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="32" pageNumber="33" unit="m" value="1290.0">
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<elevation id="000F47649C6D700CD78AFABAED30FACF" box="[965,1048,1379,1405]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="32" pageNumber="33" unit="m" value="1290.0">1290m</elevation>
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||
</quantity>
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||
), 24-I- / 1981”, “
|
||
<specimenCode id="DB84082C9C6D700CD094FABAECBFFACF" box="[1243,1431,1379,1405]" collectionCode="FMHD" pageId="32" pageNumber="33">FMHD #81-237</specimenCode>
|
||
, semi- / dry hillside litter, /
|
||
<collectorName id="26D7C5819C6D700CD5F2FA5EEB08FA13" box="[445,544,1415,1441]" pageId="32" pageNumber="33">W. Suter</collectorName>
|
||
” (
|
||
<collectionCode id="ED3338929C6D700CD678FA5EEBA6FA13" box="[567,654,1415,1441]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="32" pageNumber="33" type="Museum">FMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D246BDE9C6D700CD6EEFA5EEBF1FA13" box="[673,729,1415,1441]" pageId="32" pageNumber="33" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectionCode id="ED3338929C6D700CD6A1FA5EEA03FA13" box="[750,811,1415,1441]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13841" lsid="urn:lsid:biocol.org:col:13841" name="Panjab University" pageId="32" pageNumber="33" type="Herbarium">PAN</collectionCode>
|
||
:
|
||
<collectingRegion id="49E66EB59C6D700CD77BFA5EEAF3FA13" box="[820,987,1415,1441]" country="Panama" name="Bocas del Toro" pageId="32" pageNumber="33">Bocas del Toro</collectingRegion>
|
||
/ Prov.,
|
||
<quantity id="4CDA0DB29C6D700CD078FA5FEDADFA13" box="[1079,1157,1414,1441]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="32" pageNumber="33" unit="m" value="1345.0">
|
||
<elevation id="000F47649C6D700CD078FA5FEDADFA13" box="[1079,1157,1414,1441]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="32" pageNumber="33" unit="m" value="1345.0">1345m</elevation>
|
||
</quantity>
|
||
<collectingMunicipality id="6BF93A2D9C6D700CD0C4FA5EEDE9FA13" box="[1163,1217,1415,1441]" pageId="32" pageNumber="33">Qda.</collectingMunicipality>
|
||
/
|
||
<location id="8EFDF68C9C6D700CD09AFA5EEC26FA13" LSID="urn:lsid:plazi:treatment:038B11419C6D700ED4D8FB77EA05FBA3:8EFDF68C9C6D700CD09AFA5EEC26FA13" box="[1237,1294,1415,1441]" country="Panama" county="Cerro" municipality="Qda." name="Gato" pageId="32" pageNumber="33" stateProvince="Chiriqui">Gato</location>
|
||
,
|
||
<date id="FF9C86979C6D700CD156FA5EECA4FA13" box="[1305,1420,1415,1441]" pageId="32" pageNumber="33" value="1981-01-21">
|
||
<collectingDate id="EFD87F7F9C6D700CD156FA5EECA4FA13" box="[1305,1420,1415,1441]" pageId="32" pageNumber="33" value="1981-01-21">21-I-1981</collectingDate>
|
||
</date>
|
||
”, “
|
||
<specimenCode id="DB84082C9C6D700CD4EEFA72E865FA77" box="[161,333,1451,1477]" collectionCode="FMHD" pageId="32" pageNumber="33">FMHD #81-72</specimenCode>
|
||
, log, W. / Suter” (
|
||
<collectionCode id="ED3338929C6D700CD65AFA72EB44FA77" box="[533,620,1451,1477]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="32" pageNumber="33" type="Museum">FMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C6D700CD488FA17E9FFF89F" pageId="32" pageNumber="33" type="diagnosis">
|
||
<paragraph id="8B9DA0579C6D700CD488FA17E9FFF89F" blockId="32.[151,1437,1306,2017]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9567C459C6D700CD488FA17E86AFA5A" bold="true" box="[199,322,1486,1512]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
|
||
Body circular and strongly convex, dark brown, nearly black; antennal club over twice as long as broad; anterior clypeal margin smooth; pronotum with distinct, sulciform lateral longitudinal impressions, lacking sublateral impressions; frons and pronotum lacking microreticulation, vertex finely, transversely microreticulate, clypeus with isodiametric meshes; pronotum and elytra with very fine, inconspicuous punctures, elytra lacking large punctures arranged in rows; prosternal carinae reaching anterior ~1/3 of prosternum; mesoventral plate with evenly rounded anterior margin and submarginal carina interrupted just at middle, with strongly oblique lateral carinae; elytral epipleura not narrowed at the level of metaventrite and extremely broad, near middle broader than width of mesofemur; metaventrite entirely impunctate; penis stout, in lateral view weakly curved; in ventral view with broadly subtriangular apex; tegmen short and broad, with slightly emarginate apex bearing two pairs of short setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C6D700ED488F8EFE859FD37" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8B9DA0579C6D700CD488F8EFE826F80F" blockId="32.[151,1437,1306,2017]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9567C459C6D700CD488F8EFE871F8E2" bold="true" box="[199,345,1846,1872]" pageId="32" pageNumber="33">Description.</emphasis>
|
||
BL
|
||
<quantity id="4CDA0DB29C6D700CD5C2F8EEEB1DF8E3" box="[397,565,1847,1873]" metricMagnitude="-3" metricUnit="m" metricValue="1.465" metricValueMax="1.6" metricValueMin="1.33" pageId="32" pageNumber="33" unit="mm" value="1.465" valueMax="1.6" valueMin="1.33">1.33–1.60 mm</quantity>
|
||
; BL/EW 1.15–1.20. Body circular (
|
||
<figureCitation id="1319BCD29C6D700CD780F8EEED79F8E3" box="[975,1105,1847,1873]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="32" pageNumber="33">Figs 81–82</figureCitation>
|
||
) and in lateral view strongly convex (
|
||
<figureCitation id="1319BCD29C6D700CD4B7F882E865F8C7" box="[248,333,1883,1909]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="32" pageNumber="33">Fig. 84</figureCitation>
|
||
); pigmentation dark brown, nearly black, legs slightly lighter; dorsum very finely setose and with inconspicuous, fine and sparse punctures; cuticle strongly glossy, only clypeus and vertex finely microreticulate (
|
||
<figureCitation id="1319BCD29C6D700CD4EFF87AE82AF80F" box="[160,258,1955,1981]" captionStart="FIGURES 123–128" captionStartId="44.[151,264,1783,1807]" captionTargetBox="[151,1436,181,1735]" captionTargetId="figure-19@44.[151,1436,181,1759]" captionTargetPageId="44" captionText="FIGURES 123–128. Murmidius campbelli group, comparison of microsculpture on heads of M. campbelli sp. n. (123), M. panamanus sp. n. (124), M. araguanus sp. n. (125), M. meridensis sp. n. (126), M. tachiranus sp. n. (127), and M. trujilloensis sp. n. (128)." figureDoi="http://doi.org/10.5281/zenodo.6336983" httpUri="https://zenodo.org/record/6336983/files/figure.png" pageId="32" pageNumber="33">Fig. 123</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C6D700DD488F81EEA1FFF67" blockId="32.[151,1437,1306,2017]" lastBlockId="33.[151,1436,151,213]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
Head (
|
||
<figureCitation id="1319BCD29C6D700CD55EF81EE847F853" box="[273,367,1991,2017]" captionStart="FIGURES 123–128" captionStartId="44.[151,264,1783,1807]" captionTargetBox="[151,1436,181,1735]" captionTargetId="figure-19@44.[151,1436,181,1759]" captionTargetPageId="44" captionText="FIGURES 123–128. Murmidius campbelli group, comparison of microsculpture on heads of M. campbelli sp. n. (123), M. panamanus sp. n. (124), M. araguanus sp. n. (125), M. meridensis sp. n. (126), M. tachiranus sp. n. (127), and M. trujilloensis sp. n. (128)." figureDoi="http://doi.org/10.5281/zenodo.6336983" httpUri="https://zenodo.org/record/6336983/files/figure.png" pageId="32" pageNumber="33">Fig. 123</figureCitation>
|
||
)
|
||
<quantity id="4CDA0DB29C6D700CD531F81FEB08F853" box="[382,544,1990,2017]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="32" pageNumber="33" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.40–0.50 mm</quantity>
|
||
wide; frons weakly convex, with fine, inconspicuous punctures, glossy; clypeus matt and finely microreticulate, vertex with transverse microreticulation but glossy; eyes large, strongly convex, coarsely faceted. Antennal club over twice as long as broad.
|
||
</paragraph>
|
||
<caption id="DF5DF0DF9C6C700DD4D8F9B3EA74F915" ID-DOI="http://doi.org/10.5281/zenodo.6336969" ID-Zenodo-Dep="6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,264,1642,1666]" targetBox="[151,1436,237,1619]" targetPageId="33">
|
||
<paragraph id="8B9DA0579C6C700DD4D8F9B3EA74F915" blockId="33.[151,1437,1642,1703]" pageId="33" pageNumber="34">
|
||
<emphasis id="B9567C459C6C700DD4D8F9B3E87BF930" bold="true" box="[151,339,1642,1666]" pageId="33" pageNumber="34">FIGURES 81–87.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C22DBD49C6C700DD638F9B3EA78F930" authority="Jałoszyński & Ślipiński, 2022" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[631,848,1642,1666]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli" status="sp. nov.">
|
||
<emphasis id="B9567C459C6C700DD638F9B3EA78F930" box="[631,848,1642,1666]" italics="true" pageId="33" pageNumber="34">Murmidius campbelli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9567C459C6C700DD71AF9B2EAA5F931" bold="true" box="[853,909,1643,1667]" pageId="33" pageNumber="34">
|
||
<taxonomicNameLabel id="A265C13E9C6C700DD71AF9B2EAA5F931" box="[853,909,1643,1667]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9DA0579C6C700DD488F90CEB25F859" blockId="33.[151,1437,1749,2027]" pageId="33" pageNumber="34">
|
||
Pronotum (
|
||
<figureCitation id="1319BCD29C6C700DD507F90CE8E6F95D" box="[328,462,1749,1775]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="33" pageNumber="34">Figs 81–82</figureCitation>
|
||
,
|
||
<figureCitation id="1319BCD29C6C700DD593F90CEB2FF95D" box="[476,519,1749,1775]" captionStart="FIGURES 129–134" captionStartId="45.[151,264,1173,1197]" captionTargetBox="[151,1429,181,1137]" captionTargetId="figure-17@45.[151,1436,181,1149]" captionTargetPageId="45" captionText="FIGURES 129–134. Murmidius campbelli group, comparison of microsculpture on pronota of M. campbelli sp. n. (129), M. panamanus sp. n. (130), M. araguanus sp. n. (131), M. meridensis sp. n. (132), M. tachiranus sp. n. (133), and M. trujilloensis sp. n. (134)." figureDoi="http://doi.org/10.5281/zenodo.6336985" httpUri="https://zenodo.org/record/6336985/files/figure.png" pageId="33" pageNumber="34">129</figureCitation>
|
||
) strongly transverse, widest at base; PL
|
||
<quantity id="4CDA0DB29C6C700DD7AAF90CED67F95D" box="[997,1103,1749,1776]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="33" pageNumber="34" unit="mm" value="0.25">0.25 mm</quantity>
|
||
, PW 0.88–1.00 mm, PL/PW 0.25–0.29; lateral margins weakly rounded and very strongly convergent anterad, with narrow lateral carinae; mesal corners of antennal cavities distinctly projecting anterad; lateral longitudinal impressions distinct, broad and deep anteriorly and reducing in width and depth toward pronotal base, posteriorly convergent with lateral pronotal margins; sublateral longitudinal impressions lacking; anterior pronotal margin not swollen and with barely discernible marginal carina. Punctures on disc (
|
||
<figureCitation id="1319BCD29C6C700DD661F850EBA7F811" box="[558,655,1929,1955]" captionStart="FIGURES 129–134" captionStartId="45.[151,264,1173,1197]" captionTargetBox="[151,1429,181,1137]" captionTargetId="figure-17@45.[151,1436,181,1149]" captionTargetPageId="45" captionText="FIGURES 129–134. Murmidius campbelli group, comparison of microsculpture on pronota of M. campbelli sp. n. (129), M. panamanus sp. n. (130), M. araguanus sp. n. (131), M. meridensis sp. n. (132), M. tachiranus sp. n. (133), and M. trujilloensis sp. n. (134)." figureDoi="http://doi.org/10.5281/zenodo.6336985" httpUri="https://zenodo.org/record/6336985/files/figure.png" pageId="33" pageNumber="34">Fig. 129</figureCitation>
|
||
) very fine, barely discernible, unevenly distributed, those on median area separated by spaces 2–5 times as wide as their diameters, large punctures lacking, surface of disc strongly glossy, lacking microreticulation.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C6F700ED488FF4EEB6EFEAF" blockId="34.[151,1437,151,1041]" pageId="34" pageNumber="35">
|
||
Prosternum (
|
||
<figureCitation id="1319BCD29C6F700ED519FF4EE88EFF03" box="[342,422,151,177]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="34" pageNumber="35">Fig. 83</figureCitation>
|
||
) with barely discernible (even with SEM), superficial transverse microreticulation and with fine, inconspicuous punctures; near middle with pair of small glandular openings; prosternal carinae short, anteriorly nearly reaching 1/3 of prosternum; notosternal carinae weakly sinuate, not reaching anterior prosternal margin, the latter with narrow marginal carina.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C6F700ED488FEFEEC64FED7" blockId="34.[151,1437,151,1041]" pageId="34" pageNumber="35">
|
||
Elytra (
|
||
<figureCitation id="1319BCD29C6F700ED552FEFEE8E5FEF3" box="[285,461,295,321]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="34" pageNumber="35">Figs 81–82, 84</figureCitation>
|
||
) together oval, with strongly rounded sides, broadest slightly in front of middle, EL 1.00–
|
||
<quantity id="4CDA0DB29C6F700ED498FE92E814FED7" box="[215,316,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="34" pageNumber="35" unit="mm" value="1.28">1.28 mm</quantity>
|
||
, EW
|
||
<quantity id="4CDA0DB29C6F700ED535FE93EB37FED7" box="[378,543,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.265" metricValueMax="1.38" metricValueMin="1.15" pageId="34" pageNumber="35" unit="mm" value="1.265" valueMax="1.38" valueMin="1.15">1.15–1.38 mm</quantity>
|
||
, EL/EW 0.87–0.94. Humerus lacking callus; elytra virtually impunctate.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C6F700ED488FEB6EAF2FE3B" blockId="34.[151,1437,151,1041]" box="[199,986,367,393]" pageId="34" pageNumber="35">Hind wings reduced, narrow and about as long as elytra, unfunctional.</paragraph>
|
||
<paragraph id="8B9DA0579C6F700ED488FE4AEC6DFE63" blockId="34.[151,1437,151,1041]" pageId="34" pageNumber="35">
|
||
Mesoventral plate (
|
||
<figureCitation id="1319BCD29C6F700ED5D0FE4AE8D9FE1F" box="[415,497,402,429]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="34" pageNumber="35">Fig. 85</figureCitation>
|
||
) with lateral longitudinal carinae present and strongly oblique, running at 45°-angle in relation to long body axis; anterior margin evenly rounded, with submarginal carina disrupted at middle.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C6F700ED488FE02ED35FE47" blockId="34.[151,1437,151,1041]" box="[199,1053,474,501]" pageId="34" pageNumber="35">
|
||
Metaventrite (
|
||
<figureCitation id="1319BCD29C6F700ED529FE02E89FFE47" box="[358,439,474,501]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="34" pageNumber="35">Fig. 85</figureCitation>
|
||
) entirely impunctate. Discrimen externally not visible.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C6F700ED488FE26EAB3FDAB" blockId="34.[151,1437,151,1041]" box="[199,923,511,537]" pageId="34" pageNumber="35">
|
||
Tarsi with conspicuously long setae on tarsomeres 1–3 (
|
||
<figureCitation id="1319BCD29C6F700ED775FE26EAA5FDAB" box="[826,909,511,537]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="34" pageNumber="35">Fig. 83</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C6F700ED488FDFAE859FD37" blockId="34.[151,1437,151,1041]" pageId="34" pageNumber="35">
|
||
Aedeagus (
|
||
<figureCitation id="1319BCD29C6F700ED509FDFAE8EFFD8C" box="[326,455,547,574]" captionStart="FIGURES 81–87" captionStartId="33.[151,264,1642,1666]" captionTargetBox="[151,1436,237,1619]" captionTargetId="figure-41@33.[151,1436,237,1619]" captionTargetPageId="33" captionText="FIGURES 81–87. Morphological structures of Murmidius campbelli sp. n. Body in dorsal (81–82), ventral (83) and lateral (84) views; pterothorax in ventral view (85); aedeagus (79) and penis (87)." figureDoi="http://doi.org/10.5281/zenodo.6336969" httpUri="https://zenodo.org/record/6336969/files/figure.png" pageId="34" pageNumber="35">Figs 86–87</figureCitation>
|
||
)
|
||
<quantity id="4CDA0DB29C6F700ED598FDFAEB15FD8F" box="[471,573,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="34" pageNumber="35" unit="mm" value="0.98">0.98 mm</quantity>
|
||
long, conspicuously stout; penis in ventral view with broadly subtriangular, blunt apex, in lateral view weakly curved and extremely thick; tegmen with shallowly emarginate apex bearing 2 pairs of short setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C6F700ED488FD57EA22FD1B" box="[199,778,654,681]" pageId="34" pageNumber="35" type="distribution">
|
||
<paragraph id="8B9DA0579C6F700ED488FD57EA22FD1B" blockId="34.[151,1437,151,1041]" box="[199,778,654,681]" pageId="34" pageNumber="35">
|
||
<emphasis id="B9567C459C6F700ED488FD57E849FD1A" bold="true" box="[199,353,654,680]" pageId="34" pageNumber="35">Distribution.</emphasis>
|
||
Neotropical (
|
||
<collectingCountry id="F335E0C79C6F700ED5B5FD56EB42FD1B" box="[506,618,655,681]" name="Colombia" pageId="34" pageNumber="35">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F335E0C79C6F700ED6EEFD56EBD7FD1B" box="[673,767,655,681]" name="Panama" pageId="34" pageNumber="35">Panama</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C6F700ED488FD6AE8A7FD43" pageId="34" pageNumber="35" type="etymology">
|
||
<paragraph id="8B9DA0579C6F700ED488FD6AE8A7FD43" blockId="34.[151,1437,151,1041]" pageId="34" pageNumber="35">
|
||
<emphasis id="B9567C459C6F700ED488FD6AE860FD7F" bold="true" box="[199,328,691,717]" pageId="34" pageNumber="35">Etymology</emphasis>
|
||
. Name dedicated to Dr. Milton Campbell, the former curator of the
|
||
<taxonomicName id="4C22DBD49C6F700ED072FD6AED90FD7F" authorityName="Linnaeus" authorityYear="1758" box="[1085,1208,691,717]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="100" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
at CNC and the collector of the
|
||
<typeStatus id="54991EF59C6F700ED56AFD0EE8A3FD43" box="[293,395,727,753]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C6F700ED488FD22EA05FBA3" pageId="34" pageNumber="35" type="discussion">
|
||
<paragraph id="8B9DA0579C6F700ED488FD22EA05FBA3" blockId="34.[151,1437,151,1041]" pageId="34" pageNumber="35">
|
||
<emphasis id="B9567C459C6F700ED488FD22E813FCA7" bold="true" box="[199,315,763,789]" pageId="34" pageNumber="35">Remarks.</emphasis>
|
||
Within this species group,
|
||
<taxonomicName id="4C22DBD49C6F700ED625FD22EBD5FCA7" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[618,765,763,789]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="campbelli">
|
||
<emphasis id="B9567C459C6F700ED625FD22EBD5FCA7" box="[618,765,763,789]" italics="true" pageId="34" pageNumber="35">M. campbelli</emphasis>
|
||
</taxonomicName>
|
||
is the only member that lacks any large punctures on elytra, and has a microreticulate clypeus and vertex, but not the frons and pronotum. Its aedeagus is most similar to that of
|
||
<taxonomicName id="4C22DBD49C6F700ED4FAFC9AE873FCEE" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[181,347,835,860]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="panamanus">
|
||
<emphasis id="B9567C459C6F700ED4FAFC9AE873FCEE" box="[181,347,835,860]" italics="true" pageId="34" pageNumber="35">M. panamanus</emphasis>
|
||
</taxonomicName>
|
||
, except for having more subtriangular apex. Externally these two species can be distinguished by the prosternal carinae in
|
||
<taxonomicName id="4C22DBD49C6F700ED5FEFCBEEB6DFC33" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[433,581,871,897]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="campbelli">
|
||
<emphasis id="B9567C459C6F700ED5FEFCBEEB6DFC33" box="[433,581,871,897]" italics="true" pageId="34" pageNumber="35">M. campbelli</emphasis>
|
||
</taxonomicName>
|
||
nearly reaching 1/3 of prosternum and straight (in
|
||
<taxonomicName id="4C22DBD49C6F700ED0C3FCBEEC1CFC32" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[1164,1332,871,896]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="panamanus">
|
||
<emphasis id="B9567C459C6F700ED0C3FCBEEC1CFC32" box="[1164,1332,871,896]" italics="true" pageId="34" pageNumber="35">M. panamanus</emphasis>
|
||
</taxonomicName>
|
||
reaching median glandular openings and each carina with its anterior end bent mesad at a right angle), the mesoventral plate with strongly oblique, complete lateral carinae (lacking any trace of lateral carinae in
|
||
<taxonomicName id="4C22DBD49C6F700ED01BFC76EDD3FC7A" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[1108,1275,943,968]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="panamanus">
|
||
<emphasis id="B9567C459C6F700ED01BFC76EDD3FC7A" box="[1108,1275,943,968]" italics="true" pageId="34" pageNumber="35">M. panamanus</emphasis>
|
||
</taxonomicName>
|
||
), microreticulate vertex (smooth in
|
||
<taxonomicName id="4C22DBD49C6F700ED5DEFC0AEB1EFC5E" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[401,566,979,1004]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="panamanus">
|
||
<emphasis id="B9567C459C6F700ED5DEFC0AEB1EFC5E" box="[401,566,979,1004]" italics="true" pageId="34" pageNumber="35">M. panamanus</emphasis>
|
||
</taxonomicName>
|
||
), and elytra lacking any large punctures (in
|
||
<taxonomicName id="4C22DBD49C6F700ED06FFC0AEDEEFC5E" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[1056,1222,979,1004]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="panamanus">
|
||
<emphasis id="B9567C459C6F700ED06FFC0AEDEEFC5E" box="[1056,1222,979,1004]" italics="true" pageId="34" pageNumber="35">M. panamanus</emphasis>
|
||
</taxonomicName>
|
||
there are ~15 large punctures on sides arranged in two slightly irregular rows).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |